NORTHERN
SUDAN
WADI MILK
FORMATION
?CENOMANIAN STAGE
GALLIC
SUBEPOCH
LOWER GULF
EPOCH
MIDDLE
CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: CRUROTARSI SERENO & ARCUCCI, 1990
COHORT: CROCODYLOTARSI BENTON & CLARK, 1988
MAGNORDER: SUCHIA KREBS, 1974
SUPERORDER: CROCODYLOMORPHA WALKER, 1968
GRANDORDER: CROCODILIFORMES HAY, 1930
MICRORDER: MESOEUROCROCODYLIA WHETSTONE & WHYBROW, 1983
PARVORDER: METASUCHIA BENTON & CLARK, 1988
GONDWANASUCHIA DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004
NOTOSUCHIMORPHA DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004
NOTOSUCHIFORMES DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004
TERRIASUCHIA DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004
PEIROSAURIMORPHA DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004
Family: CANDIDODONTIDAE de Souza Carvalho, Ribeiro & dos Santos Avilla, 2004
PEIROSAURIFORMES DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004
Superfamily: PEIROSAURIDEA Gasparini, 1982 emend de Souza Carvalho, Ribeiro & dos Santos Avilla, 2004
Family: PEIROSAURIDAE Gasparini, 1982
Gen. sp indet.
_____________________________________________________________________________________
NORTHERN SUDAN
ABOUT 200 KM NORTHWEST OF KHARTOUM
F1/89, F2/89
UPPERMOST ALBIAN-LOWERMOST CENOMANIAN STAGE
UPPER GALLIC SUBEPOCH
LOWEST GULF EPOCH
LATE EARLY TO EARLY MIDDLE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: CHELONOMORPHA KUHN, 1960
SUPERCOHORT: TESTUDINES LINNAEUS, 1758
INCERTAE SEDIS
Gen. sp indet.
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: CRUROTARSI SERENO & ARCUCCI, 1990
COHORT: CROCODYLOTARSI BENTON & CLARK, 1988
MAGNORDER: SUCHIA KREBS, 1974
SUPERORDER: CROCODYLOMORPHA WALKER, 1968
GRANDORDER: CROCODILIFORMES HAY, 1930
MICRORDER: MESOEUROCROCODYLIA WHETSTONE & WHYBROW, 1983
ORDER: MESOSUCHIA EMEND HUXLEY, 1875
PARVORDER: METASUCHIA BENTON & CLARK, 1988
INFRAORDER: TETHYSUCHIA BUFFETAUT, 1982
Family: DYROSAURIDAE de STEFANO, 1903
Gen. sp indet.
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
INCERTAE SEDIS
Gen. sp indet.
COHORT: ORNITHISCHIFORMES COOPER, 1985
SUPERORDER: SAUROPODOMORPHA HUENE, 1932
GRANDORDER: BRONTOSAURIA OLSHEVSKY, 1991 (emend)
MICRORDER: SAUROPODA MARSH, 1878
ORDER: EUSAUROPODA
SUBORDER: NEOSAUROPODA BONAPARTE, 1986
INFRAORDER: MACRONARIA WILSON & SERENO, 1998
PARVORDER: TITANOSAURIFORMES SALGADO ET AL, 1997
Superfamily: TITANOSAUROIDEA Lydekker, 1885 (sensu Upchurch, 1999)
Family: TITANOSAURIDAE Lydekker, 1885
Gen. sp indet.
N SUDAN
CENOMANIAN STAGE
UPPER GALIC SUBEPOCH
LOWER UPPER UPPER GULF EPOCH
MIDDLE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
____________________________________________________________________________________SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: CHELONOMORPHA KUHN, 1960
SUPERCOHORT: TESTUDINES LINNAEUS, 1758
Incertae sedis
Genus sp indet.
SUDAN
CENOMANIAN STAGE
UPPER GALLIC SUBEPOCH
LOWERMOST GULF EPOCH
MIDDLE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
____________________________________________________________________________________SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: CRUROTARSI SERENO & ARCUCCI, 1990
COHORT: CROCODYLOTARSI BENTON & CLARK, 1988
MAGNORDER: SUCHIA KREBS, 1974
SUPERORDER: CROCODYLOMORPHA WALKER, 1968
ORDER: MESOSUCHIA EMEND HUXLEY, 1875
SUBORDER: MESOEUROCROCODYLIA WHETSTONE & WHYBROW, 1983
Incertae sedis
Gen. sp indet.
SUDAN
WADI ABU HASHIM
SHENDI FORMATION
CENOMANIAN
MIDDLE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
____________________________________________________________________________________SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
ORDER: Incertae sedis
Gen. sp indet.
SUDAN
C. 200 KM NORTH-WEST OF KHARTOUM
WADI ABU HASHIM
HASHIM MEMBER
WADI MILK FORMATION
CENOMANIAN STAGE
UPPER GALIC SUBEPOCH
LOWER UPPER UPPER GULF EPOCH
MIDDLE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
____________________________________________________________________________________SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: AMPHIBIA LINNAEUS, 1758
SUBCLASS: LISSAMPHIBIA, HAECKEL, 1866?
ORDER: GYMNOPHIONA RAFINESQUE, 1814
Incertae sedis
Gen. sp indet.
SUPERORDER: BATRACHIA BRONGNIART, 1800
ORDER: CAUDATA SCOPOLI, 1777
Incertae sedis
Gen. sp indet.
SUBORDER: SALAMANDROIDEA NOBLE, 1931
Family: SIRENIDAE Gray, 1825
*Kababisha sudanensis
*Kababisha humarensis
Kababisha spSUPERORDER: SALIENTIA LAURENTI, 1768
Incertae sedis
Gen. sp indet.
ORDER: ANURA RAFINESQUE, 1815
SUBORDER: NEOBATRACHIA REIG, 1958
Superfamily: RANOIDEA Gray, 1825 (sensu Fitzinger, 1826)
Family: RANIDAE GRAY, 1825
Subfamly: RANINAE Gray, 1825
Rana sp
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: CHELONOMORPHA KUHN, 1960
SUPERCOHORT: TESTUDINES LINNAEUS, 1758
COHORT: CASICHELYDIA emend GAFFNEY & MEYLAN, 1988
MAGNORDER: PLEURODIRA GAFFNEY & MEYLAN, 1988
ORDER: PLEURODIRIDA COPE, 1864
SUBORDER: EUPLEURODIRA GAFFNEY & MEYLAN, 1988
Superfamily: PELOMEDUSOIDES Cope, 1868 Cope, 1868
Family: PELOMEDUSIDAE Cope, 1868
Gen. sp indet.
SUDAN
WADI ABU HASHIM
WADI MILK FORMATION
CENOMANIAN STAGE
UPPER GALLIC SUBEPOCH
LOWERMOST GULF EPOCH
MIDDLE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
INFRACLASS: NEODIAPSIDA BENTON, 1985
LEGION: LEPIDOSAUROMORPHA BENTON, 1983
SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)
MAGNORDER: SQUAMATA, OPPEL, 1811
SUPERORDER: SERPENTES LINNAEUS, 1759
ORDER: OPHIDIA BRONGNIART, 1800
SUBORDER: ALETHINOPHIDIA NOPCSCA, 1923
Superfamily: ACROCHORDOIDEA Bonaparte, 1838
Family: PALAEOPHIDAE LYDEKKER, 1888
Gen. sp indet.
____________________________________________________________________________________LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: CRUROTARSI SERENO & ARCUCCI, 1990
COHORT: CROCODYLOTARSI BENTON & CLARK, 1988
MAGNORDER: SUCHIA KREBS, 1974
SUPERORDER: CROCODYLOMORPHA WALKER, 1968
GRANDORDER: CROCODILIFORMES HAY, 1930
MICRORDER: MESOEUROCROCODYLIA WHETSTONE & WHYBROW, 1983
Incertae sedis
Gen. sp indet
ORDER: THALATTOSUCHIA EMEND FRAAS, 1901
Family: TELEOSAURIDAE Geoffroy, 1831
Machimosaurus sp
ORDER: MESOSUCHIA EMEND HUXLEY, 1875
PARVORDER: METASUCHIA BENTON & CLARK, 1988
INFRAORDER: METAMESOSUCHIA HULKE, 1878
NEOSUCHIA CLARK, IN BENTON & CLARK, 1988
Family: TREMATOCHAMPSIDAE Buffetaut, 1974
Incertae sedis
Gen. sp indet.
INFRAORDER: TETHYSUCHIA BUFFETAUT, 1982
Family: DYROSAURIDAE de STEFANO, 1903
Incertae sedis
Gen. sp indet.
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
Incertae sedis
Gen. sp indet.
COHORT: THEROPODA EMEND MARSH, 1881
ORDER: Incertae sedis
Gen. sp indet.
AVEPODA PAUL, 2002
SUPERORDER: NEOTHEROPODA BAKKER, 1986
GRANDORDER: TETANURAE GAUTHIER, 1986
ORDER: NEOTETANURAE SERENO, WILSON, LARSSON, DUTHEIL &
SUBORDER: AVETHEROPODA PAUL, 1988
Superfamily: ALLOSAUROIDEA Marsh, 1878 (sensu Currie & Zhao, 1994)
Family: CARCHARODONTOSAURIDAE Stromer, 1931
Gen. sp indet.
GRANDORDER: TETANURAE GAUTHIER, 1986
MICRORDER: COELUROSAURIA HUENE, 1914
ORDER: MANIRAPTORFORMES HOLTZ, 1996
AVETHEROPOA PAUL, 2002
SUBORDER: COELURIA MARSH, 1881
Incertae sedis
Gen. sp indet.
AVEPECTORA PAUL, 2002
SUBORDER: ORNITHOMIMOSAURIA BARSBOLD, 1976
Family: ORNITHOMIMIDAE Marsh, 1890
Incertae sedis
Gen. sp indet.
SUBORDER: MANIRAPTORA GAUTHIER, 1986
INFRAORDER: PARVES SERENO, 1997
PARVORDER: DEINONYCHOSAURIA COLBERT & D. A. RUSSELL, 1969
Family: DROMAEOSAURIDAE Russell, 1969
Incerate sedis
Gen. sp indet.
COHORT: ORNITHISCHIFORMES COOPER, 1985
SUPERORDER: SAUROPODOMORPHA HUENE, 1932
Incertae sedis
Gen. sp indet.
GRANDORDER: BRONTOSAURIA OLSHEVSKY, 1991 (emend)
MICRORDER: SAUROPODA MARSH, 1878
ORDER: EUSAUROPODA
SUBORDER: NEOSAUROPODA BONAPARTE, 1986
INFRAORDER: HOMALOSAUROPIA JANENSCH, 1929
PARVORDER: DIPLODOCIMORPHA CALVO & SALGADO, 1995
Superfamily: DIPLODOCOIDEA Upchurch, 1995
Family: DIPLODOCIDAE Marsh, 1884
Subfamily: DICRAEOSAURINAE Janensch, 1929
Gen. sp indet.
INFRAORDER: MACRONARIA WILSON & SERENO, 1998
PARVORDER: TITANOSAURIFORMES SALGADO ET AL, 1997
Superfamily: TITANOSAUROIDEA Lydekker, 1885 (sensu Upchurch, 1999)
Family: TITANOSAURIDAE Lydekker, 1885
Subfamily: TITANOSAURINAE Lydekker, 1885 (sensu Nopcsa, 1928)
Gen. sp indet.
MAGNORDER: ORNITHISCHIA SEELEY, 1888
Incertae sedis
Gen. sp indet.
SUPERORDER: NEORNITHISCHIA COOPER, 1985
GRANDORDER: ORNITHOPODA MARSH, 1871
MICRORDER: EUORNITHOPODA SERENO, 1986
ORDER: HYPSILOPHODONTIA COOPER, 1983
Family: HYPSILOPHODONTIDAE Dollo, 1882
Incertae sedis? Or new families? (pending further research)
Gen. sp indet.
ORDER: IGUANODONTIA DOLLO, 1888
SUBORDER: ANKYLOPOLLEXIA SERENO, 1986
INFRAORDER: STYRACOSTERNA SERENO, 1986
PARVORDER: HADROSAURIFORMES SERENO, 1997
Superfamily: IGUANODONTIDEA Cope, 1869 (sensu Sereno, 1986)
Family: IGUANODONTIDAE Cope, 1869
Incertae sedis
Gen. sp indet.
Superfamily: OURANOSAURIDEA Nova
Family: OURANOSAURIDAE nova
cf. Ouranosaurus sp
SUDAN
WADI ABU HASHIM
200 KM NW OF KHARTOUM
WADI ABU HASHIM MEMBER
WADI MILK FORMATION
CENOMANIAN STAGE
UPPERMIDDLE GALLIC SUBEPOCH
LOWERMOST GULF EPOCH
LATE MIDDLE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
INFRACLASS: NEODIAPSIDA BENTON, 1985
LEGION: LEPIDOSAUROMORPHA BENTON, 1983
SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)
MAGNORDER: SQUAMATA, OPPEL, 1811
SUPERORDER: SERPENTES LINNAEUS, 1759
Incertae sedis
Gen. sp indet.
ORDER: OPHIDIA BRONGNIART, 1800
SUBORDER: ALETHINOPHIDIA NOPCSCA, 1923
Superfamily: SIMOLIOPHEOIDEA Nopcsa, 1925
Family: LAPPARENTOPHEIDAE Hoffstetter, 1968
Genus a
Genus b
Superfamily: ANILIODEA Fitzinger, 1826
Family: ANILIIDAE Fitzenger, 1826
*Coniophis dabiebus
Coniophis cf. C. dabiebusSuperfamily: BOOIDEA Gray, 1825
Family: BOIDAE Gray, 1825
Subfamily: MADTSOINAE Hoffstetter, 1961
Gen. sp indet.
Superfamily: ACROCHORDOIDEA Bonaparte, 1838
Family: ?NIGEROPHEIDAE Rage, 1975
*Nubianophis afaahus
Nubianophis cf. N. afaahusFamily: PALAEOPHIDAE LYDEKKER, 1888
Gen. sp indet.
Superfamily: COLUBROIDEA Oppel, 1811
Incerte sedis
Gen. sp indet.
Family: RUSSELLOPHEIDAE Rage, 1978
*Krebsophis thobanus
____________________________________________________________________________________SUPERORDER: LACERTILIA OWEN, 1842
Incertae sedis
Gen. sp indet.
SUDAN
WADI ABU HASHIM
SHENDI FORMATION
CENOMANIAN STAGE
UPPERMIDDLE GALLIC SUBEPOCH
LOWERMOST GULF EPOCH
MIDDLE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
ORDER: Incertae sedis
Gen. sp indet.
____________________________________________________________________________________