SOUTHERN UNITED KINGDOM
ENGLAND
EAST SUSSEX COUNTY
BATTLE
HASTINGS SUB-GROUP
BERRIASIAN-VALANGINIAN STAGE
LOWER NEOCOMIAN SUBSTAGE
LOWERMOST EARLY CRETACEOUS EPOCH
EARLY CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: CHELONOMORPHA KUHN, 1960
SUPERCOHORT: TESTUNINATA KLEIN, 1760 (sensu JOYCE et al, 2004)
CLADE NAME: MESOCHELYDIA JOYCE, 2017
CLADE NAME: PERICHELYDIA JOYCE, 2017
Family: HELOCHELYDRIDAE Nopcsa, 1928
Helochelydra bakewelli
= Compsemys bakewelli
= Trionyx bakewelli
= Helochelydra bakewelli
= Tretosternon bakewelli
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ENGLAND
EAST SUSSEX COUNTY
HASTINGS
WADHURST CLAY
HASTINGS BEDS
(ASHDOWN BEDS, TUNBRIDGE WELLS SAND)
BERRIASIAN-VALANGINIAN STAGE
LOWER NEOCOMIAN SUBEPOCH
LOWER EARLY CRETACEOUS SUBEPOCH
EARLY CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
INFRACLASS: EURYAPSIDA COLBERT, 1945
SUPERORDER: SAUROPTERYGIA OWEN 1860
GRANDORDER: EOSAUROPTERYGIA emend RIEPPEL, 1994
ORDER: PLESIOSAURIA OWEN, 1840 NON BLAINVILLE, 1835
Incertae sedis
Gen. sp. indet.
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: CRUROTARSI SERENO & ARCUCCI, 1990
COHORT: CROCODYLOTARSI BENTON & CLARK, 1988
MAGNORDER: SUCHIA KREBS, 1974
SUPERORDER: CROCODYLOMORPHA WALKER, 1968
GRANDORDER: CROCODILIFORMES HAY, 1930
MICRORDER: MESOEUROCROCODYLIA WHETSTONE & WHYBROW, 1983
ORDER: MESOSUCHIA EMEND HUXLEY, 1875
PARVORDER: METASUCHIA BENTON & CLARK, 1988
INFRAORDER: METAMESOSUCHIA HULKE, 1878
NEOSUCHIA CLARK, IN BENTON & CLARK, 1988
Family: BERNISSARTIIDAE Dollo, 1883
Bernissartia fagesii
= *Crocodilus saulii
Bernissartia spFamily: ATOPOSAURIDAE Gervais, 1871
Theriosuchus pusillus
Theriosuchus spFamily: GONIOPHOLIDIDAE Cope, 1875
Incertae sedis
Goniopholis sp
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ENGLAND
EAST SUSSEX COUNTY
HASTINGS SAND
HASTINGS BEDS
(ASHDOWN BEDS, TUNBRIDGE WELLS SAND)
BERRIASIAN-VALANGINIAN STAGE
LOWER-MIDDLE NEOCOMIAN SUBEPOCH
LOWER EARLY CRETACEOUS EPOCH
EARLY CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: CRUROTARSI SERENO & ARCUCCI, 1990
COHORT: CROCODYLOTARSI BENTON & CLARK, 1988
MAGNORDER: SUCHIA KREBS, 1974
SUPERORDER: CROCODYLOMORPHA WALKER, 1968
GRANDORDER: CROCODILIFORMES HAY, 1930
MICRORDER: MESOEUROCROCODYLIA WHETSTONE & WHYBROW, 1983
ORDER: EUSUCHIA emend HUXLEY, 1875
Family: HYLAEOCHAMPSIDAE Andrews, 1913
Hylaeochampsa vectiana
= *Heterosuchus valdensis
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SOUTHERN UNITED KINGDOM
ENGLAND
EAST SUSSEX COUNTY
UNCERTAIN, BUT PRESUMABLY HASTINGS
UNCERTAIN, BUT PRESUMABLY HASTINGS GROUP
BERRIASIAN-VALANGINIAN STAGE
LOWER-MIDDLE NEOCOMIAN SUBEPOCH
LOWER EARLY CRETACEOUS EPOCH
EARLY CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
AVEPODA PAUL, 2002
SUPERORDER: NEOTHEROPODA BAKKER, 1986
GRANDORDER: TETANURAE GAUTHIER, 1986
ORDER: NEOTETANURAE SERENO, WILSON, LARSSON, DUTHEIL & SUES, 1994
SUBORDER: AVETHEROPODA PAUL, 1988
Superfamily: ALLOSAUROIDEA Marsh, 1878 (sensu Currie & Zhao, 1994)
Family: ALLOSAURIDAE Marsh, 1878
Incertae sedis
Gen. sp indet.
_____________________________________________________________________________________
ENGLAND
EAST SUSSEX COUNTY
HASTINGS
WADHURST CLAY
BERRIASIAN-VALANGINIAN STAGE
LOWER NEOCOMIAN SUBEPOCH
LOWER EARLY CRETACEOUS SUBEPOCH
EARLY CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
INFRACLASS: EURYAPSIDA COLBERT, 1945
SUPERORDER: SAUROPTERYGIA OWEN, 1860
GRANDORDER: EOSAUROPTERYGIA emend RIEPPEL, 1994
ORDER: PLESIOSAURIA OWEN, 1840 NON BLAINVILLE, 1835
SUBORDER: PLESIOSAURIA OWEN, 1840 NON BLAINVILLE, 1835
INFRAORDER: PLIOSAUROMORPHA NOVA
Superfamily: PLIOSAUROIDEA Seeley, 1874
Family: PLIOSAURIDAE Seeley, 1874
*Hastanects valdensis
= Cimoliasaurus valdensis
= Plesiosaurus valdensis
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SOUTHERN UNITED KINGDOM
ENGLAND
EAST SUSSEX COUNTY
ON THE COAST NEAR HASTINGS
HASTINGS BEDS GROUP
UPPER BERRIASIAN-VALANGININ STAGE
LOWER GALLIC SUBEPOCH
MIDDLE EARLY CRETACEOUS EPOCH
EARLY CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: ORNITHISCHIFORMES COOPER, 1985
SUPERORDER: SAUROPODOMORPHA HUENE, 1932
GRANDORDER: BRONTOSAURIA OLSHEVSKY, 1991 (emend)
MICRORDER: SAUROPODA MARSH, 1878
ORDER: EUSAUROPODA
SUBORDER: NEOSAUROPODA BONAPARTE, 1986
INFRAORDER: MACRONARIA WILSON & SERENO, 1998
Incertae sedis
*Haestasaurus becklesii
= Pelorosaurus becklesi
= Morosaurus becklesi
____________________________________________________________________________________
EAST SUSSEX COUNTY
HASTINGS
HASTINGS BEDS
BERRIASIAN-VALANGINIAN STAGE
LOWER NEOCOMIAN SUBSTAGE
LOWERMOST EARLY CRETACEOUS EPOCH
EARLY CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: CHELONOMORPHA KUHN, 1960
SUPERCOHORT: TESTUDINES LINNAEUS, 1758
COHORT: CASICHELYDIA emend GAFFNEY & MEYLAN, 1988
MAGANORDER: CRYPTODIRA GAFFNEY & MEYLAN, 1988
SUPERORDER: SELMACRYPTODIRA GAFFNEY, HUTCHISON, JENKINS & MEEKR, 1987
GRANDORDER: DAIOCRYPTODIRA GAFFNEY & MEYLAN, 1988
MICRORDER: BAENOIDEA WILLIAMS, 1960
Family: NEURANKYLIDAE Lambe, 1902
Compsemys bakewelli
= *Helochelydra bakewelli
= Trionyx bakewelli
= Helochelydra bakewelli
= Tretosternon bakewelliMICRORDER: EUCRYPTODIRA GAFFNEY, 1975
Family: PLESIOCHELYIDAE Rutimeyer, 1873
Plesiochelys brodiei
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: AVEMETATARSALIA BENTON, 1999
COHORT: PROLACERTIFORMES CAMP, 1945 NOVA
SUPERORDER: TAPINOPLATIA PETERS, 2000
GRANDORDER: CHARACIOPODA PETERS, 2000
MICRORDER: FENESTRASAURIA PETERS, 2000
ORDER: PTEROSAURIA KAUP, 1834
SUBORDER: EUPTERODACTYLOIDEA BENNETT, 1994
Superfamily: ORNITHOCHEIROIDEA Seeley, 1870 (sensu Seeley, 1891)
Family: ORNITHOCHEIRIDAE Seeley, 1870
Incertae sedis
Ornithocheirus sp
SUPERCOHORT: CRUROTARSI SERENO & ARCUCCI, 1990
COHORT: CROCODYLOTARSI BENTON & CLARK, 1988
MAGNORDER: SUCHIA KREBS, 1974
SUPERORDER: CROCODYLOMORPHA WALKER, 1968
GRANDORDER: CROCODILIFORMES HAY, 1930
MICRORDER: MESOEUROCROCODYLIA WHETSTONE & WHYBROW, 1983
ORDER: MESOSUCHIA EMEND HUXLEY, 1875
PARVORDER: METASUCHIA BENTON & CLARK, 1988
INFRAORDER: METAMESOSUCHIA HULKE, 1878
NEOSUCHIA CLARK, IN BENTON & CLARK, 1988
Family: GONIOPHOLIDIDAE Cope, 1875
Goniopholis crassidens
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
Incertae sedis
Gen. sp indet.
AVEPODA PAUL, 2002
SUPERORDER: NEOTHEROPODA BAKKER, 1986
GRANDORDER: TETANURAE GAUTHIER, 1986
ORDER: NEOTETANURAE SERENO, WILSON, LARSSON, DUTHEIL & SUES, 1994
SUBORDER: CARNOSAURIA HUENE, 1920
Superfamily: Incertae sedis
Family: MEGALOSAURIDAE Huxley, 1889
Incertae sedis
Megalosaurus sp
SUBORDER: AVETHEROPODA PAUL, 1988
Superfamily: ALLOSAUROIDEA Marsh, 1878 (sensu Currie & Zhao, 1994)
Family: ALLOSAURIDAE Marsh, 1878
Incertae sedis
Becklespinax altispinax
= Acrocanthosaurus altispinaxCOHORT: ORNITHISCHIFORMES COOPER, 1985
SUPERORDER: SAUROPODOMORPHA HUENE, 1932
GRANDORDER: BRONTOSAURIA OLSHEVSKY, 1991 (emend)
MICRORDER: SAUROPODA MARSH, 1878
ORDER: EUSAUROPODA
SUBORDER: NEOSAUROPODA BONAPARTE, 1986
INFRAORDER: MACRONARIA WILSON & SERENO, 1998
PARVORDER: TITANOSAURIFORMES SALGADO ET AL, 1997
Family: BRACHIOSAURIDAE Riggs, 1904
Subfamily: BRACHIOSAURINAE Riggs, 1904 (sensu Janensch, 1929)
Ornithopsis hulkei
= Pelorosaurus hulkeiPelorosaurus conybearei
= Cetiosaurus brevis
= Morosaurus brevisMAGNORDER: ORNITHISCHIA SEELEY, 1888
SUPERORDER: NEORNITHISCHIA COOPER, 1985
GRANDORDER: ORNITHOPODA MARSH, 1871
SUPERORDER: EURYOPDA SERENO, 1986
GRANDORDER: STEGOSAUROMORPHA COOPER, 1985 (EMEND)
ORDER: STEGOSAURIA MARSH, 1877
Incertae sedis
Gen. sp indet.
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ENGLAND
EAST SUSSEX COUNTY
ECCLESBOURNE
HASTINGS BEDS
BERRIASIAN-VALANGINIAN STAGE
LOWER-MIDDLE NEOCOMIAN SUBEPOCH
LOWER EARLY CRETACEOUS EPOCH
EARLY CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: CRUROTARSI SERENO & ARCUCCI, 1990
COHORT: CROCODYLOTARSI BENTON & CLARK, 1988
MAGNORDER: SUCHIA KREBS, 1974
SUPERORDER: CROCODYLOMORPHA WALKER, 1968
GRANDORDER: CROCODILIFORMES HAY, 1930
MICRORDER: MESOEUROCROCODYLIA WHETSTONE & WHYBROW, 1983
ORDER: MESOSUCHIA EMEND HUXLEY, 1875
PARVORDER: METASUCHIA BENTON & CLARK, 1988
INFRAORDER: METAMESOSUCHIA HULKE, 1878
NEOSUCHIA CLARK, IN BENTON & CLARK, 1988
Family: GONIOPHOLIDIDAE Cope, 1875
Goniopholis crassidens
Incertae sedis
Gen. sp. Indet.
_____________________________________________________________________________________
SOUTHERN UNITED KINGDOM
ENGLAND
EAST SUSSEX COUNTY
ECCLESBOURNE GLEN
HASTINGS BEDS
BERRIASIAN-VALANGINIAN STAGE
LOWER-MIDDLE NEOCOMIAN SUBEPOCH
LOWER EARLY CRETACEOUS EPOCH
EARLY CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: ORNITHISCHIFORMES COOPER, 1985
SUPERORDER: SAUROPODOMORPHA HUENE, 1932
GRANDORDER: BRONTOSAURIA OLSHEVSKY, 1991 (emend)
MICRORDER: SAUROPODA MARSH, 1878
ORDER: EUSAUROPODA
SUBORDER: NEOSAUROPODA BONAPARTE, 1986
INFRAORDER: HOMALOSAUROPIA JANENSCH, 1929
PARVORDER: DIPLODOCIMORPHA CALVO & SALGADO, 1995
Superfamily: DIPLODOCOIDEA Marsh, 1884 (sensu Upchurch, 1995)
Family: REBBACHISAURIDAE Bonaparte, 1996
*Xenoposeidon proneneukos
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