SOUTEHRN TUNESIA
JEBEL EL MRA
OUM ED DIAB MEMBER
AIN EL GUETTAR FORMATION
EARLY ALBIAN STAGE
MIDDLE GALLIC SUBEPOCH
LATE EARLY CRETACEOUS EPOCH
EARLY CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: ORNITHISCHIFORMES COOPER, 1985
SUPERORDER: SAUROPODOMORPHA HUENE, 1932
GRANDORDER: BRONTOSAURIA OLSHEVSKY, 1991 (emend)
MICRORDER: SAUROPODA MARSH, 1878
ORDER: EUSAUROPODA
SUBORDER: NEOSAUROPODA BONAPARTE, 1986
INFRAORDER: HOMALOSAUROPIA JANENSCH, 1929
PARVORDER: DIPLODOCIMORPHA CALVO & SALGADO, 1995
Superfamily: DIPLODOCOIDEA Upchurch, 1995
Family: REBBACHISAURIDAE Bonaparte, 1996
*Tataouinea hannibalis
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TUNISIA
BATEUN EL HMAIMA
CHENINI MEMBER
APTIAN-ALBIAN STAGE
GALLIC SUBEPOCH
EARLY CRETACEOUS EPOCH
EARLY CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: CRUROTARSI SERENO & ARCUCCI, 1990
COHORT: CROCODYLOTARSI BENTON & CLARK, 1988
MAGNORDER: SUCHIA KREBS, 1974
SUPERORDER: CROCODYLOMORPHA WALKER, 1968
GRANDORDER: CROCODILIFORMES HAY, 1930
MICRORDER: MESOEUROCROCODYLIA WHETSTONE & WHYBROW, 1983
ORDER: EUSUCHIA emend HUXLEY, 1875
INFRAORDER: CROCODYLIA emend GMELIN, 1877
Incertae sedis
Gen. sp indet.
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
Incertae sedis
Gen. sp indet.
AVEPODA PAUL, 2002
SUPERORDER: NEOTHEROPODA BAKKER, 1986
GRANDORDER: TETANURAE GAUTHIER, 1986
ORDER: SPINOSAURIA OLSHEVSKY, 1991
Family: SPINOSAURIDAE Stromer, 1915
CF. Spinosaurus sp
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SOUTHERN TUNISIA
NEAR TATAOUINE
CHENINI SITE
5 M BELOWE THE TOP OF THE OUM ED DIAB MEMBER
MIDDLE ALBIAN STAGE
MIDDLE GALLIC SUBEPOCH
LATE EARLY CRETACEOUS EPOCH
LATEST EARLY CRETACEOUS
ICHNOSTEM: TETRAPODIPIDA NOVA
ICHNOSUPERCLASS: AMNIOTAIPIDII NOVA
ICHNOCLASS: REPTILIPEDIA VIALOV, 1966
ICHNOSUBCLASS: DIAPSIDIPIDII NOVA
ICHNOSTEM-GROUP: NEODIAPSIDIPEDII NOVA
ICHNOINFRACLASS: NEODIAPSIDIPEDII NOVA
ICHNOSUBDIVISION: ARCHOSAURIPEDIDII NOVA
ICHNOLEGION: DINOSAUROMORPHIPEDA NOVA
ICHNOSUBLEGION: DINOSAURIFORMEIPEDIDII NOVA
ICHNOSUPERCOHORT: DINOSAUROPEDIDA NOVA
ICHNOCOHORT: ORNITHISCHIFORMEIPIDA NOVA
ICHNOMAGNORDER: ORNITHISCHIPIDA NOVA
ICHNOSUPERORDER: NEORNITHISCHIPIDA NOVA
ICHNOGRANDORDER: ORNITHOPODIPDIA NOVA
ICHNOMICRORDER: EUORNITHOPODIPIDA NOVA
ICHNOORDER: IGUANODONTIPIDIA NOVA
ICHNOSUBORDER: ANKYLOPOLLEXIPEDIDA NOVA
ICHNOINFRAORDER: STYRACOSTERNIPIDA NOVA
ICHNOPARVORDER: HADROSAURIFORMIPEDIDA NOVA
ICHNOSUPERFAMILY: HADROSAUROIDIPEDIA NOVA
Ichnofamily: HADROSAURIOPDIDAE Vylov, 1988
cf. Apulosauripus fredericianus
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TUNISIA
APPROXIMATELY 50 KM SOUTH
OF TATAOUINE
SOUTH-WEST OF THE EL MRA
LOCALITY
10 M ABOVE THE CHENINI/OUM
ED DIAB CONTACT
ALBIAN STAGE
MIDDLE GALLIC SUBEPOCH
UPPER EARLY CRETACEOUS
EPOCH
EARLY CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: CRUROTARSI SERENO & ARCUCCI, 1990
COHORT: CROCODYLOTARSI BENTON & CLARK, 1988
MAGNORDER: SUCHIA KREBS, 1974
SUPERORDER: CROCODYLOMORPHA WALKER, 1968
GRANDORDER: CROCODILIFORMES HAY, 1930
Gen. sp indet.
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: ORNITHISCHIFORMES COOPER, 1985
SUPERORDER: SAUROPODOMORPHA HUENE, 1932
GRANDORDER: BRONTOSAURIA OLSHEVSKY, 1991 (emend)
MICRORDER: SAUROPODA MARSH, 1878
Gen. sp indet.
MAGNORDER: ORNITHISCHIA SEELEY, 1888
SUPERORDER: NEORNITHISCHIA COOPER, 1985
GRANDORDER: ORNITHOPODA MARSH, 1871
MICRORDER: EUORNITHOPODA SERENO, 1986
ORDER: IGUANODONTIA DOLLO, 1888
SUBORDER: ANKYLOPOLLEXIA SERENO, 1986
INFRAORDER: STYRACOSTERNA SERENO, 1986
PARVORDER: HADROSAURIFORMES SERENO, 1997
IGUANODONTIA DOLLO, 1888
Superfamily: HADROSAUROIDEA Cope, 1869 (sensu Sereno, 1986)
Incertae sedis
Gen. sp indet.
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TUNISIA
5 KM TO THE SOUTH OF THE
BIR AMIR VILLAGE
APPROXIMATELY 50 KM SOUTH
OF TATAOUINE
EL MRA LOCALITY
LOWER OUM ED DIAB MEMBER
APPROXIMATELY 5 METERS
ABOVE THE BASE OF THE UNIT
ALBIAN STAGE
MIDDLE GALLIC SUBEPOCH
UPPER EARLY CRETACEOUS
EPOCH
EARLY CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: ORNITHISCHIFORMES COOPER, 1985
MAGNORDER: ORNITHISCHIA SEELEY, 1888
SUPERORDER: NEORNITHISCHIA COOPER, 1985
GRANDORDER: ORNITHOPODA MARSH, 1871
MICRORDER: EUORNITHOPODA SERENO, 1986
ORDER: IGUANODONTIA DOLLO, 1888
SUBORDER: ANKYLOPOLLEXIA SERENO, 1986
INFRAORDER: STYRACOSTERNA SERENO, 1986
PARVORDER: HADROSAURIFORMES SERENO, 1997
IGUANODONTIA DOLLO, 1888
Superfamily: HADROSAUROIDEA Cope, 1869 (sensu Sereno, 1986)
Incertae sedis
Gen. sp indet.
_____________________________________________________________________________________
TUNISIA
TATAOUINE REGION
OUM ED DHIAB MEMBER
AIN EL GUETTAR FORMATION
APTIAN-ALBIAN STAGE
MIDDLE GALLIC SUBEPOCH
UPPER EARLY CRETACEOUS EPOCH
MIDDLE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: CRUROTARSI SERENO & ARCUCCI, 1990
COHORT: CROCODYLOTARSI BENTON & CLARK, 1988
MAGNORDER: SUCHIA KREBS, 1974
SUPERORDER: CROCODYLOMORPHA WALKER, 1968
GRANDORDER: CROCODILIFORMES HAY, 1930
Incertae sedis
Gen. sp indet.
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
Indertae sedis
Morphotype 1
Morphotype 2
Morphotype 3
Morphotype 4
Morphotype 5
Morphotype 6
Morphotype 7
Morphotype 8
Small theropod
Large theropod
AVEPODA PAUL, 2002
GRANDORDER: CERATOSAURIA NOVAS, 1992
AVEROSTRA PAUL, 2001
ORDER: CERATOSAURIA MARSH, 1884
Superfamily: ABELISAUROIDEA Bonaparte & Novas, 1985
Family: ABELISAURIDAE Bonaparte & Novas, 1985
Gen. sp indet.
SUPERORDER: NEOTHEROPODA BAKKER, 1986
GRANDORDER: TETANURAE GAUTHIER, 1986
ORDER: SPINOSAURIA OLSHEVSKY, 1991
Family: BARYONYCHIDAE Charig & Milner, 1986
Gen. sp indet.
Family: SPINOSAURIDAE Stromer, 1915
Spinosaurus sp
ORDER: NEOTETANURAE SERENO, WILSON, LARSSON, DUTHEIL &
SUBORDER: AVETHEROPODA PAUL, 1988
Superfamily: ALLOSAUROIDEA Marsh, 1878 (sensu Currie & Zhao, 1994)
Family: CARCHARODONTOSAURIDAE Stromer, 1931
Carcharodontosaurus sp
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SOUTH AFRICA
UMTATA RIVER MOUTH
CLIFFS OF SOUTH AFRICAN COAST
NEAR THE UMTATUNA NAD UMZAMBAWI RIVERS
ALBIAN-CENOMAINAN STAGE
MIDDLE GALLIC SUBEPOCH
UPPERMOST EARLY CRETACEOUS EPOCH OR LOWERMOST GULF EPOCH
LATE EARLY OR MIDDLE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: CHELONOMORPHA KUHN, 1960
SUPERCOHORT: TESTUDINES LINNAEUS, 1758
Chelonoidea or Dermochelyiodea incertae sedis
Gen sp indet.
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TUNISIA
ER RONDA-CHENINI TRAIL
GUERMESSA
DEHIBAT-GARA ER REHI
UPPERMOST ALBIAN-LOWERMOST CENOMANIAN STAGE
UPPER GALLIC SUBEPOCH
LOWEST GULF EPOCH
LATE EARLY TO EARLY MIDDLE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: CHELONOMORPHA KUHN, 1960
SUPERCOHORT: TESTUDINES LINNAEUS, 1758
COHORT: CASICHELYDIA emend GAFFNEY & MEYLAN, 1988
MAGNORDER: PLEURODIRA GAFFNEY & MEYLAN, 1988
ORDER: PLEURODIRIDA COPE, 1864
SUBORDER: EUPLEURODIRA GAFFNEY & MEYLAN, 1988
Superfamily: PELOMEDUSOIDES Cope, 1868
Family: PELOMEDUSIDAE Cope, 1868
Gen. sp indet.
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TUNISIA
TATOUINE REGION
CHENINI FORMATION
ALBIAN STAGE
MIDDLE GALLIC SUBEPOCH
UPPER EARLY CRETACEOUS EPOCH
EARLY CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
NEODIAPSIDA: BENTON, 1985
LEPIDOSAUROMORPHA: BENTON 1985
INFRACLASS: EURYAPSIDA COLBERT, 1945
SUPERORDER: SAUROPTERYGIA OWEN 1860
GRANDORDER: EOSAUROPTERYGIA emend RIEPPEL, 1994
ORDER: PLESIOSAURIA OWEN, 1840 NON BLAINVILLE, 1835
SUBORDER: PLESIOSAURIA OWEN, 1840 NON BLAINVILLE, 1835
INFRAORDER: PLESIOSAUROMORPHA NOVAA
Incertae sedis
Gen. sp indet.
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: AVEMETATARSALIA BENTON, 1999
COHORT: PROLACERTIFORMES CAMP, 1945 NOVA
SUPERORDER: TAPINOPLATIA PETERS, 2000
GRANDORDER: CHARACIOPODA PETERS, 2000
MICRORDER: FENESTRASAURIA PETERS, 2000
ORDER: PTEROSAURIA KAUP, 1834
SUBORDER: EUPTERODACTYLOIDEA BENNETT, 1994
Superfamily: ORNITHOCHEIROIDEA Seeley, 1870 (sensu Seeley, 1891)
Family: ORNITHOCHEIRIDAE Seeley, 1870
Incertae sedis
Gen sp indet.
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
ORDER: Incertae sedis
Gen. sp indet.
AVEPODA PAUL, 2002
SUPERORDER: NEOTHEROPODA BAKKER, 1986
GRANDORDER: TETANURAE GAUTHIER, 1986
ORDER: SPINOSAURIA OLSHEVSKY, 1991
Family: SPINOSAURIDAE Stromer, 1915
Spinosaurus sp
GRANDORDER: TETANURAE GAUTHIER, 1986
ORDER: NEOTETANURAE SERENO, WILSON, LARSSON, DUTHEIL &
SUBORDER: AVETHEROPODA PAUL, 1988
Superfamily: ALLOSAUROIDEA Marsh, 1878 (sensu Currie & Zhao, 1994)
Family: CARCHARODONTOSAURIDAE Stromer, 1931
Carcharodontosaurus sp
COHORT: ORNITHISCHIFORMES COOPER, 1985
SUPERORDER: SAUROPODOMORPHA HUENE, 1932
Incertae sedis
Gen. sp indet.
MAGNORDER: ORNITHISCHIA SEELEY, 1888
SUPERORDER: NEORNITHISCHIA COOPER, 1985
GRANDORDER: ORNITHOPODA MARSH, 1871
MICRORDER: EUORNITHOPODA SERENO, 1986
ORDER: IGUANODONTIA DOLLO, 1888
SUBORDER: ANKYLOPOLLEXIA SERENO, 1986
INFRAORDER: STYRACOSTERNA SERENO, 1986
PARVORDER: HADROSAURIFORMES SERENO, 1997
Superfamily: IGUANODONTIDEA Cope, 1869 (sensu Sereno, 1986)
Family: IGUANODONTIDAE Cope, 1869
Incertae sedis
Gen. sp indet.
____________________________________________________________________________________
TUNISIA
FOUM TAAHOUNE REGION
LOCALITY RH 45
CHENINNI FORMATION
ALBIAN STAGE
MIDDLE GALLIC SUBEPOCH
UPPER EARLY CRETACEOUS EPOCH
EARLY CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
AVEPODA PAUL, 2002
SUPERORDER: NEOTHEROPODA BAKKER, 1986
GRANDORDER: TETANURAE GAUTHIER, 1986
ORDER: SPINOSAURIA OLSHEVSKY, 1991
Family: SPINOSAURIDAE Stromer, 1915
Spinosaurus sp
GRANDORDER: TETANURAE GAUTHIER, 1986
ORDER: NEOTETANURAE SERENO, WILSON, LARSSON, DUTHEIL &
SUBORDER: AVETHEROPODA PAUL, 1988
Superfamily: ALLOSAUROIDEA Marsh, 1878 (sensu Currie & Zhao, 1994)
Family: CARCHARODONTOSAURIDAE Stromer, 1931
Carcharodontosaurus saharicus
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TUNISIA
TATAHOUME REGION
4 KM NW OF BIR MITEUR
CHENINI FORMATION
ALBIAN STAGE
MIDDLE GALLIC SUBEPOCH
UPPER EARLY CRETACEOUS EPOCH
EARLY CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: ORNITHISCHIFORMES COOPER, 1985
SUPERORDER: SAUROPODOMORPHA HUENE, 1932
Incertae sedis
Gen. sp indet.
_____________________________________________________________________________________
TUNISIA
TATAOUINE GOVERNORATE
JEBEL MITEUR
CHENINI FORMATION
ALBIAN STAGE
MIDDLE GALLIC SUBEPOCH
UPPER EARLY CRETACEOUS EPOCH
EARLY CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: CRUROTARSI SERENO & ARCUCCI, 1990
COHORT: CROCODYLOTARSI BENTON & CLARK, 1988
MAGNORDER: SUCHIA KREBS, 1974
SUPERORDER: CROCODYLOMORPHA WALKER, 1968
GRANDORDER: CROCODILIFORMES HAY, 1930
Incertae sedis
Gen sp indet.
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
AVEPODA PAUL, 2002
SUPERORDER: NEOTHEROPODA BAKKER, 1986
GRANDORDER: TETANURAE GAUTHIER, 1986
ORDER: SPINOSAURIA OLSHEVSKY, 1991
Family: SPINOSAURIDAE Stromer, 1915
Spinosaurus cf. aegyptiacus
GRANDORDER: TETANURAE GAUTHIER, 1986
ORDER: NEOTETANURAE SERENO, WILSON, LARSSON, DUTHEIL &
SUBORDER: AVETHEROPODA PAUL, 1988
Superfamily: ALLOSAUROIDEA Marsh, 1878 (sensu Currie & Zhao, 1994)
Family: CARCHARODONTOSAURIDAE Stromer, 1931
Carcharodontosaurus sp
COHORT: ORNITHISCHIFORMES COOPER, 1985
SUPERORDER: SAUROPODOMORPHA HUENE, 1932
Incertae sedis
Gen. sp indet.
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