UZBEKISTAN
KARAKALPAKISTAN
SOUTHWESTERN KYXYL KUM
SERIES OF STEEP SLOPES NORTH OF DRY LAKE KHODZHAKUL
KHODZHAKUL
LOWER OR MIDDLE PART OF THE KHODZHAKUL FORMATION
LOWERMOST CENOMANIAN STAGE
UPPER GALLIC SUBEPOCH
LOWEST GULF EPOCH
LATE EARLY TO EARLY MIDDLE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: AVEMETATARSALIA BENTON, 1999
COHORT: PROLACERTIFORMES CAMP, 1945 NOVA
SUPERORDER: TAPINOPLATIA PETERS, 2000
GRANDORDER: CHARACIOPODA PETERS, 2000
MICRORDER: FENESTRASAURIA PETERS, 2000
ORDER: PTEROSAURIA KAUP, 1834
SUBORDER: EUPTERODACTYLOIDEA BENNETT, 1994
Superfamily: ORNITHOCHEIROIDEA Seeley, 1870 (sensu Seeley, 1891)
Family: ORNITHOCHEIRIDAE Seeley, 1870
Incertae sedis
Gen. sp indet.
_____________________________________________________________________________________
UZBEKISTAN
KARAKALPAKISTAN
SOUTHWESTERN KYXYL KUM
A LARGE RAVINE APPROXIMATELY 1.5 KM NORTHEAST OF THE SHEIKHDZHEILI LOCALITY
KHODZHAKULSAI
UPPER PART OF THE KHODZHAKUL FORMATION
LOWERMOST CENOMANIAN STAGE
UPPER GALLIC SUBEPOCH
LOWEST GULF EPOCH
LATE EARLY TO EARLY MIDDLE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: AVEMETATARSALIA BENTON, 1999
COHORT: PROLACERTIFORMES CAMP, 1945 NOVA
SUPERORDER: TAPINOPLATIA PETERS, 2000
GRANDORDER: CHARACIOPODA PETERS, 2000
MICRORDER: FENESTRASAURIA PETERS, 2000
ORDER: PTEROSAURIA KAUP, 1834
SUBORDER: EUPTERODACTYLOIDEA BENNETT, 1994
Superfamily: ORNITHOCHEIROIDEA Seeley, 1870 (sensu Seeley, 1891)
Family: ORNITHOCHEIRIDAE Seeley, 1870
Incertae sedis
Gen. sp indet.
_____________________________________________________________________________________
UZBEKISTAN
KARAKALPAKISTAN
SOUTHWESTERN KYXYL KUM
A RESIDUAL MOUNTAIN NEAR THE RIGHT BANK OF THE AMU DARYA
CHELPYK
UPPER PART OF THE KHODZHAKUL FORMATION
LOWERMOST CENOMANIAN STAGE
UPPER GALLIC SUBEPOCH
LOWEST GULF EPOCH
LATE EARLY TO EARLY MIDDLE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: AVEMETATARSALIA BENTON, 1999
COHORT: PROLACERTIFORMES CAMP, 1945 NOVA
SUPERORDER: TAPINOPLATIA PETERS, 2000
GRANDORDER: CHARACIOPODA PETERS, 2000
MICRORDER: FENESTRASAURIA PETERS, 2000
ORDER: PTEROSAURIA KAUP, 1834
SUBORDER: EUPTERODACTYLOIDEA BENNETT, 1994
Superfamily: ORNITHOCHEIROIDEA Seeley, 1870 (sensu Seeley, 1891)
Family: ORNITHOCHEIRIDAE Seeley, 1870
Incertae sedis
Gen. sp indet.
_____________________________________________________________________________________
UZBEKISTAN
KARAKALPAKISTAN
SOUTHWESTERN KYXYL KUM
A LARGE GULLY IN THE EATERN MARGINAL AREA OF THE SULTANUIZDAG (= ZULTANUVAIS) MOUNTAINS
KARACHADALYSAI
UPPER PART OF THE KHODZHAKUL FORMATION
LOWERMOST CENOMANIAN STAGE
UPPER GALLIC SUBEPOCH
LOWEST GULF EPOCH
LATE EARLY TO EARLY MIDDLE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: AVEMETATARSALIA BENTON, 1999
COHORT: PROLACERTIFORMES CAMP, 1945 NOVA
SUPERORDER: TAPINOPLATIA PETERS, 2000
GRANDORDER: CHARACIOPODA PETERS, 2000
MICRORDER: FENESTRASAURIA PETERS, 2000
ORDER: PTEROSAURIA KAUP, 1834
SUBORDER: EUPTERODACTYLOIDEA BENNETT, 1994
Superfamily: ORNITHOCHEIROIDEA Seeley, 1870 (sensu Seeley, 1891)
Family: ORNITHOCHEIRIDAE Seeley, 1870
Incertae sedis
Gen. sp indet.
_____________________________________________________________________________________
UZBEKISTAN
KARAKALPAKISTAN
SOUTHWESTERN KIZYLKUM DESERT AREA
UNKNOWN
KHODZHAKUL FORMATION
EARLY CENOMANIAN STAGE
LOWER LATE CRETACEOUS EPOCH
LOWER MIDDLE CRETACEOUS
MIDDLE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: CHELONOMORPHA KUHN, 1960
SUPERCOHORT: TESTUDINES LINNAEUS, 1758
COHORT: CASICHELYDIA emend GAFFNEY & MEYLAN, 1988
MAGANORDER: CRYPTODIRA GAFFNEY & MEYLAN, 1988
SUPERORDER: SELMACRYPTODIRA GAFFNEY, HUTCHISON, JENKINS & MEEKR, 1987
GRANDORDER: DAIOCRYPTODIRA GAFFNEY & MEYLAN, 1988
MICRORDER: CENTROCRYPTODIRA GAFFNEY & MEYLAN, 1988
ORDER: POLYCRPTODIRA GAFFNEY & MEYLAN, 1988
SUBORDER: PROCOELOCRYPTODIRA GAFFNEY & MEYLAN, 1988
INFRAORDER: CHELOMACRYPTODIRA GAFFNEY & MEYLAN, 1988
PARVORDER: TRIONYCHIDA GRAY, 1870
Superfamily: TRIONYCHIDEA Bell, 1828 (sensu Gray, 1870)
Family: TRIONYCHIDAE Bell, 1828 sa
Gen. sp. indet.
Subfamily: TRIONYCHINAE Bell, 1828 (sensu Lydekker, 1889)
Tribe: TRIONYCHINI Bell, 1828 (sensu Gaffney & Meylan, 1988)
Trionyx dissolutus
____________________________________________________________________________________
UZBEKISTAN
KARAKALPAKISTAN
SOUTHWESTERN KIZYLKUM DESERT AREA
SKH-"V"
KHODZHAKUL FORMATION
EARLY CENOMANIAN STAGE
LOWER LATE CRETACEOUS EPOCH
LOWER MIDDLE CRETACEOUS
MIDDLE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: CHELONOMORPHA KUHN, 1960
SUPERCOHORT: TESTUDINES LINNAEUS, 1758
COHORT: CASICHELYDIA emend GAFFNEY & MEYLAN, 1988
MAGANORDER: CRYPTODIRA GAFFNEY & MEYLAN, 1988
SUPERORDER: SELMACRYPTODIRA GAFFNEY, HUTCHISON, JENKINS & MEEKR, 1987
GRANDORDER: DAIOCRYPTODIRA GAFFNEY & MEYLAN, 1988
MICRORDER: CENTROCRYPTODIRA GAFFNEY & MEYLAN, 1988
ORDER: POLYCRPTODIRA GAFFNEY & MEYLAN, 1988
SUBORDER: PROCOELOCRYPTODIRA GAFFNEY & MEYLAN, 1988
INFRAORDER: CHELOMACRYPTODIRA GAFFNEY & MEYLAN, 1988
PARVORDER: TRIONYCHIDA GRAY, 1870
Superfamily: TRIONYCHIDEA Bell, 1828 (sensu Gray, 1870)
Family: TRIONYCHIDAE Bell, 1828
Gen. sp indet.
____________________________________________________________________________________
UZBEKISTAN
KARAKALPAKIAN AUTONOMOUS REPUBLIC
CHELPYK
SCH-1
LOWER OR MIDDLE PART OF KHODZHAKULIAN SUITE
CENOMANIAN STAGE
MIDDLE GALLIC SUBEPOCH
LOWERMOST GULF EPOCH
MIDDLE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: CHELONOMORPHA KUHN, 1960
SUPERCOHORT: TESTUDINES LINNAEUS, 1758
COHORT: CASICHELYDIA emend GAFFNEY & MEYLAN, 1988
MAGANORDER: CRYPTODIRA GAFFNEY & MEYLAN, 1988
SUPERORDER: SELMACRYPTODIRA GAFFNEY, HUTCHISON, JENKINS & MEEKR, 1987
GRANDORDER: DAIOCRYPTODIRA GAFFNEY & MEYLAN, 1988
MICRORDER: CENTROCRYPTODIRA GAFFNEY & MEYLAN, 1988
ORDER: POLYCRPTODIRA GAFFNEY & MEYLAN, 1988
SUBORDER: PROCOELOCRYPTODIRA GAFFNEY & MEYLAN, 1988
INFRAORDER: CHELOMACRYPTODIRA GAFFNEY & MEYLAN, 1988
PARVORDER: TRIONYCHIDA GRAY, 1870
Superfamily: TRIONYCHIDEA Bell, 1828 (sensu Gray, 1870)
Family: TRIONYCHIDAE Bell, 1828
Subfamily: TRIONYCHINAE Bell, 1828 (sensu Lydekker, 1889)
Tribe: TRIONYCHINI Bell, 1828 (sensu Gaffney & Meylan, 1988)
Trionyx kyrgyzensis
Family: CARETTOCHELYIDAE Boulenger, 1887
Subfamily: ANOSTERINAE Lydekker, 1889
Kizylkumemys sp
_____________________________________________________________________________________
UZBEKISTAN
KARAKALPAKISTAN
SOUTHWESTERN KIZYLKUM DESERT AREA
KHODZHAKUL II
SKH-4
KHODZHAKUL FORMATION
EARLY CENOMANIAN STAGE
LOWER LATE CRETACEOUS EPOCH
LOWER MIDDLE CRETACEOUS
MIDDLE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: CHELONOMORPHA KUHN, 1960
SUPERCOHORT: TESTUDINES LINNAEUS, 1758
COHORT: CASICHELYDIA emend GAFFNEY & MEYLAN, 1988
MAGANORDER: CRYPTODIRA GAFFNEY & MEYLAN, 1988
SUPERORDER: SELMACRYPTODIRA GAFFNEY, HUTCHISON, JENKINS & MEEKR, 1987
GRANDORDER: DAIOCRYPTODIRA GAFFNEY & MEYLAN, 1988
MICRORDER: CENTROCRYPTODIRA GAFFNEY & MEYLAN, 1988
ORDER: POLYCRPTODIRA GAFFNEY & MEYLAN, 1988
SUBORDER: PROCOELOCRYPTODIRA GAFFNEY & MEYLAN, 1988
INFRAORDER: CHELOMACRYPTODIRA GAFFNEY & MEYLAN, 1988
PARVORDER: TRIONYCHIDA GRAY, 1870
Superfamily: TRIONYCHIDEA Bell, 1828 (sensu Gray, 1870)
Family: TRIONYCHIDAE Bell, 1828
Subfamily: TRIONYCHINAE Bell, 1828 (sensu Lydekker, 1889)
Tribe: TRIONYCHINI Bell, 1828 (sensu Gaffney & Meylan, 1988)
Trionyx kyrgyzensis Trionyx dissolutus
____________________________________________________________________________________
UZBEKISTAN
KARAKALPAKISTAN
SOUTHWESTERN KIZYLKUM DESERT AREA
KHODZHAKUL II
SKH-4
KHODZHAKUL FORMATION
EARLY CENOMANIAN STAGE
LOWER LATE CRETACEOUS EPOCH
LOWER MIDDLE CRETACEOUS
MIDDLE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: CHELONOMORPHA KUHN, 1960
SUPERCOHORT: TESTUDINES LINNAEUS, 1758
COHORT: CASICHELYDIA emend GAFFNEY & MEYLAN, 1988
MAGANORDER: CRYPTODIRA GAFFNEY & MEYLAN, 1988
SUPERORDER: SELMACRYPTODIRA GAFFNEY, HUTCHISON, JENKINS & MEEKR, 1987
GRANDORDER: DAIOCRYPTODIRA GAFFNEY & MEYLAN, 1988
MICRORDER: CENTROCRYPTODIRA GAFFNEY & MEYLAN, 1988
ORDER: POLYCRPTODIRA GAFFNEY & MEYLAN, 1988
SUBORDER: PROCOELOCRYPTODIRA GAFFNEY & MEYLAN, 1988
INFRAORDER: CHELOMACRYPTODIRA GAFFNEY & MEYLAN, 1988
PARVORDER: TRIONYCHIDA GRAY, 1870
Superfamily: TRIONYCHIDEA Bell, 1828 (sensu Gray, 1870)
Family: TRIONYCHIDAE Bell, 1828
Subfamily: TRIONYCHINAE Bell, 1828 (sensu Lydekker, 1889)
Tribe: TRIONYCHINI Bell, 1828 (sensu Gaffney & Meylan, 1988)
Trionyx dissolutus
____________________________________________________________________________________
UZBEKISTAN
KARAKALPAKISTAN
SOUTHWESTERN KIZYLKUM DESERT AREA
KHODZHAKUL II
SKH-26
KHODZHAKUL FORMATION
EARLY CENOMANIAN STAGE
LOWER LATE CRETACEOUS EPOCH
LOWER MIDDLE CRETACEOUS
MIDDLE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: CHELONOMORPHA KUHN, 1960
SUPERCOHORT: TESTUDINES LINNAEUS, 1758
COHORT: CASICHELYDIA emend GAFFNEY & MEYLAN, 1988
MAGANORDER: CRYPTODIRA GAFFNEY & MEYLAN, 1988
SUPERORDER: SELMACRYPTODIRA GAFFNEY, HUTCHISON, JENKINS & MEEKR, 1987
GRANDORDER: DAIOCRYPTODIRA GAFFNEY & MEYLAN, 1988
MICRORDER: CENTROCRYPTODIRA GAFFNEY & MEYLAN, 1988
ORDER: POLYCRPTODIRA GAFFNEY & MEYLAN, 1988
SUBORDER: PROCOELOCRYPTODIRA GAFFNEY & MEYLAN, 1988
INFRAORDER: CHELOMACRYPTODIRA GAFFNEY & MEYLAN, 1988
PARVORDER: TRIONYCHIDA GRAY, 1870
Superfamily: TRIONYCHIDEA Bell, 1828 (sensu Gray, 1870)
Family: TRIONYCHIDAE Bell, 1828
Subfamily: TRIONYCHINAE Bell, 1828 (sensu Lydekker, 1889)
Tribe: TRIONYCHINI Bell, 1828 (sensu Gaffney & Meylan, 1988)
Trionyx kyrgyzensis *Trionyx dissolutus
____________________________________________________________________________________
UZBEKISTAN
KARAKALPAKISTAN
SOUTHWESTERN KIZYLKUM DESERT AREA
KHODZHAKUL II
SKH-25
KHODZHAKUL FORMATION
EARLY CENOMANIAN STAGE
LOWER LATE CRETACEOUS EPOCH
LOWER MIDDLE CRETACEOUS
MIDDLE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: CHELONOMORPHA KUHN, 1960
SUPERCOHORT: TESTUDINES LINNAEUS, 1758
COHORT: CASICHELYDIA emend GAFFNEY & MEYLAN, 1988
MAGANORDER: CRYPTODIRA GAFFNEY & MEYLAN, 1988
SUPERORDER: SELMACRYPTODIRA GAFFNEY, HUTCHISON, JENKINS & MEEKR, 1987
GRANDORDER: DAIOCRYPTODIRA GAFFNEY & MEYLAN, 1988
MICRORDER: CENTROCRYPTODIRA GAFFNEY & MEYLAN, 1988
ORDER: POLYCRPTODIRA GAFFNEY & MEYLAN, 1988
SUBORDER: PROCOELOCRYPTODIRA GAFFNEY & MEYLAN, 1988
INFRAORDER: CHELOMACRYPTODIRA GAFFNEY & MEYLAN, 1988
PARVORDER: TRIONYCHIDA GRAY, 1870
Superfamily: TRIONYCHIDEA Bell, 1828 (sensu Gray, 1870)
Family: TRIONYCHIDAE Bell, 1828
Gen. sp indet.
Subfamily: TRIONYCHINAE Bell, 1828 (sensu Lydekker, 1889)
Tribe: TRIONYCHINI Bell, 1828 (sensu Gaffney & Meylan, 1988)
Trionyx kyrgyzensis Trionyx dissolutus
____________________________________________________________________________________
UZBEKISTAN
KARAKALPAKISTAN
SOUTHWESTERN KIZYLKUM DESERT AREA
KHODZHAKUL II
SSHD-8
KHODZHAKUL FORMATION
EARLY CENOMANIAN STAGE
LOWER LATE CRETACEOUS EPOCH
LOWER MIDDLE CRETACEOUS
MIDDLE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: CHELONOMORPHA KUHN, 1960
SUPERCOHORT: TESTUDINES LINNAEUS, 1758
COHORT: CASICHELYDIA emend GAFFNEY & MEYLAN, 1988
MAGANORDER: CRYPTODIRA GAFFNEY & MEYLAN, 1988
SUPERORDER: SELMACRYPTODIRA GAFFNEY, HUTCHISON, JENKINS & MEEKR, 1987
GRANDORDER: DAIOCRYPTODIRA GAFFNEY & MEYLAN, 1988
MICRORDER: CENTROCRYPTODIRA GAFFNEY & MEYLAN, 1988
ORDER: POLYCRPTODIRA GAFFNEY & MEYLAN, 1988
SUBORDER: PROCOELOCRYPTODIRA GAFFNEY & MEYLAN, 1988
INFRAORDER: CHELOMACRYPTODIRA GAFFNEY & MEYLAN, 1988
PARVORDER: TRIONYCHIDA GRAY, 1870
Superfamily: TRIONYCHIDEA Bell, 1828 (sensu Gray, 1870)
Family: TRIONYCHIDAE Bell, 1828
Gen. sp indet.
Subfamily: TRIONYCHINAE Bell, 1828 (sensu Lydekker, 1889)
Tribe: TRIONYCHINI Bell, 1828 (sensu Gaffney & Meylan, 1988)
Trionyx kyrgyzensis Trionyx dissolutus
____________________________________________________________________________________
UZBEKISTAN
KARAKALPAKISTAN
SOUTHWESTERN KIZYLKUM DESERT AREA
KHODZHAKUL II
SSHD-8a
KHODZHAKUL FORMATION
EARLY CENOMANIAN STAGE
LOWER LATE CRETACEOUS EPOCH
LOWER MIDDLE CRETACEOUS
MIDDLE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: CHELONOMORPHA KUHN, 1960
SUPERCOHORT: TESTUDINES LINNAEUS, 1758
COHORT: CASICHELYDIA emend GAFFNEY & MEYLAN, 1988
MAGANORDER: CRYPTODIRA GAFFNEY & MEYLAN, 1988
SUPERORDER: SELMACRYPTODIRA GAFFNEY, HUTCHISON, JENKINS & MEEKR, 1987
GRANDORDER: DAIOCRYPTODIRA GAFFNEY & MEYLAN, 1988
MICRORDER: CENTROCRYPTODIRA GAFFNEY & MEYLAN, 1988
ORDER: POLYCRPTODIRA GAFFNEY & MEYLAN, 1988
SUBORDER: PROCOELOCRYPTODIRA GAFFNEY & MEYLAN, 1988
INFRAORDER: CHELOMACRYPTODIRA GAFFNEY & MEYLAN, 1988
PARVORDER: TRIONYCHIDA GRAY, 1870
Superfamily: TRIONYCHIDEA Bell, 1828 (sensu Gray, 1870)
Family: TRIONYCHIDAE Bell, 1828
Subfamily: TRIONYCHINAE Bell, 1828 (sensu Lydekker, 1889)
Tribe: TRIONYCHINI Bell, 1828 (sensu Gaffney & Meylan, 1988)
Trionyx kyrgyzensis Trionyx dissolutus
____________________________________________________________________________________
UZBEIKSTAN
KARAKALPAKIA
SULTANUVAIS RIDGE
CHODZHAKULSAY
SHEICHDZHEILI
UPPER PART OF THE CHODZHAKUL FORMATION
APPROXIMATELY 14 M ABOVE THE LAYER 4
LOWER CENOMANIAN STAGE
UPPER GALIC SUBEPOCH
LOWER GULF EPOCH
MIDDLE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: AMPHIBIA LINNAEUS, 1758
SUBCLASS: LISSAMPHIBIA, HAECKEL, 1866?
SUPERORDER: BATRACHIA BRONGNIART, 1800
ORDER: CAUDATA SCOPOLI, 1777
SUBORDER: AMBYSTOMATOIDEA NOBLE, 1931
Family: SCAPHERPETONTIDAE Auffenberg & Goin, 1959
Horezmia gracile new subspecies?
SUPERORDER: SALIENTIA LAURENTI, 1768
ORDER: ANURA RAFINESQUE, 1815
SUBORDER: DISCOGLOSSOIDEI SOKOL, 1977
Family: DISCOGLOSSIDAE Guenther, 1859
Incertae sedis
Gen. sp indet.
SUBORDER: MESOBATRACHIA LAURENT, 1979
Superfamily: PELOBATOIDEA Stanius, 1856 , nom. corr. Spinar, 1971 (sensu Bolkay, 1919)
Family: PELOBATIDAE Stanius, 1856 , nom. corr. Spinar, 1971
Gen. sp indet.
CLASS: MAMMALIA LINNAEUS, 1758
SUBCLASS: PROTOTHERIA GILL, 1872
INFRACLASS: TRECHNOTHERIA McKENNA, 1975
LEGION: CLADOTHERIA McKENNA, 1975
SUBLEGION: ZATHERIA MCKENNA, 1975
INFRALEGION: TRIBOSPHENIDA (McKENNA, 1975) McKENNA & BELL, 1997
SUPERCOHORT: THERIA (PARKER & HASWELL, 1987) McKENNA & BELL, 1997
Incertae sedis
Family: PAPPOTHERIIDAE Slaughter, 1965
Gen. sp indet.
____________________________________________________________________________________
UZBEIKSTAN
KARAKALPAKIA
SULTANUVAIS RIDGE
CHODZHAKULSAY
SHEICHDZHEILI
UPPER PART OF THE CHODZHAKUL FORMATION
APPROXIMATELY 14 M ABOVE THE LAYER 4
LOWER CENOMANIAN STAGE
UPPER GALIC SUBEPOCH
LOWER GULF EPOCH
MIDDLE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: AMPHIBIA LINNAEUS, 1758
SUBCLASS: LISSAMPHIBIA, HAECKEL, 1866?
SUPERORDER: BATRACHIA BRONGNIART, 1800
ORDER: CAUDATA SCOPOLI, 1777
SUBORDER: AMBYSTOMATOIDEA NOBLE, 1931
Family: SCAPHERPETONTIDAE Auffenberg & Goin, 1959
Horezmia gracile new subspecies?
SUPERORDER: SALIENTIA LAURENTI, 1768
ORDER: ANURA RAFINESQUE, 1815
SUBORDER: DISCOGLOSSOIDEI SOKOL, 1977
Family: DISCOGLOSSIDAE Guenther, 1859
Incertae sedis
Gen. sp indet.
SUBORDER: MESOBATRACHIA LAURENT, 1979
Superfamily: PELOBATOIDEA Stanius, 1856 , nom. corr. Spinar, 1971 (sensu Bolkay, 1919)
Family: PELOBATIDAE Stanius, 1856 , nom. corr. Spinar, 1971
Gen. sp indet.
CLASS: MAMMALIA LINNAEUS, 1758
SUBCLASS: PROTOTHERIA GILL, 1872
INFRACLASS: TRECHNOTHERIA McKENNA, 1975
LEGION: CLADOTHERIA McKENNA, 1975
SUBLEGION: ZATHERIA MCKENNA, 1975
INFRALEGION: TRIBOSPHENIDA (McKENNA, 1975) McKENNA & BELL, 1997
SUPERCOHORT: THERIA (PARKER & HASWELL, 1987) McKENNA & BELL, 1997
Incertae sedis
Family: PAPPOTHERIIDAE Slaughter, 1965
Gen. sp indet.
____________________________________________________________________________________
UZBEIKSTAN
KARAKALPAKIA
SULTANUVAIS RIDGE
SHEICHDZHEILI
UPPER PART OF THE CHODZHAKUL FORMATION
LOWER CENOMANIAN STAGE
UPPERMIDDLE GALLIC SUBEPOCH
LOWERMOST GULF EPOCH
LATE MIDDLE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
INFRACLASS: NEODIAPSIDA BENTON, 1985
LEGION: LEPIDOSAUROMORPHA BENTON, 1983
SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)
MAGNORDER: SQUAMATA, OPPEL, 1811
SUPERORDER: SERPENTES LINNAEUS, 1759
ORDER: OPHIDIA BRONGNIART, 1800
SUBORDER: ALETHINOPHIDIA NOPCSCA, 1923
Superfamily: SIMOLIOPHEOIDEA Nopcsa, 1925
Family: SIMOLIOHEIDAE Nopcsa, 1925
Gen sp indet.
____________________________________________________________________________________
UZBEKISTAN
KARKALPAK AUTONOMOUS SSR
SHEILKHDZHELI
KHOZHAKUL FORMATION
LOWER CENOMANIAN STAGE
UPPER GALIC SUBEPOCH
LOWER GULF EPOCH
MIDDLE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: AVES LINNAEUS, 1758
Incertae sedis
Gen. sp indet.
____________________________________________________________________________________
UZBEKISTAN
NAVOISKAYA OBLAST
BELEUTINSKAYA SVITA
LATE TURONIAN STAGE
UPPERMOST GALLIC SUBEPOCH
LOWER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
AVEPODA PAUL, 2002
SUPERORDER: NEOTHEROPODA BAKKER, 1986
GRANDORDER: TETANURAE GAUTHIER, 1986
MICRORDER: COELUROSAURIA HUENE, 1914
ORDER: MANIRAPTORFORMES HOLTZ, 1996
AVETHEROPODA PAUL, 2001
AVEPECTORA PAUL, 2001
SUBORDER: MANIRAPTORA GAUTHIER, 1986
INFRAORDER: PARVES SERENO, 1997
PARVORDER: DEINONYCHOSAURIA COLBERT & D. A. RUSSELL, 1969
Family: DROMAEOSAURIDAE Russell, 1969
*Itemirus medullaris
_____________________________________________________________________________________
UZBEKISTAN
FROM THE NORTHERN BORDER OF SHEYKHDZHEYLI
UPPER KHODZHAKUL FORMATION
LOWER CENOMANIAN STAGE
UPPERMIDDLE GALLIC SUBEPOCH
LOWERMOST GULF EPOCH
MIDDLE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
AVEPODA PAUL, 2002
SUPERORDER: NEOTHEROPODA BAKKER, 1986
GRANDORDER: TETANURAE GAUTHIER, 1986
MICRORDER: COELUROSAURIA HUENE, 1914
ORDER: MANIRAPTORFORMES HOLTZ, 1996
AVETHEROPODA PAUL, 2001
AVEPECTORA PAUL, 2001
SUBORDER: MANIRAPTORA GAUTHIER, 1986
INFRAORDER: PARVES SERENO, 1997
PARVORDER: DEINONYCHOSAURIA COLBERT & D. A. RUSSELL, 1969
Family: TROODONTIDAE Gilmore, 1924
*Saurornithoides? asiamericanus
= Pectinodon asiamericanus
= Saurornithoides? asiamericanus
= Troodon asiamericanus
_____________________________________________________________________________________
UZBEKISTAN
TCELPYK
SOUTHWEST KYZYLKUM
?KHODZAKUL FORMATION
CENOMANIAN STAGE
MIDDLE GALLIC SUBEPOCH
LOWERMOST GULF EPOCH
MIDDLE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: AVEMETATARSALIA BENTON, 1999
COHORT: PROLACERTIFORMES CAMP, 1945 NOVA
SUPERORDER: TAPINOPLATIA PETERS, 2000
GRANDORDER: CHARACIOPODA PETERS, 2000
MICRORDER: FENESTRASAURIA PETERS, 2000
ORDER: PTEROSAURIA KAUP, 1834
Incertae sedis
Gen. sp indet.
____________________________________________________________________________________
UZBEIKSTAN
KARAKALPAKIA
TCELPYK HILL
UPPER PART OF THE CHODZHAKUL FORMATION OR LOWER BEDS OF THE BESTUBE FORMATION
CENOMANIAN-?LOWER TURONIAN STAGE
UPPER GALIC SUBEPOCH
LOWER GULF EPOCH
MIDDLE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: AMPHIBIA LINNAEUS, 1758
SUBCLASS: LISSAMPHIBIA, HAECKEL, 1866?
SUPERORDER: BATRACHIA BRONGNIART, 1800
ORDER: CAUDATA SCOPOLI, 1777
SUBORDER: AMBYSTOMATOIDEA NOBLE, 1931
Family: SCAPHERPETONTIDAE Auffenberg & Goin, 1959
Eoscapherpeton asiaticum
Horezmia gracile new subspecies?
SUPERORDER: SALIENTIA LAURENTI, 1768
Incertae sedis
Gen. sp indet.
ORDER: ANURA RAFINESQUE, 1815
Incertae sedis
Aralobatrachus robustus
Itemirella cretacea
SUBORDER: DISCOGLOSSOIDEI SOKOL, 1977
Family: DISCOGLOSSIDAE Guenther, 1859
Subfamily: ALYTINAE Fitzinger, 1943
Kizylkuma antiqua
SUBORDER: MESOBATRACHIA LAURENT, 1979
Superfamily: PELOBATOIDEA Stanius, 1856 , nom. corr. Spinar, 1971 (sensu Bolkay, 1919)
Family: PELOBATIDAE Stanius, 1856 , nom. corr. Spinar, 1971
Gen. sp indet.
____________________________________________________________________________________
UZBEIKSTAN
KARAKALPAKIA
SULTANUVAIS RIDGE
CHODZHAKULSAY
SHEICHDZHEILI
UPPER PART OF THE CHODZHAKUL FORMATION
APPROXIMATELY 14 M ABOVE THE LAYER 4
LOWER CENOMANIAN STAGE
UPPER GALIC SUBEPOCH
LOWER GULF EPOCH
MIDDLE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: AMPHIBIA LINNAEUS, 1758
SUBCLASS: LISSAMPHIBIA, HAECKEL, 1866?
SUPERORDER: BATRACHIA BRONGNIART, 1800
ORDER: CAUDATA SCOPOLI, 1777
SUBORDER: AMBYSTOMATOIDEA NOBLE, 1931
Family: SCAPHERPETONTIDAE Auffenberg & Goin, 1959
Horezmia gracile new subspecies?
SUPERORDER: SALIENTIA LAURENTI, 1768
ORDER: ANURA RAFINESQUE, 1815
SUBORDER: DISCOGLOSSOIDEI SOKOL, 1977
Family: DISCOGLOSSIDAE Guenther, 1859
Incertae sedis
Gen. sp indet.
SUBORDER: MESOBATRACHIA LAURENT, 1979
Superfamily: PELOBATOIDEA Stanius, 1856 , nom. corr. Spinar, 1971 (sensu Bolkay, 1919)
Family: PELOBATIDAE Stanius, 1856 , nom. corr. Spinar, 1971
Gen. sp indet.
____________________________________________________________________________________
UZBEKISTAN
KARAKALPAKIA
SOUTH WEST KYZYLKUM
EAST OF THE SULTAN-UVAYS RANGE
AYAZKALA UPLAND
KHODZHAKULIAN SUITE
CENOMANIAN STAGE
MIDDLE GALLIC SUBEPOCH
LOWERMOST GULF EPOCH
MIDDLE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: CHELONOMORPHA KUHN, 1960
SUPERCOHORT: TESTUDINES LINNAEUS, 1758
COHORT: CASICHELYDIA emend GAFFNEY & MEYLAN, 1988
MAGANORDER: CRYPTODIRA GAFFNEY & MEYLAN, 1988
SUPERORDER: SELMACRYPTODIRA GAFFNEY, HUTCHISON, JENKINS & MEEKR, 1987
GRANDORDER: DAIOCRYPTODIRA GAFFNEY & MEYLAN, 1988
MICRORDER: CENTROCRYPTODIRA GAFFNEY & MEYLAN, 1988
ORDER: POLYCRPTODIRA GAFFNEY & MEYLAN, 1988
SUBORDER: PROCOELOCRYPTODIRA GAFFNEY & MEYLAN, 1988
INFRAORDER: CHELOMACRYPTODIRA GAFFNEY & MEYLAN, 1988
PARVORDER: TRIONYCHIDA GRAY, 1870
Superfamily: TRIONYCHIDEA Bell, 1828 (sensu Gray, 1870)
Family: CARETTOCHELYIDAE Boulenger, 1887
Subfamily: ANOSTERINAE Lydekker, 1889
Kizylkumemys schultzi
____________________________________________________________________________________
UZBEKISTAN
KHODZHAKULSAY
SOUTHWEST KYZYLKUM
KHODZAKUL FORMATION
CENOMANIAN STAGE
MIDDLE GALLIC SUBEPOCH
LOWERMOST GULF EPOCH
MIDDLE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: AVEMETATARSALIA BENTON, 1999
COHORT: PROLACERTIFORMES CAMP, 1945 NOVA
SUPERORDER: TAPINOPLATIA PETERS, 2000
GRANDORDER: CHARACIOPODA PETERS, 2000
MICRORDER: FENESTRASAURIA PETERS, 2000
ORDER: PTEROSAURIA KAUP, 1834
Incertae sedis
Gen. sp indet.
____________________________________________________________________________________
UZBEKISTAN
KHODZHAKULSAY
SOUTHWEST KYZYLKUM
KHODZAKUL FORMATION
CENOMANIAN STAGE
MIDDLE GALLIC SUBEPOCH
LOWERMOST GULF EPOCH
MIDDLE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: AVEMETATARSALIA BENTON, 1999
COHORT: PROLACERTIFORMES CAMP, 1945 NOVA
SUPERORDER: TAPINOPLATIA PETERS, 2000
GRANDORDER: CHARACIOPODA PETERS, 2000
MICRORDER: FENESTRASAURIA PETERS, 2000
ORDER: PTEROSAURIA KAUP, 1834
Incertae sedis
Gen. sp indet.
____________________________________________________________________________________
UZBEKISTAN
KARAKALPAK ASSR
WESTERN PART OF SULTAN-AVAYS RANGE
RIGHT SLOPE OF THE KHODZHAKULSAY
UPPER PART OF KHODZHAKULIAN SUITE
CENOMANIAN STAGE
MIDDLE GALLIC SUBEPOCH
LOWERMOST GULF EPOCH
MIDDLE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: CHELONOMORPHA KUHN, 1960
SUPERCOHORT: TESTUDINES LINNAEUS, 1758
COHORT: CASICHELYDIA emend GAFFNEY & MEYLAN, 1988
MAGANORDER: CRYPTODIRA GAFFNEY & MEYLAN, 1988
SUPERORDER: SELMACRYPTODIRA GAFFNEY, HUTCHISON, JENKINS & MEEKR, 1987
GRANDORDER: DAIOCRYPTODIRA GAFFNEY & MEYLAN, 1988
MICRORDER: CENTROCRYPTODIRA GAFFNEY & MEYLAN, 1988
ORDER: POLYCRPTODIRA GAFFNEY & MEYLAN, 1988
SUBORDER: PROCOELOCRYPTODIRA GAFFNEY & MEYLAN, 1988
INFRAORDER: CHELOMACRYPTODIRA GAFFNEY & MEYLAN, 1988
PARVORDER: TRIONYCHIDA GRAY, 1870
Superfamily: TRIONYCHIDEA Bell, 1828 (sensu Gray, 1870)
Family: CARETTOCHELYIDAE Boulenger, 1887
Subfamily: ANOSTERINAE Lydekker, 1889
*Kizylkumemys schultzi
____________________________________________________________________________________
UZBEKISTAN
KARAKALPAKIAN AUTONOMOUS REPUBLIC
SKH-1
CHELPYK LOCALITY
LOWER? CENOMANIAN STAGE
MIDDLE GALLIC SUBEPOCH
LOWERMOST GULF EPOCH
LATE EARLY CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
Incertae sedis
Gen sp indet.
COHORT: ORNITHISCHIFORMES COOPER, 1985
MAGNORDER: ORNITHISCHIA SEELEY, 1888
SUPERORDER: NEORNITHISCHIA COOPER, 1985
GRANDORDER: ORNITHOPODA MARSH, 1871
MICRORDER: EUORNITHOPODA SERENO, 1986
ORDER: IGUANODONTIA DOLLO, 1888
INFRAORDER: STYRACOSTERNA SERENO, 1986
PARVORDER: HADROSAURIFORMES SERENO, 1997
SUPERFAMILY: HADROSAUROIDEA COPE, 1869 (SENSU SERENO, 1986)
Family: HADROSAURIDAE COPE, 1869
Subfamily: HADROSAURINAE Cope, 1869 (sensu Lambe, 1918)
Gilmoreosaurus sp
MICRORDER: MARGINOCEPHALIA SERENO, 1986
ORDER: CERATOPIA MARSH, 1890
SUBORDER: NEOCERATOPIA SERENO, 1986
INFRAORDER: CORONOSAURIA SERENO, 1986
Family: PROTOCERATOPIDAE Granger & Gregory, 1923
Incertae sedis
Asiaceratops salsopaludalis
SUPERORDER: EURYOPDA SERENO, 1986
GRANDORDER: ARMATOSAURIA ZHAO, 1983 (emend)
ORDER: ANKYLOSAURIA OSBORN, 1923
INFRAORDER: ANKYLOSAURA OSBORN, 1923
Family: ANKYLOSAURIDAE Brown, 1908
Incertae sedis
Gen. sp indet.
CLASS: MAMMALIA LINNAEUS, 1758
SUBCLASS: PROTOTHERIA GILL, 1872
INFRACLASS: TRECHNOTHERIA McKENNA, 1975
LEGION: CLADOTHERIA McKENNA, 1975
SUBLEGION: ZATHERIA MCKENNA, 1975
INFRALEGION: TRIBOSPHENIDA (McKENNA, 1975) McKENNA & BELL, 1997
SUPERCOHORT: THERIA (PARKER & HASWELL, 1987) McKENNA & BELL, 1997
Incertae sedis
Gen. sp indet.
____________________________________________________________________________________
UZBEKISTAN
KARAKALPAKIAN AUTONOMOUS REPBULIC
KHODZHAKULSAY LOCALITY
SITE SKH-5
CENOMANIAN STAGE
MIDDLE GALLIC SUBEPOCH
LOWERMOST GULF EPOCH
MIDDLE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: MAMMALIA LINNAEUS, 1758
SUBCLASS: PROTOTHERIA GILL, 1872
INFRACLASS: TRECHNOTHERIA McKENNA, 1975
LEGION: CLADOTHERIA McKENNA, 1975
SUBLEGION: ZATHERIA MCKENNA, 1975
INFRALEGION: TRIBOSPHENIDA (McKENNA, 1975) McKENNA & BELL, 1997
SUPERCOHORT: THERIA (PARKER & HASWELL, 1987) McKENNA & BELL, 1997
Family: Uncertain
Gen. sp indet.
_____________________________________________________________________________________
UZBEKISTAN
KARAKALPAKISTAN
SOUTHWESTERN KYZYLKUM
KHODZHAKULSAI
UPPER PART OF KHODZHAKUL FORMATION (KHODZHAKUL’SKAYA SVITA)
LOWER CENOMANIAN STAGE
UPPERMIDDLE GALLIC SUBEPOCH
LOWERMOST GULF EPOCH
MIDDLE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
INFRACLASS: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
AVEPODA PAUL, 2002
SUPERORDER: NEOTHEROPODA BAKKER, 1986
GRANDORDER: TETANURAE GAUTHIER, 1986
MICRORDER: COELUROSAURIA HUENE, 1914
ORDER: MANIRAPTORFORMES HOLTZ, 1996
AVETHEROPODA PAUL, 2001
AVEPECTORA PAUL, 2001
SUBORDER: MANIRAPTORA GAUTHIER, 1986
INFRAORDER: PARVES SERENO, 1997
PARVORDER: DEINONYCHOSAURIA COLBERT & D. A. RUSSELL, 1969
Family: DROMAEOSAURIDAE Russell, 1969
Incerate sedis
non Dromaeosaurus, *D? explanatus
= Laelaps explanatusGen. sp indet.
COHORT: ORNITHISCHIFORMES COOPER, 1985
SUPERORDER: SAUROPODOMORPHA HUENE, 1932
GRANDORDER: BRONTOSAURIA OLSHEVSKY, 1991 (emend)
MICRORDER: SAUROPODA MARSH, 1878
ORDER: EUSAUROPODA
SUBORDER: NEOSAUROPODA BONAPARTE, 1986
INFRAORDER: MACRONARIA WILSON & SERENO, 1998
PARVORDER: TITANOSAURIFORMES SALGADO ET AL, 1997
Superfamily: TITANOSAUROIDEA Lydekker, 1885 (sensu Upchurch, 1999)
Family: TITANOSAURIDAE Lydekker, 1885
Incertae sedis
Gen. sp indet.
MAGNORDER: ORNITHISCHIA SEELEY, 1888
SUPERORDER: NEORNITHISCHIA COOPER, 1985
GRANDORDER: ORNITHOPODA MARSH, 1871
MICRORDER: EUORNITHOPODA SERENO, 1986
ORDER: IGUANODONTIA DOLLO, 1888
INFRAORDER: STYRACOSTERNA SERENO, 1986
PARVORDER: HADROSAURIFORMES SERENO, 1997
SUPERFAMILY: HADROSAUROIDEA COPE, 1869 (SENSU SERENO, 1986)
Family: HADROSAURIDAE COPE, 1869
Incertae sedis
Gen. sp indet.
MICRORDER: MARGINOCEPHALIA SERENO, 1986
ORDER: CERATOPIA MARSH, 1890
SUBORDER: NEOCERATOPIA SERENO, 1986
INFRAORDER: CORONOSAURIA SERENO, 1986
Family: PROTOCERATOPIDAE Granger & Gregory, 1923
Incertae sedis
Asiaceratops sp
SUPERORDER: EURYOPDA SERENO, 1986
GRANDORDER: ARMATOSAURIA ZHAO, 1983 (emend)
ORDER: ANKYLOSAURIA OSBORN, 1923
INFRAORDER: ANKYLOSAURA OSBORN, 1923
Family: ANKYLOSAURIDAE Brown, 1908
Incertae sedis
Gen. sp indet.
____________________________________________________________________________________
UZBEKISTAN
KARAKALPAKISTAN
KARATEPA (SMALL ELEVATION NEAR THE CHELPYK MOUNTAINS)
UPPER PART OF KHODZHAKUL FORMATION (KHODZHAKUL’SKAYA SVITA)
LOWER CENOMANIAN STAGE
UPPERMIDDLE GALLIC SUBEPOCH
LOWERMOST GULF EPOCH
MIDDLE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
Incertae sedis
Gen sp indet.
COHORT: ORNITHISCHIFORMES COOPER, 1985
MAGNORDER: ORNITHISCHIA SEELEY, 1888
SUPERORDER: NEORNITHISCHIA COOPER, 1985
GRANDORDER: ORNITHOPODA MARSH, 1871
MICRORDER: EUORNITHOPODA SERENO, 1986
ORDER: IGUANODONTIA DOLLO, 1888
INFRAORDER: STYRACOSTERNA SERENO, 1986
PARVORDER: HADROSAURIFORMES SERENO, 1997
SUPERFAMILY: HADROSAUROIDEA COPE, 1869 (SENSU SERENO, 1986)
Family: HADROSAURIDAE COPE, 1869
Gen. sp indet.
_____________________________________________________________________________________
UZBEKISTAN
KARAKALPAKISTAN
RAVINE ON THE EASTERN EDGE OF THE SULTAN-UVAIS RANGE
KARACHADALYSAI
UPPER (?) PART OF THE KHODZHAKUL FORMATION (KHODZHAKUL’SKAYA SVITA)
CENOMANIAN STAGE
UPPER GALLIC SUBEPOCH
LOWEST GULF EPOCH
LATE EARLY TO EARLY MIDDLE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: ORNITHISCHIFORMES COOPER, 1985
MAGNORDER: ORNITHISCHIA SEELEY, 1888
SUPERORDER: NEORNITHISCHIA COOPER, 1985
GRANDORDER: ORNITHOPODA MARSH, 1871
MICRORDER: EUORNITHOPODA SERENO, 1986
ORDER: IGUANODONTIA DOLLO, 1888
INFRAORDER: STYRACOSTERNA SERENO, 1986
PARVORDER: HADROSAURIFORMES SERENO, 1997
SUPERFAMILY: HADROSAUROIDEA COPE, 1869 (SENSU SERENO, 1986)
Family: HADROSAURIDAE COPE, 1869
Gen. sp indet.
_____________________________________________________________________________________
UZBEKISTAN
KARAKALPAKISTAN
EAST OF THE SULTAN-UVAIS RANGE
HILLS WITH FORTRESS ON THE TOP
KYRKKYZ
UPPER PART OF THE KHODZHAKUL FORMATION (KHODZHAKUL’SKAYA SVITA)
CENOMANIAN STAGE
UPPER GALLIC SUBEPOCH
LOWEST GULF EPOCH
LATE EARLY TO EARLY MIDDLE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
INFRACLASS: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
Incertae sedis
Gen. sp indet.
COHORT: ORNITHISCHIFORMES COOPER, 1985
MAGNORDER: ORNITHISCHIA SEELEY, 1888
SUPERORDER: NEORNITHISCHIA COOPER, 1985
GRANDORDER: ORNITHOPODA MARSH, 1871
MICRORDER: EUORNITHOPODA SERENO, 1986
ORDER: IGUANODONTIA DOLLO, 1888
INFRAORDER: STYRACOSTERNA SERENO, 1986
PARVORDER: HADROSAURIFORMES SERENO, 1997
SUPERFAMILY: HADROSAUROIDEA COPE, 1869 (SENSU SERENO, 1986)
Family: HADROSAURIDAE COPE, 1869
Gen. sp indet.
_____________________________________________________________________________________
UZBEKISTAN
KARAKALPAKISTAN
HILLS BETWEEN THE EASTERN EXTREMITY OF THE SULTAN-UVAIS RANGE AND THE AYAZKALA ELEVATION
KULKALA
UPPER PART OF THE KHODZHAKUL FORMATION (KHODZHAKUL’SKAYA SVITA)
CENOMANIAN STAGE
UPPER GALLIC SUBEPOCH
LOWEST GULF EPOCH
LATE EARLY TO EARLY MIDDLE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
INFRACLASS: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: ORNITHISCHIFORMES COOPER, 1985
MAGNORDER: ORNITHISCHIA SEELEY, 1888
SUPERORDER: NEORNITHISCHIA COOPER, 1985
GRANDORDER: ORNITHOPODA MARSH, 1871
MICRORDER: EUORNITHOPODA SERENO, 1986
ORDER: IGUANODONTIA DOLLO, 1888
INFRAORDER: STYRACOSTERNA SERENO, 1986
PARVORDER: HADROSAURIFORMES SERENO, 1997
SUPERFAMILY: HADROSAUROIDEA COPE, 1869 (SENSU SERENO, 1986)
Family: HADROSAURIDAE COPE, 1869
Gen. sp indet.
_____________________________________________________________________________________
UZBEKISTAN
KARAKALPAKISTAN
SHALLOW RAVINE 7 KM EAST-NORTH-EAST OF THE SETTLEMENT OF KYZYLKALA
KYZYLKALA II
UPPER PART OF THE KHODZHKUL FORMATION OR BESHTYUBIN FORMATION
CENOMANIAN STAGE
UPPERMIDDLE GALLIC SUBEPOCH
LOWERMOST GULF EPOCH
LATE EARLY CRETACEOUS-LOWER TURONIAN STAGE
UPPERMOST GALLIC SUBEPOCH
LOWER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
Incertae sedis
Gen. sp indet.
_____________________________________________________________________________________
UZBEKISTAN
WESTERN SULTANUVAIS RIDGE
CHODZHAKUL LAKE LOCALITY
KHODZHAKUL I (KHODZHAKUL LOCALITY)
LOWER OR MIDDLE KHODZHAKUL (CHODZHAKUL) FORMATION
EARLY CENOMANIAN STAGE
UPPERMIDDLE GALLIC SUBEPOCH
LOWERMOST GULF EPOCH
MIDDLE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: CHELONOMORPHA KUHN, 1960
SUPERCOHORT: TESTUDINES LINNAEUS, 1758
COHORT: CASICHELYDIA emend GAFFNEY & MEYLAN, 1988
MAGANORDER: CRYPTODIRA GAFFNEY & MEYLAN, 1988
SUPERORDER: SELMACRYPTODIRA GAFFNEY, HUTCHISON, JENKINS & MEEKR, 1987
GRANDORDER: DAIOCRYPTODIRA GAFFNEY & MEYLAN, 1988
MICRORDER: CENTROCRYPTODIRA GAFFNEY & MEYLAN, 1988
ORDER: POLYCRPTODIRA GAFFNEY & MEYLAN, 1988
SUBORDER: PROCOELOCRYPTODIRA GAFFNEY & MEYLAN, 1988
INFRAORDER: CHELOMACRYPTODIRA GAFFNEY & MEYLAN, 1988
PARVORDER: TRIONYCHIDA GRAY, 1870
Superfamily: TRIONYCHIDEA Bell, 1828 (sensu Gray, 1870)
Family: ADOCIDAE Cope, 1870
*Adocus kizylkumensis
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UZBEKISTAN
KARAKALPAKISTAN
SOUTHWESTERN KYZYLKUM
KHODZHAKULSAI
UPPER PART OF KHODZHAKUL FORMATION (KHODZHAKUL’SKAYA SVITA)
LOWER CENOMANIAN STAGE
UPPERMIDDLE GALLIC SUBEPOCH
LOWERMOST GULF EPOCH
MIDDLE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
INFRACLASS: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
AVEPODA PAUL, 2002
SUPERORDER: NEOTHEROPODA BAKKER, 1986
GRANDORDER: TETANURAE GAUTHIER, 1986
MICRORDER: COELUROSAURIA HUENE, 1914
ORDER: MANIRAPTORFORMES HOLTZ, 1996
AVETHEROPODA PAUL, 2001
AVEPECTORA PAUL, 2001
SUBORDER: MANIRAPTORA GAUTHIER, 1986
INFRAORDER: PARVES SERENO, 1997
PARVORDER: DEINONYCHOSAURIA COLBERT & D. A. RUSSELL, 1969
Family: DROMAEOSAURIDAE Russell, 1969
Incerate sedis
non Dromaeosaurus, *D? explanatus
= Laelaps explanatusGen. sp indet.
COHORT: ORNITHISCHIFORMES COOPER, 1985
SUPERORDER: SAUROPODOMORPHA HUENE, 1932
GRANDORDER: BRONTOSAURIA OLSHEVSKY, 1991 (emend)
MICRORDER: SAUROPODA MARSH, 1878
ORDER: EUSAUROPODA
SUBORDER: NEOSAUROPODA BONAPARTE, 1986
INFRAORDER: MACRONARIA WILSON & SERENO, 1998
PARVORDER: TITANOSAURIFORMES SALGADO ET AL, 1997
Superfamily: TITANOSAUROIDEA Lydekker, 1885 (sensu Upchurch, 1999)
Family: TITANOSAURIDAE Lydekker, 1885
Incertae sedis
Gen. sp indet.
MAGNORDER: ORNITHISCHIA SEELEY, 1888
SUPERORDER: NEORNITHISCHIA COOPER, 1985
GRANDORDER: ORNITHOPODA MARSH, 1871
MICRORDER: EUORNITHOPODA SERENO, 1986
ORDER: IGUANODONTIA DOLLO, 1888
INFRAORDER: STYRACOSTERNA SERENO, 1986
PARVORDER: HADROSAURIFORMES SERENO, 1997
SUPERFAMILY: HADROSAUROIDEA COPE, 1869 (SENSU SERENO, 1986)
Family: HADROSAURIDAE COPE, 1869
Incertae sedis
Gen. sp indet.
MICRORDER: MARGINOCEPHALIA SERENO, 1986
ORDER: CERATOPIA MARSH, 1890
SUBORDER: NEOCERATOPIA SERENO, 1986
INFRAORDER: CORONOSAURIA SERENO, 1986
Family: PROTOCERATOPIDAE Granger & Gregory, 1923
Incertae sedis
Asiaceratops salsopaludalis