SPAIN
BASQUE COUNTY
DISUSED SAND QUARRY NEAR THE VILLAGE OF LANO
SOBREPENA FORMATION
LATEST CAMPANIAN – EARLIEST MAASTRICHTIAN STAGE
UPPER SENONIAN SUBEPOCH
UPPER UPPER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
____________________________________________________________________________________SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: AMPHIBIA LINNAEUS, 1758
SUBCLASS: LISSAMPHIBIA, HAECKEL, 1866?
ORDER: ALLOCAUDATA FOX & NAYLOR, 1982
Family: ALBANERPETONTIDAE Fox & Naylor, 1982
Cf. Albanerpeton sp
SUPERORDER: BATRACHIA BRONGNIART, 1800
ORDER: CAUDATA SCOPOLI, 1777
Incertae sedis
Gen. sp indet.
SUPERORDER: SALIENTIA LAURENTI, 1768
Incertae sedis
Gen. sp indet.
ORDER: ANURA RAFINESQUE, 1815
SUBORDER: DISCOGLOSSOIDEI SOKOL, 1977
Family: DISCOGLOSSIDAE Guenther, 1859
Incertae sedis
Gen. sp indet.
SUBORDER: MESOBATRACHIA LAURENT, 1979
Superfamily: PIPOIDEA Gray, 1825 (sensu Fitzinger, 1843)
Family: PALAEOBATRACHIDAE Cope, 1865
Gen. sp indet.
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: CHELONOMORPHA KUHN, 1960
SUPERCOHORT: TESTUDINES LINNAEUS, 1758
COHORT: CASICHELYDIA emend GAFFNEY & MEYLAN, 1988
MAGNORDER: PLEURODIRA GAFFNEY & MEYLAN, 1988
ORDER: PLEURODIRIDA COPE, 1864
Family: DORTOKIDAE Lapparent & Murelaga, 1997
*Dortoka vasconica
ORDER: PLEURODIRIDA COPE, 1864
SUBORDER: EUPLEURODIRA GAFFNEY & MEYLAN, 1988
Superfamily: PELOMEDUSOIDES Cope, 1868 Cope, 1868
Family: PELOMEDUSIDAE Cope, 1868
Subfamily: PELOMEDUSINAE Cope, 1868
Elochelys sp
*Polysternon atlanticum
MAGANORDER: CRYPTODIRA GAFFNEY & MEYLAN, 1988
ORDER: CRYPTODIRA COPE, 1868
Family: SOLEMYDIDAE Lapparent & Murelaga, 1997
*Solemys vermiculata
Solemys spSUBCLASS: DIAPSIDA OSBORN, 1904
INFRACLASS: NEODIAPSIDA BENTON, 1985
LEGION: LEPIDOSAUROMORPHA DUMERIL & BIBRON, 1839
SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)
MAGNORDER: SQUAMATA OPPEL, 1811
SUPERORDER: LACERTILIA OWEN, 1842
Incertae sedis
Gen. sp indet.
GRANDORDER: ASCALABOTA GILMORE, 1928
ORDER: IGUANIA LATREILLE, 1825
SUBORDER: DRACOMORPHA NORTHCUTT, 1978
INFRAORDER: IGUANOMORPHA FURBRINGER, 1900
Family: IGUANIDAE Gray, 1827
Gen. sp indet.
GRANDORDER: SCINCOGEKKONOMORPHA SUKHANOV, 1961
MICRORDER: SCINCOMORPHA CAMP, 1923
Incertae sedis
Gen. sp indet.
ORDER: LACERTOGNATHA EMEND ALIFANOV, 2000
PARVORDER: AMPHISBAENIA GRAY, 1844
Family: AMPHISBAENIDAE Gray, 1865
Gen. sp indet.
SUPERORDER: SERPENTES LINNAEUS, 1759
ORDER: OPHIDIA BRONGNIART, 1800
SUBORDER: ALETHINOPHIDIA NOPCSCA, 1923
Superfamily: BOOIDEA Gray, 1825
Family: BOIDAE Gray, 1825
Subfamily: MADTSOINAE Hoffstetter, 1961
*Herensugea caristiorum
*Madtsoia laurasiae
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: AVEMETATARSALIA BENTON, 1999
COHORT: PROLACERTIFORMES CAMP, 1945 NOVA
SUPERORDER: TAPINOPLATIA PETERS, 2000
GRANDORDER: CHARACIOPODA PETERS, 2000
MICRORDER: FENESTRASAURIA PETERS, 2000
ORDER: PTEROSAURIA KAUP, 1834
SUBORDER: EUPTERODACTYLOIDEA BENNETT, 1994
Superfamily: AZHDARCHIDEA Nesov, 1984
Family: AZHDARCHIDAE Nesov, 1984 emend Padian, 1986
Gen. sp indet.
SUPERCOHORT: CRUROTARSI SERENO & ARCUCCI, 1990
COHORT: CROCODYLOTARSI BENTON & CLARK, 1988
MAGNORDER: SUCHIA KREBS, 1974
SUPERORDER: CROCODYLOMORPHA WALKER, 1968
GRANDORDER: CROCODILIFORMES HAY, 1930
MICRORDER: MESOEUROCROCODYLIA WHETSTONE & WHYBROW, 1983
ORDER: MESOSUCHIA EMEND HUXLEY, 1875
PARVORDER: METASUCHIA BENTON & CLARK, 1988
INFRAORDER: METAMESOSUCHIA HULKE, 1878
NEOSUCHIA CLARK, IN BENTON & CLARK, 1988
Family: TREMATOCHAMPSIDAE Buffetaut, 1974
Incertae sedis
Gen. sp indet.
ORDER: EUSUCHIA emend HUXLEY, 1875
INFRAORDER: CROCODYLIA emend GMELIN, 1877
PARVORDER: BREVIROSTRES
Superfamily: ALLIGATORIDEA Buffetaut, 1985
GLOBIDONTA BROUCHU, 1999
Family: ALLIGATORIDAE Gray, 1844
*Acynodon iberoccitanus
*Musturzabalsuchus buffetauti
Genus sp indet.
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
Incertae sedis
Gen. sp indet.
(Small theropod)AVEPODA PAUL, 2002
SUPERORDER: NEOTHEROPODA BAKKER, 1986
GRANDORDER: CERATOSAURIA NOVAS, 1992
AVEROSTRA PAUL, 2001
ORDER: CERATOSAURIA MARSH, 1884
Superfamily: ABELISAUROIDEA Novas, 1989
Family: ABELISAURIDAE Bonaparte & Novas, 1985
Gen sp indet
COHORT: ORNITHISCHIFORMES COOPER, 1985
SUPERORDER: SAUROPODOMORPHA HUENE, 1932
GRANDORDER: BRONTOSAURIA OLSHEVSKY, 1991 (emend)
MICRORDER: SAUROPODA MARSH, 1878
ORDER: EUSAUROPODA
SUBORDER: NEOSAUROPODA BONAPARTE, 1986
INFRAORDER: MACRONARIA WILSON & SERENO, 1998
PARVORDER: TITANOSAURIFORMES SALGADO ET AL, 1997
Family: TITANOSAURIDAE Lydekker, 1885
Subfamily: TITANOSAURINAE Lydekker, 1885 (sensu Nopcsa, 1928)
*Lirainosaurus astibiae
MAGNORDER: ORNITHISCHIA SEELEY, 1888
SUPERORDER: NEORNITHISCHIA COOPER, 1985
GRANDORDER: ORNITHOPODA MARSH, 1871
MICRORDER: EUORNITHOPODA SERENO, 1986
ORDER: IGUANODONTIA DOLLO, 1888
Family: RHABDODONTIDAE Weishampel, Jainu, Csiki & Norman, 2003
Rhabdodon priscus
= Rhabdodon priscumSUPERORDER: EURYOPDA SERENO, 1986
GRANDORDER: ARMATOSAURIA ZHAO, 1983 (emend)
ORDER: ANKYLOSAURIA OSBORN, 1923
SUBORDER: SCELIDOSAURIA COOPER, 1985
INFRAORDER: ANKYLOSAURA OSBORN, 1923
Family: NODOSAURIDAE Marsh, 1890
Gen. sp indet.
CLASS: MAMMALIA LINNAEUS, 1758
SUBCLASS: PROTOTHERIA GILL, 1872
INFRACLASS: TRECHNOTHERIA McKENNA, 1975
LEGION: CLADOTHERIA McKENNA, 1975
SUBLEGION: ZATHERIA MCKENNA, 1975
INFRALEGION: TRIBOSPHENIDA (McKENNA, 1975) McKENNA & BELL, 1997
SUPERCOHORT: THERIA (PARKER & HASWELL, 1987) McKENNA & BELL, 1997
Incrtae sedis
Gen. sp indet.
COHORT: PLACENTALIA (OWEN, 1837) McKENNA & BELL, 1997
MAGNORDER: EPITHERIA (McKENNA, 1975) McKENNA & BELL, 1997
SUPERORDER: LEPTICTIDAE McKENNA, 1975
Incertae sedis
Labes? sp
*Lainodon orueetxebarriai
Lainodon orueetxebarriai or L. new species
Lainodon sp
SPAIN
BASQUE-CANTABRIC BASIN
BASQUE COUNTY
LANO
LATE CAMPANIAN-EARLY MAASTRICHTIAN STAGE
UPPERMOST SENONIAN
UPPERMOST GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
AVEPODA PAUL, 2002
SUPERORDER: NEOTHEROPODA BAKKER, 1986
GRANDORDER: CERATOSAURIA NOVAS, 1992
AVEROSTRA PAUL, 2001
ORDER: CERATOSAURIA MARSH, 1884
Superfamily: ABELISAUROIDEA Novas, 1989
Family: ABELISAURIDAE Bonaparte & Novas, 1985
Tarascosaurus sp
COHORT: ORNITHISCHIFORMES COOPER, 1985
MAGNORDER: ORNITHISCHIA SEELEY, 1888
SUPERORDER: NEORNITHISCHIA COOPER, 1985
GRANDORDER: ORNITHOPODA MARSH, 1871
SUPERORDER: EURYOPDA SERENO, 1986
GRANDORDER: ARMATOSAURIA ZHAO, 1983 (emend)
ORDER: ANKYLOSAURIA OSBORN, 1923
SUBORDER: SCELIDOSAURIA COOPER, 1985
INFRAORDER: ANKYLOSAURA OSBORN, 1923
Family: NODOSAURIDAE Marsh, 1890
Subfamily: STRUTHIOSAURINAE Nopcsa, 1929
Struthiosaurus austraicus
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SPAIN
IBERIAN PENINSULA
BASQUE PROVINCE
ALAVA
CASTILLO-LASARTE
VITORIA FORMATION
H. MARROTI ZONE
UPPERMOST CAMPANIAN STAGE
UPPER SENONIAN SUBEPOCH
UPPER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
INFRACLASS: NEODIAPSIDA BENTON, 1985
LEGION: LEPIDOSAUROMORPHA BENTON, 1983
SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)
MAGNORDER: SQUAMATA, OPPEL, 1811
SUPERORDER: LACERTILIA OWEN, 1842
GRANDORDER: SCINCOGEKKONOMORPHA SUKHANOV, 1961
MICRORDER: SCINCOMORPHA CAMP, 1923
ORDER: ANGUIMORPHA FURBRINGER, 1900
SUBORDER: DIPLOGLOSSA COPE, 1864
INFRAORDER: THECAGLOSSA COPE, 1890
PARVORDER: MONSTERSAURIA NORELL & GAO, 1977
Superfamily: VARANOIDEA Gray, 1827 (sensu Camp, 1923)
Family: MOSASAURIDAE Gervais, 1853
Infrafamily: NATANIA
Subfamily: TYLOSAURINAE Williston, 1897
Tylosaurus sp
= cf. Mosasaurus sp
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