UTAH
GRANDSTAIRCASE-ESCALANTE NATIONAL MONUMENT
UMNH LOCALTY 303
KAIPAROWITS FORMATION
CAMPANIAN STAGE
SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: CRUROTARSI SERENO & ARCUCCI, 1990
COHORT: CROCODYLOTARSI BENTON & CLARK, 1988
MAGNORDER: SUCHIA KREBS, 1974
SUPERORDER: CROCODYLOMORPHA WALKER, 1968
GRANDORDER: CROCODILIFORMES HAY, 1930
Incertae sedis
Gen. sp indet.
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: ORNITHISCHIFORMES COOPER, 1985
MAGNORDER: ORNITHISCHIA SEELEY, 1888
SUPERORDER: NEORNITHISCHIA COOPER, 1985
GRANDORDER: ORNITHOPODA MARSH, 1871
MICRORDER: EUORNITHOPODA SERENO, 1986
ORDER: HYPSILOPHODONTIA COOPER, 1983
Family: HYPSILOPHODONTIDAE Dollo, 1882
Gen. sp indet.
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EC UTAH
NESLEN FORMATION
CAMPANIAN STAGE
SENONIAN SUBEPOCH
LOWER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: ORNITHISCHIFORMES COOPER, 1985
MAGNORDER: ORNITHISCHIA SEELEY, 1888
SUPERORDER: NEORNITHISCHIA COOPER, 1985
GRANDORDER: ORNITHOPODA MARSH, 1871
MICRORDER: EUORNITHOPODA SERENO, 1986
ORDER: IGUANODONTIA DOLLO, 1888
SUBORDER: ANKYLOPOLLEXIA SERENO, 1986
INFRAORDER: STYRACOSTERNA SERENO, 1986
PARVORDER: HADROSAURIFORMES SERENO, 1997
Superfamily: HADROSAUROIDEA Cope, 1869 (sensu Sereno, 1986)
Family: HADROSAURIDAE COPE, 1869
Incertae sedis
Gen. sp indet.
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UTAH
KENILWORTH MINE
BLACKHAWK FORMATION
CAMPANIAN STAGE
SENONIAN SUBEPOCH
LOWER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
AVEPODA PAUL, 2002
SUPERORDER: NEOTHEROPODA BAKKER, 1986
GRANDORDER: TETANURAE GAUTHIER, 1986
MICRORDER: COELUROSAURIA HUENE, 1914
ORDER: MANIRAPTORFORMES HOLTZ, 1996
AVETHEROPODA PAUL, 2001
SUBORDER: COELURIA MARSH, 1881
Superfamily: TYRANNOSAUROIDEA Osborn, 1906 (sensu Bonaparte et. al., 1990)
Family: TYRANNOSAURIDAE Osborn, 1906
Subfamily: ALBERTOSAURINAE Currie, Hurum & Sabath, 2003
Albertosaurus sp
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UTAH
PAUNSAUGUNT PLATEAU
MNA LOCALITY 1071
DAKOTA FORMATION
AGE?
STAGE?
UPPER MIDDLE CAMPANIAN STAGE
UPPER MIDDLE SENONIAN SUBEPOCH
LOWER GULF EPOCH
MIDDLE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: MAMMALIA LINNAEUS, 1758
SUBCLASS: PROTOTHERIA GILL, 1872
INFRACLASS: TRECHNOTHERIA McKENNA, 1975
LEGION: CLADOTHERIA McKENNA, 1975
SUBLEGION: ZATHERIA MCKENNA, 1975
INFRALEGION: TRIBOSPHENIDA (McKENNA, 1975) McKENNA & BELL, 1997
SUPERCOHORT: THERIA (PARKER & HASWELL, 1987) McKENNA & BELL, 1997
Incertae sedis
Dakotadens morrowi
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UTAH
PAUNSAUGUNT PLETEAU
MNA LOCALITY 1073
UPPER KAIPAROWITS FORMATION
JUDITHIAN AGE
LATE CAMPANIAN STAGE
SENONIAN SUBEPOCH
UPPER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: MAMMALIA LINNAEUS, 1758
SUBCLASS: THERIIFORMES (ROWE, 1988) McKENNA & BELL, 1997
INFRACLASS: ALLOTHERIA (MARSH, 1880) McKENNA & BELL, 1997
ORDER: MULTITUBERCULATA COPE, 1884
SUBORDER: CIMOLODONTA McKENNA, 1975
Incertae sedis
*Cimexomys gregoryi
Paracimexomys nova
Superfamily: PTILODONTOIDEA Cope, 1887 (sensu McKenna & Bell, 1997)
Family: CIMOLODONTIDAE Marsh, 1889
Cimolodon sp. cf. C. nitidus
Cimolodon spFamily: PTILODONTIDAE Cope, 1887 (sensu McKenna & Bell, 1997)
Subfamily: NEOPLAGIAULACINAE Ameghino, 1890 (sensu McKenna & Bell, 1997)
Mesodma sp. cf. formosa
Mesodma sp. cf. hensleighi
Mesodma spSUBCLASS: PROTOTHERIA GILL, 1872
INFRACLASS: HOLOTHERIA (WIBEL, ROUGIER, NOVACEK, MCKENNA & DASHZEVEG, 1995) MCKENNA & BELL, 1997
SUPERLEGION: SYMMETRODONTA (SIMPSON, 1925) McKENNA & BELL, 1997
LEGION: SYMMETRODONTA SIMPSON, 1925
ORDER: SPALACOTHERIIDA PROTHERO, 1981
Family: SPALACOTHERIIDAE Marsh, 1887
Subfamily: SPALACOLESTINAE Cifelli & Madsen, 1999
Symmeterodontoides foxi
SUBCLASS: PROTOTHERIA GILL, 1872
INFRACLASS: TRECHNOTHERIA McKENNA, 1975
LEGION: CLADOTHERIA McKENNA, 1975
SUBLEGION: ZATHERIA MCKENNA, 1975
INFRALEGION: TRIBOSPHENIDA (McKENNA, 1975) McKENNA & BELL, 1997
SUPERCOHORT: THERIA (PARKER & HASWELL, 1987) McKENNA & BELL, 1997
COHORT: MARSUPIALIA (ILLIGER, 1811) McKENNA & BELL, 1997
MAGNORDER: AMERIDELPHIA (SZALAY, in ARCHER, ED, 1982) McKENNA & BELL, 1997
ORDER: DIDELPHIMORPHIA GILL, 1872
Family: DIDELPHIDAE Gray 1821
Turgidodon cf. T. russelli
= Alaphadon cf. A. russelli
Turgidodon spSubfamily: ALPHADONTIDAE Marshall, Case & Woodburne, 1990
Alphadon sp cf. A. marshi
= Alphadon sp cf. A. wilsoni
Alphadon sp. cf. A. attaragos
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UTAH
PAUNSAUGUNT PLATEAU
MNA LOCALITY 1074
UPPER KAIPRAROWITS FORMATION
JUDITHIAN AGE
LATE CAMPANIAN STAGE
SENONIAN SUBEPOCH
UPPER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: MAMMALIA LINNAEUS, 1758
SUBCLASS: PROTOTHERIA GILL, 1872
INFRACLASS: TRECHNOTHERIA McKENNA, 1975
LEGION: CLADOTHERIA McKENNA, 1975
SUBLEGION: ZATHERIA MCKENNA, 1975
INFRALEGION: TRIBOSPHENIDA (McKENNA, 1975) McKENNA & BELL, 1997
SUPERCOHORT: THERIA (PARKER & HASWELL, 1987) McKENNA & BELL, 1997
COHORT: MARSUPIALIA (ILLIGER, 1811) McKENNA & BELL, 1997
MAGNORDER: AMERIDELPHIA (SZALAY, in ARCHER, ED, 1982) McKENNA & BELL, 1997
ORDER: DIDELPHIMORPHIA GILL, 1872
Family: DIDELPHIDAE Gray 1821
Turgidodon sp cf. T. russelli
= Alaphadon sp. cf. T. russelliSubfamily: ALPHADONTIDAE Marshall, Case & Woodburne, 1990
Alphadon sp cf. A. marshi
= Alphadon sp cf. A. wilsoni
Alphadon sp cf. A. attaragos
____________________________________________________________________________________
UTAH
KAIPAROWITS FORMATION
JUDITHIAN AGE
LATE CAMPANIAN STAGE
SENONIAN SUBEPOCH
UPPER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: AVES LINNAEUS, 1758
SUBCLASS: SAURORNITHES NICHOLSON, 1879
AVEBREVICAUDA PAUL, 2001
INFRACLASS: ORNITHOTHORACES NOVA
LEGION: ORNITHURAE NOVA
COHORT: ENANTIORNITHES NOVA
SUPERORDER: EUENANTIORNITHES CHIAPPE, 2002
ORDER: ALEXORNITHIFORMES BRODKORB, 1976
Family: AVISAURIDAE Brett-Surman & Paul, 1985
Avisaurus new species
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UTAH
KAIPAROWITZ PLATEAU
KAIPAROWITZ FORMATION
JUDITHIAN AGE
LATE CAMPANIAN STAGE
SENONIAN SUBEPOCH
UPPER GULF EPOCH
LATE CRETACEOUS
VETEROVATA
OOSUPERCOHORT: DINOSAURIOVUM VIALOV, 1975
Incertae sedis
Gen. sp indet.
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UTAUTAH
OMNH V5
KAIPAROWITS FORMATION
JUDITHIAN AGE
LATE CAMPANIAN STAGE
SENONIAN SUBEPOCH
UPPER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: AMPHIBIA LINNAEUS, 1758
SUBCLASS: LISSAMPHIBIA, HAECKEL, 1866?
ORDER: ALLOCAUDATA FOX & NAYLOR, 1982
Family: ALBANERPETONTIDAE Fox & Naylor, 1982
Gen. sp indet.
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
INFRACLASS: NEODIAPSIDA BENTON, 1985
LEGION: LEPIDOSAUROMORPHA DUMERIL & BIBRON, 1839
SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)
MAGNORDER: SQUAMATA OPPEL, 1811
SUPERORDER: LACERTILIA OWEN, 1842
GRANDORDER: ASCALABOTA GILMORE, 1928
ORDER: IGUANIA LATREILLE, 1825
SUBORDER: ACRODONTIA COPE, 1864
INFRAORDER: AGAMOGNATHA EMEND ALIFANOV, 2000
PARVORDER: POLYGLYPHANODONTIA EMEND ALIFANOV, 2000
Family: POLYGLPHANODONTIDAE Gilmore, 1942
Peneteius saueri
= Manangyasaurus sauer
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UTAH
OMNH V6
KAIPAROWITS FORMATION
JUDITHIAN AGE
LATE CAMPANIAN STAGE
SENONIAN SUBEPOCH
UPPER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: AMPHIBIA LINNAEUS, 1758
SUBCLASS: LISSAMPHIBIA, HAECKEL, 1866?
ORDER: ALLOCAUDATA FOX & NAYLOR, 1982
Family: ALBANERPETONTIDAE Fox & Naylor, 1982
Albanerpeton nexuosus
Albanerpeton galaktion
Albanerpeton gracilisGen. sp indet
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
INFRACLASS: NEODIAPSIDA BENTON, 1985
LEGION: LEPIDOSAUROMORPHA DUMERIL & BIBRON, 1839
SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)
MAGNORDER: SQUAMATA OPPEL, 1811
SUPERORDER: LACERTILIA OWEN, 1842
GRANDORDER: ASCALABOTA GILMORE, 1928
ORDER: IGUANIA LATREILLE, 1825
SUBORDER: ACRODONTIA COPE, 1864
INFRAORDER: AGAMOGNATHA EMEND ALIFANOV, 2000
PARVORDER: POLYGLYPHANODONTIA EMEND ALIFANOV, 2000
Family: POLYGLPHANODONTIDAE Gilmore, 1942
Peneteius saueri
= *Manangyasaurus sauer
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UTAH
OMNH V61
KAIPAROWITS FORMATION
JUDITHIAN AGE
LATE CAMPANIAN STAGE
SENONIAN SUBEPOCH
UPPER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: AMPHIBIA LINNAEUS, 1758
SUBCLASS: LISSAMPHIBIA, HAECKEL, 1866?
ORDER: ALLOCAUDATA FOX & NAYLOR, 1982
Family: ALBANERPETONTIDAE Fox & Naylor, 1982
Albanerpeton nexuosus
Gen. sp indet.
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UTAH
PARIA RIVER
PARIA RIVER AMPHITHEATRE
KAIPAROWITS FORMATION
JUDITHIAN AGE
LATE CAMPANIAN STAGE
SENONIAN SUBEPOCH
UPPER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
AVEPODA PAUL, 2002
SUPERORDER: NEOTHEROPODA BAKKER, 1986
GRANDORDER: TETANURAE GAUTHIER, 1986
MICRORDER: COELUROSAURIA HUENE, 1914
ORDER: MANIRAPTORFORMES HOLTZ, 1996
SUBORDER: ORNITHOMIMOSAURIA BARSBOLD, 1976
Family: ORNITHOMIMIDAE Marsh, 1890
Ornithomimus antiquus
= Ornithomimus velox
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UTAH
PRICE RIVER COAL COMPANY ROOF
BLACKHAWK FORMATION
CAMPANIAN STAGE
SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
ICHNOSTEM: TETRAPODIPIDA NOVA
ICHNOSUPERCLASS: AMNIOTAIPIDII NOVA
ICHNOCLASS: REPTILIPEDIA VIALOV, 1966
ICHNOSUBCLASS: DIAPSIDIPIDII NOVA
ICHNOSTEM-GROUP: NEODIAPSIDIPEDII NOVA
ICHNOINFRACLASS: NEODIAPSIDIPEDII NOVA
ICHNOSUBDIVISION: ARCHOSAURIPEDIDII NOVA
ICHNOLEGION: DINOSAUROMORPHIPEDA NOVA
ICHNOSUBLEGION: DINOSAURIFORMEIPEDIDII NOVA
ICHNOSUPERCOHORT: DINOSAUROPEDIDA NOVA
ICHNOCOHORT: THEROPODIPEDIA NOVA
ICHNOSUPERORDER: NEOTHEROPODOPEDIDA NOVA
ICHNOORDER: NEOTETANURIPEDIDII NOVA
ICHNOSUBORDER: AVETHEROPODIPIDA NOVA
ICHNOMICRORDER: COELUROSAURIPIDIDA NOVA
Incertae sedis
Gen. sp indet.
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UTAH
PRICE RIVER COAL COMPANY ROOF
BLACKHAWK FORMATION
CAMPANIAN STAGE
SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
ICHNOSTEM: TETRAPODIPIDA NOVA
ICHNOSUPERCLASS: AMNIOTAIPIDII NOVA
ICHNOCLASS: REPTILIPEDIA VIALOV, 1966
ICHNOSUBCLASS: DIAPSIDIPIDII NOVA
ICHNOSTEM-GROUP: NEODIAPSIDIPEDII NOVA
ICHNOINFRACLASS: NEODIAPSIDIPEDII NOVA
ICHNOSUBDIVISION: ARCHOSAURIPEDIDII NOVA
ICHNOLEGION: DINOSAUROMORPHIPEDA NOVA
ICHNOSUBLEGION: DINOSAURIFORMEIPEDIDII NOVA
ICHNOSUPERCOHORT: DINOSAUROPEDIDA NOVA
ICHNOCOHORT: THEROPODIPEDIA NOVA
ICHNOSUPERORDER: NEOTHEROPODOPEDIDA NOVA
ICHNOORDER: MANIRAPTORFORMIPIDA NOVA
ICHNOSUBORDER: COELURIPIDA NOVA
Ichnosuperfamily: TYRANNOSAUROIPIDEA Nova
Ichnofamily: TYRANNOSAURIPODIDAE emend Vylov, 1988
Gen. sp indet.
ICHNOCOHORT: ORNITHISCHIFORMEIPIDA NOVA
ICHNOMAGNORDER: ORNITHISCHIPIDA NOVA
ICHNOSUPERORDER: NEORNITHISCHIPIDA NOVA
ICHNOGRANDORDER: ORNITHOPODIPDIA NOVA
ICHNOMICRORDER: EUORNITHOPODIPIDA NOVA
ICHNOORDER: IGUANODONTIPIDIA NOVA
ICHNOSUBORDER: ANKYLOPOLLEXIPEDIDA NOVA
ICHNOINFRAORDER: STYRACOSTERNIPIDA NOVA
ICHNOPARVORDER: HADROSAURIFORMIPEDIDA NOVA
ICHNOSUPERFAMILY: HADROSAUROIDIPEDIA NOVA
Ichnofamily: HADROSAURIOPDIDAE Vylov, 1988
Gen. sp indet.
ICHNOMICRORDER: MARGINOCEPHALIPIDA SERENO, 1986
INCHNOORDER: CERATOPIAIPIDA NOVA
ICHNOSUBORDER: NEOCERATOPIAIPIDA NOVA
ICHNOINFRAORDER: CORONOSAURIPIDA emend SERENO, 1986
ICHNOPARVORDER: CERATOPOMORPIDA NOVA
ICHNOORDER: CERATOPIPIDA MARSH, 1890
ICHNOSUBORDER: NEOCERATOPIPIDA NOVA
Ichnofamily: CERATOPOPIDII Nova
Ceratopsipes sp
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