UZBEKISTAN
NAVOI VILOYAK
32 KM SW OF MYNBULAK
CENTRAL KYZYLKUM DESERT
DZHARAKUDUK
BISSEKTY FORMATION
TURONIAN STAGE
UPPERMOST GALLIC SUBEPOCH
LOWER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: ORNITHISCHIFORMES COOPER, 1985
SUPERORDER: SAUROPODOMORPHA HUENE, 1932
GRANDORDER: BRONTOSAURIA OLSHEVSKY, 1991 (emend)
MICRORDER: SAUROPODA MARSH, 1878
ORDER: EUSAUROPODA UPCHURCH, 1995
SUBORDER: NEOSAUROPODA BONAPARTE, 1986
INFRAORDER: MACRONARIA WILSON & SERENO, 1998
PARVORDER: TITANOSAURIFORMES SALGADO ET AL, 1997
Superfamily: TITANOSAUROIDEA Lydekker, 1885 (sensu Upchurch, 1999)
Family: TITANOSAURIDAE Lydekker, 1885
?COLOSSOSAURIA GONZALEZ RIGA, LAMANNA, OTERO, ORTIZ, KELLNER & IBIRICU, 2019
CLADE NAME:LOGNKOSAURIA CALVO, PORFIRI, GONZALEZ-RIGA & KELLNER, 2007
*Dzharatitanis kingi
_____________________________________________________________________________________
UZBEKISTAN
NAVOI DISTRICT
32 KM SW OF MYNBULAK
CENTRAL KYZYLKUM DESERT
DZHARAKUDUK
UNKNOWN
BISSEKTY FORMATION
MIDDLE TO UPPER TURONIAN STAGE
UPPERMOST GALLIC SUBEPOCH
LOWER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
AVEPODA PAUL, 2002
SUPERORDER: NEOTHEROPODA BAKKER, 1986
GRANDORDER: TETANURAE GAUTHIER, 1986
ORDER: NEOTETANURAE SERENO, WILSON, LARSSON, DUTHEIL &
SUBORDER: AVETHEROPODA PAUL, 1988
Superfamily: ALLOSAUROIDEA Marsh, 1878 (sensu Currie & Zhao, 1994)
Clade ALLOSAURIA PAUL, 1988
CARCHARODONTOSAURIA BENSON, CARRANO & BRUSATTE, 2009
Incertae sedis
*Ulughbegsaurus uzbekistanensis
____________________________________________________________________________________
UZBEKISTAN
NAVOI DISTRICT
32 KM SW OF MYNBULAK
CENTRAL KYZYLKUM DESERT
APPROXIMATELY 8 KM LONG ESCARPEMTN NEAR THE SMALL SETTLEMENT OF DZHARAKUDUK (ALSO CALLED DZHARA-KUDUK, DZHIRAKUDUK, DZHYRAKUDUK, BISSEKTY, AND KUL'BEKE)
BISSEKTY FORMATION
TURONIAN STAGE
UPPERMOST GALLIC SUBEPOCH
LOWER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
AVEPODA PAUL, 2002
SUPERORDER: NEOTHEROPODA BAKKER, 1986
GRANDORDER: TETANURAE GAUTHIER, 1986
ORDER: NEOTETANURAE SERENO, WILSON, LARSSON, DUTHEIL &
SUBORDER: AVETHEROPODA PAUL, 1988
Superfamily: ALLOSAUROIDEA Marsh, 1878 (sensu Currie & Zhao, 1994)
Clade ALLOSAURIA PAUL, 1988
CARCHARODONTOSAURIA BENSON, CARRANO & BRUSATTE, 2009
Incertae sedis
Ulughbegsaurus uzbekistanensis
MICRORDER: COELUROSAURIA HUENE, 1914
ORDER: MANIRAPTORFORMES HOLTZ, 1996
AVETHEROPODA PAUL, 2001
AVEPECTORA PAUL, 2001
SUBORDER: MANIRAPTORA GAUTHIER, 1986
INFRAORDER: PARVES SERENO, 1997
PARVORDER: DEINONYCHOSAURIA COLBERT & D. A. RUSSELL, 1969
Family: DROMAEOSAURIDAE Russell, 1969
Itemirus medullaris
____________________________________________________________________________________
UZBEKISTAN
KYZYL KUM DESERT
ZENGE KURGAN 3 LOCALITY
BISSEKTY FORMATION
TURONIAN STAGE
UPPERMOST GALLIC SUBEPOCH
EARLY LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: AVES LINNAEUS, 1758
Incertae sedis
Gen. sp indet.
_____________________________________________________________________________________
UZBEKISTAN
KHOREZM DISTRICT
RIGHT BANK OF THE AMU DARYA RIVER BETWEEN THE VILLAGES OF BEZERGEN AND KULATAU
ZENGE KURGAN 3
BISSEKTY (?) FORMATION
TURONIAN STAGE
UPPER GALLIC SUBEPOCH
LOWER GULF EPOCH
LATE MIDDLE TO EARLY LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: AVEMETATARSALIA BENTON, 1999
COHORT: PROLACERTIFORMES CAMP, 1945 NOVA
SUPERORDER: TAPINOPLATIA PETERS, 2000
GRANDORDER: CHARACIOPODA PETERS, 2000
MICRORDER: FENESTRASAURIA PETERS, 2000
ORDER: PTEROSAURIA KAUP, 1834
SUBORDER: EUPTERODACTYLOIDEA BENNETT, 1994
Superfamily: AZHDARCHIDEA Nesov, 1984
Family: AZHDARCHIDAE Nesov, 1984 emend Padian, 1986
Gen. sp indet.
_____________________________________________________________________________________
UZBEKISTAN
CENTRAL KYZYLKUM
NORTHEASTERN PART OF THE BUKANTAU RIDGE
KARAMURUM
BISSEKTY FORMATION
LATE TURONIAN
UPPERMOST GALLIC SUBEPOCH- EARLY CONIACIAN STAGE
LOWER SENONIAN SUBEPOCH
LOWER GULF EPOCH
EARLY LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
Incertae sedis
Gen. sp indet.
COHORT: ORNITHISCHIFORMES COOPER, 1985
MAGNORDER: ORNITHISCHIA SEELEY, 1888
SUPERORDER: NEORNITHISCHIA COOPER, 1985
GRANDORDER: ORNITHOPODA MARSH, 1871
MICRORDER: EUORNITHOPODA SERENO, 1986
ORDER: IGUANODONTIA DOLLO, 1888
INFRAORDER: STYRACOSTERNA SERENO, 1986
PARVORDER: HADROSAURIFORMES SERENO, 1997
SUPERFAMILY: HADROSAUROIDEA COPE, 1869 (SENSU SERENO, 1986)
Family: HADROSAURIDAE COPE, 1869
Incertae sedis
Gen. sp indet.
_____________________________________________________________________________________
UZBEKISTAN
CENTRAL KYZYLKUM
LOCALITY IN THE SOUTH OF THE ARISTANTAU RIDGE
EDDUM
BISSEKTY FORMATION
LATE TURONIAN
UPPERMOST GALLIC SUBEPOCH- EARLY CONIACIAN STAGE
LOWER SENONIAN SUBEPOCH
LOWER GULF EPOCH
EARLY LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
Incertae sedis
Gen. sp indet.
COHORT: ORNITHISCHIFORMES COOPER, 1985
MAGNORDER: ORNITHISCHIA SEELEY, 1888
SUPERORDER: NEORNITHISCHIA COOPER, 1985
GRANDORDER: ORNITHOPODA MARSH, 1871
MICRORDER: EUORNITHOPODA SERENO, 1986
ORDER: IGUANODONTIA DOLLO, 1888
INFRAORDER: STYRACOSTERNA SERENO, 1986
PARVORDER: HADROSAURIFORMES SERENO, 1997
SUPERFAMILY: HADROSAUROIDEA COPE, 1869 (SENSU SERENO, 1986)
Family: HADROSAURIDAE COPE, 1869
Incertae sedis
Gen. sp indet.
_____________________________________________________________________________________
UZBEKISTAN
CENTRAL KYZYLKUM
BUKANTAU RIDGE
DZHEIRANTUI FORMATION
EARLY TURONIAN STAGE
UPPERMOST GALLIC SUBEPOCH
LOWERMOST GULF EPOCH
EARLY LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
Incertae sedis
Gen. sp indet.
_____________________________________________________________________________________
UZBEKISTAN
CENTRAL KYZYLKUM
SOUTH OF THE BUKANTAU MOUNTAINS
1-1.5 KM SOUTH OR SOUTHEAST OF THE SETTLEMENT
KULKUDUK
EARLY TURONIAN STAGE
UPPERMOST GALLIC SUBEPOCH
LOWERMOST GULF EPOCH
EARLY LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: CHELONOMORPHA KUHN, 1960
SUPERCOHORT: TESTUDINES LINNAEUS, 1758
COHORT: CASICHELYDIA emend GAFFNEY & MEYLAN, 1988
MAGANORDER: CRYPTODIRA GAFFNEY & MEYLAN, 1988
SUPERORDER: SELMACRYPTODIRA GAFFNEY, HUTCHISON, JENKINS & MEEKR, 1987
GRANDORDER: DAIOCRYPTODIRA GAFFNEY & MEYLAN, 1988
MICRORDER: CENTROCRYPTODIRA GAFFNEY & MEYLAN, 1988
ORDER: POLYCRPTODIRA GAFFNEY & MEYLAN, 1988
SUBORDER: PROCOELOCRYPTODIRA GAFFNEY & MEYLAN, 1988
INFRAORDER: CHELOMACRYPTODIRA GAFFNEY & MEYLAN, 1988
PARVORDER: TRIONYCHIDA GRAY, 1870
Superfamily: TRIONYCHIDEA Bell, 1828 (sensu Gray, 1870)
Family: CARETTOCHELYIDAE Boulenger, 1887
Gen. sp indet.
SUBCLASS: DIAPSIDA OSBORN, 1903
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: ORNITHISCHIFORMES COOPER, 1985
MAGNORDER: ORNITHISCHIA SEELEY, 1888
SUPERORDER: NEORNITHISCHIA COOPER, 1985
GRANDORDER: ORNITHOPODA MARSH, 1871
MICRORDER: EUORNITHOPODA SERENO, 1986
ORDER: IGUANODONTIA DOLLO, 1888
INFRAORDER: STYRACOSTERNA SERENO, 1986
PARVORDER: HADROSAURIFORMES SERENO, 1997
SUPERFAMILY: HADROSAUROIDEA COPE, 1869 (SENSU SERENO, 1986)
Family: HADROSAURIDAE COPE, 1869
Incertae sedis
Incertae sedis
SUPERORDER: EURYOPDA SERENO, 1986
GRANDORDER: ARMATOSAURIA ZHAO, 1983 (emend)
ORDER: ANKYLOSAURIA OSBORN, 1923
INFRAORDER: ANKYLOSAURA OSBORN, 1923
Family: ANKYLOSAURIDAE Brown, 1908
Incertae sedis
Gen. sp indet.
_____________________________________________________________________________________
UZBEKISTAN
CENTRAL KYZYLKUM
SOUTH OF THE BUKANTAU MOUNTAINS
1-1.5 KM SOUTH OR SOUTHEAST OF THE SETTLEMENT
KULKUDUK
LATE TURONIAN?
UPPERMOST GALLIC SUBEPOCH- EARLY CONIACIAN STAGE?
LOWER SENONIAN SUBEPOCH
LOWER GULF EPOCH
EARLY LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
Incertae sedis
Gen. sp indet.
COHORT: ORNITHISCHIFORMES COOPER, 1985
MAGNORDER: ORNITHISCHIA SEELEY, 1888
SUPERORDER: NEORNITHISCHIA COOPER, 1985
GRANDORDER: ORNITHOPODA MARSH, 1871
MICRORDER: EUORNITHOPODA SERENO, 1986
ORDER: IGUANODONTIA DOLLO, 1888
INFRAORDER: STYRACOSTERNA SERENO, 1986
PARVORDER: HADROSAURIFORMES SERENO, 1997
SUPERFAMILY: HADROSAUROIDEA COPE, 1869 (SENSU SERENO, 1986)
Family: HADROSAURIDAE COPE, 1869
Incertae sedis
Incertae sedis
SUPERORDER: EURYOPDA SERENO, 1986
GRANDORDER: ARMATOSAURIA ZHAO, 1983 (emend)
ORDER: ANKYLOSAURIA OSBORN, 1923
INFRAORDER: ANKYLOSAURA OSBORN, 1923
Family: ANKYLOSAURIDAE Brown, 1908
Incertae sedis
Gen. sp indet.
_____________________________________________________________________________________
UZBEKISTAN
CENTRAL KYZYLKUM
SOUTH OF THE BUKANTAU MOUNTAINS
1-1.5 KM SOUTH OR SOUTHEAST OF THE SETTLEMENT
KULKUDUK
BASE OF THE BISSEKTY FORMATION
LATE TURONIAN
UPPERMOST GALLIC SUBEPOCH- EARLY CONIACIAN STAGE
LOWER SENONIAN SUBEPOCH
LOWER GULF EPOCH
EARLY LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: ORNITHISCHIFORMES COOPER, 1985
SUPERORDER: SAUROPODOMORPHA HUENE, 1932
GRANDORDER: BRONTOSAURIA OLSHEVSKY, 1991 (emend)
MICRORDER: SAUROPODA MARSH, 1878
Incertae sedis
Gen. sp indet.
MAGNORDER: ORNITHISCHIA SEELEY, 1888
SUPERORDER: NEORNITHISCHIA COOPER, 1985
GRANDORDER: ORNITHOPODA MARSH, 1871
MICRORDER: EUORNITHOPODA SERENO, 1986
ORDER: IGUANODONTIA DOLLO, 1888
INFRAORDER: STYRACOSTERNA SERENO, 1986
PARVORDER: HADROSAURIFORMES SERENO, 1997
SUPERFAMILY: HADROSAUROIDEA COPE, 1869 (SENSU SERENO, 1986)
Family: HADROSAURIDAE COPE, 1869
Subfamily: LAMBEOSAURINAE Parks, 1923
Gen. sp indet.
SUPERORDER: EURYOPDA SERENO, 1986
GRANDORDER: ARMATOSAURIA ZHAO, 1983 (emend)
ORDER: ANKYLOSAURIA OSBORN, 1923
INFRAORDER: ANKYLOSAURA OSBORN, 1923
Family: ANKYLOSAURIDAE Brown, 1908
Incertae sedis
Gen. sp indet.
_____________________________________________________________________________________
UZEBEKISTAN
CENTRAL KYZYLKUM
WELL IN THE KULDZHUKTAU RIDGE
AIGYRBULAK
TURONIAN
UPPERMOST GALLIC SUBEPOCH
LOWER GULF EPOCH
EARLY LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
Incertae sedis
Gen. sp indet.
COHORT: THEROPODA EMEND MARSH, 1881
Incertae sedis
Gen. sp indet.
_____________________________________________________________________________________
UZBEKISTAN
CENTRAL KYZYLKUM
VICINITY OF THE WELL IN THE KARAKATIN DEPRESSION
AZNEK
TURONIAN
UPPERMOST GALLIC SUBEPOCH
LOWER GULF EPOCH
EARLY LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
Incertae sedis
Gen. sp indet.
_____________________________________________________________________________________
UZBEKISTAN
NORTHEASTERN PART OF CENTRAL KYZYLKUM
2 KM WEST-SOUTH-WEST OF THE SETTLEMENT
BESHBULAK
LATE TURONIAN
UPPERMOST GALLIC SUBEPOCH- SANTONIAN STAGE
LOWER SENONIAN SUBEPOCH
LOWER GULF EPOCH
EARLY LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: ORNITHISCHIFORMES COOPER, 1985
MAGNORDER: ORNITHISCHIA SEELEY, 1888
SUPERORDER: NEORNITHISCHIA COOPER, 1985
GRANDORDER: ORNITHOPODA MARSH, 1871
MICRORDER: EUORNITHOPODA SERENO, 1986
ORDER: IGUANODONTIA DOLLO, 1888
INFRAORDER: STYRACOSTERNA SERENO, 1986
PARVORDER: HADROSAURIFORMES SERENO, 1997
SUPERFAMILY: HADROSAUROIDEA COPE, 1869 (SENSU SERENO, 1986)
Family: HADROSAURIDAE COPE, 1869
Gen. sp indet.
_____________________________________________________________________________________
UZBEKISTAN
NORTHERN SLOPE OF THE BUKANTAU RIDGE
3-3.5 KM WEST OF THE DZHAMANURU SETTLEMENT
DZHAMANURU
LATE TURONIAN
UPPERMOST GALLIC SUBEPOCH- SANTONIAN STAGE
LOWER SENONIAN SUBEPOCH
LOWER GULF EPOCH
EARLY LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
Incertae sedis
Gen. sp indet.
COHORT: ORNITHISCHIFORMES COOPER, 1985
MAGNORDER: ORNITHISCHIA SEELEY, 1888
SUPERORDER: NEORNITHISCHIA COOPER, 1985
GRANDORDER: ORNITHOPODA MARSH, 1871
MICRORDER: EUORNITHOPODA SERENO, 1986
ORDER: IGUANODONTIA DOLLO, 1888
INFRAORDER: STYRACOSTERNA SERENO, 1986
PARVORDER: HADROSAURIFORMES SERENO, 1997
SUPERFAMILY: HADROSAUROIDEA COPE, 1869 (SENSU SERENO, 1986)
Family: HADROSAURIDAE COPE, 1869
Gen. sp indet.
_____________________________________________________________________________________
UZBEKISTAN
CENTRAL KYZYLKUM
SERIES OF SITES 3-4 KM WEST OF THE DAUGYZTAU SETTLEMENT
DAUGYZTAU II
LATE TURONIAN
UPPERMOST GALLIC SUBEPOCH- SANTONIAN STAGE
LOWER SENONIAN SUBEPOCH
LOWER GULF EPOCH
EARLY LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
Incertae sedis
Gen. sp indet.
COHORT: ORNITHISCHIFORMES COOPER, 1985
MAGNORDER: ORNITHISCHIA SEELEY, 1888
SUPERORDER: NEORNITHISCHIA COOPER, 1985
GRANDORDER: ORNITHOPODA MARSH, 1871
MICRORDER: EUORNITHOPODA SERENO, 1986
ORDER: IGUANODONTIA DOLLO, 1888
INFRAORDER: STYRACOSTERNA SERENO, 1986
PARVORDER: HADROSAURIFORMES SERENO, 1997
SUPERFAMILY: HADROSAUROIDEA COPE, 1869 (SENSU SERENO, 1986)
Family: HADROSAURIDAE COPE, 1869
Gen. sp indet.
_____________________________________________________________________________________
UZBEKISTAN
CENTRAL KYZYLKUM
NORTHWESTERN EDGE OF THE KARAKATA DEPRESSION,
KOSKUDUK
LATE TURONIAN
UPPERMOST GALLIC SUBEPOCH
SANTONIAN STAGE
LOWER SENONIAN SUBEPOCH
LOWER GULF EPOCH
EARLY LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
Incertae sedis
Gen. sp indet.
COHORT: ORNITHISCHIFORMES COOPER, 1985
MAGNORDER: ORNITHISCHIA SEELEY, 1888
SUPERORDER: NEORNITHISCHIA COOPER, 1985
GRANDORDER: ORNITHOPODA MARSH, 1871
MICRORDER: EUORNITHOPODA SERENO, 1986
ORDER: IGUANODONTIA DOLLO, 1888
INFRAORDER: STYRACOSTERNA SERENO, 1986
PARVORDER: HADROSAURIFORMES SERENO, 1997
SUPERFAMILY: HADROSAUROIDEA COPE, 1869 (SENSU SERENO, 1986)
Family: HADROSAURIDAE COPE, 1869
Gen. sp indet.
_____________________________________________________________________________________
UZBEKISTAN
CENTRAL KYZYLKUM
MOUND APPROXIMATELY 60 KM SOUTH OF KULKUDUK AND FROM THE BUKANTAU RIDGE
TAIKARSHI
BASAL PART OF THE BISSEKTY FORMATION
LATE TURONIAN
UPPERMOST GALLIC SUBEPOCH
LOWER GULF EPOCH
EARLY LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
Incertae sedis
Gen. sp indet.
COHORT: ORNITHISCHIFORMES COOPER, 1985
MAGNORDER: ORNITHISCHIA SEELEY, 1888
SUPERORDER: NEORNITHISCHIA COOPER, 1985
GRANDORDER: ORNITHOPODA MARSH, 1871
MICRORDER: EUORNITHOPODA SERENO, 1986
ORDER: IGUANODONTIA DOLLO, 1888
INFRAORDER: STYRACOSTERNA SERENO, 1986
PARVORDER: HADROSAURIFORMES SERENO, 1997
SUPERFAMILY: HADROSAUROIDEA COPE, 1869 (SENSU SERENO, 1986)
Family: HADROSAURIDAE COPE, 1869
Incertae sedis
Incertae sedis
SUPERORDER: EURYOPDA SERENO, 1986
GRANDORDER: ARMATOSAURIA ZHAO, 1983 (emend)
ORDER: ANKYLOSAURIA OSBORN, 1923
INFRAORDER: ANKYLOSAURA OSBORN, 1923
Family: ANKYLOSAURIDAE Brown, 1908
Incertae sedis
Gen. sp indet.
_____________________________________________________________________________________
UZBEKISTAN
CENTRAL KYZYLKUM
TAMDY-TRUBA
BISSEKTY FORMATION?
LATE TURONIAN STAGE
UPPERMOST GALLIC SUBEPOCH- EARLY CONIACIAN STAGE
LOWER SENONIAN SUBEPOCH
LOWER GULF EPOCH
EARLY LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: ORNITHISCHIFORMES COOPER, 1985
MAGNORDER: ORNITHISCHIA SEELEY, 1888
SUPERORDER: NEORNITHISCHIA COOPER, 1985
GRANDORDER: ORNITHOPODA MARSH, 1871
MICRORDER: EUORNITHOPODA SERENO, 1986
ORDER: IGUANODONTIA DOLLO, 1888
INFRAORDER: STYRACOSTERNA SERENO, 1986
PARVORDER: HADROSAURIFORMES SERENO, 1997
SUPERFAMILY: HADROSAUROIDEA COPE, 1869 (SENSU SERENO, 1986)
Family: HADROSAURIDAE COPE, 1869
Subfamily: LAMBEOSAURINAE Parks, 1923
Gen. sp indet.
SUPERORDER: EURYOPDA SERENO, 1986
GRANDORDER: ARMATOSAURIA ZHAO, 1983 (emend)
ORDER: ANKYLOSAURIA OSBORN, 1923
INFRAORDER: ANKYLOSAURA OSBORN, 1923
Family: ANKYLOSAURIDAE Brown, 1908
Incertae sedis
Gen. sp indet.
_____________________________________________________________________________________
UZBEKISTAN
KYZYLKUM DESERT
DZHARAKUDUK
BISSEKTY FORMATION
TURONIAN STAGE
UPPER GALLIC SUBEPOCH
LOWER GULF EPOCH
MIDDLE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: CHELONOMORPHA KUHN, 1960
SUPERCOHORT: TESTUDINES LINNAEUS, 1758
COHORT: CASICHELYDIA emend GAFFNEY & MEYLAN, 1988
MAGANORDER: CRYPTODIRA GAFFNEY & MEYLAN, 1988
SUPERORDER: SELMACRYPTODIRA GAFFNEY, HUTCHISON, JENKINS & MEEKR, 1987
GRANDORDER: DAIOCRYPTODIRA GAFFNEY & MEYLAN, 1988
MICRORDER: CENTROCRYPTODIRA GAFFNEY & MEYLAN, 1988
ORDER: POLYCRPTODIRA GAFFNEY & MEYLAN, 1988
SUBORDER: PROCOELOCRYPTODIRA GAFFNEY & MEYLAN, 1988
INFRAORDER: CHELOMACRYPTODIRA GAFFNEY & MEYLAN, 1988
PARVORDER: TRIONYCHIDA GRAY, 1870
Superfamily: TRIONYCHIDEA Bell, 1828 (sensu Gray, 1870)
Family: TRIONYCHIDAE Bell, 1828
Subfamily: TRIONYCHINAE Bell, 1828
Tribe: TRIONYCHINI Bell,1828
Subtribe: APALONINA MEYLAN, 1987
Aspidertoides sp
CLASS: MAMMALIA LINNAEUS, 1758
SUBCLASS: THERIIFORMES (ROWE, 1988) McKENNA & BELL, 1997
INFRACLASS: ALLOTHERIA (MARSH, 1880) McKENNA & BELL, 1997
ORDER: MULTITUBERCULATA COPE, 1884
Incertae sedis
Gen. sp indet.
SUBCLASS: PROTOTHERIA GILL, 1872
INFRACLASS: TRECHNOTHERIA McKENNA, 1975
LEGION: CLADOTHERIA McKENNA, 1975
SUBLEGION: ZATHERIA MCKENNA, 1975
INFRALEGION: TRIBOSPHENIDA (McKENNA, 1975) McKENNA & BELL, 1997
SUPERCOHORT: THERIA (PARKER & HASWELL, 1987) McKENNA & BELL, 1997
COHORT: MARSUPIALIA (ILLIGER, 1811) McKENNA & BELL, 1997
Incertae sedis
Metherian group 1
Metherian group 2
Metherian group 3
COHORT: PLACENTALIA (OWEN, 1837) McKENNA & BELL, 1997
Incertae sedis
Eutherian group 1
Gen. sp indet.
MAGNORDER: EPITHERIA (McKENNA, 1975) McKENNA & BELL, 1997
SUPERORDER: PREPTOTHERIA McKENNA, 1975
GRANDORDER: ANAGLIDA SZALAY & McKENNA, 1971
Family: ZALAMBDALESTIDAE Gregory & Simpson, 1926
Gen. sp indet.
GRANDORDER: UNGULATOMORPHA ARCHIBALD, 1996
Family: ZHELESTIDAE Nesov, 1990
Group 1
Group 2
Group 3
Group 4
Gen. sp indet.
_____________________________________________________________________________________
UZBEKISTAN
CENTRAL KYZYLKUM DESERT
NAVOI VIOLYAT DISTRICT
DZHARAKUDUK
BISSEKTY FORMATION
MIDDLE-LATE TURONIAN STAGE
UPPERMOST GALLIC SUBEPOCH
LOWER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
AVEPODA PAUL, 2002
SUPERORDER: NEOTHEROPODA BAKKER, 1986
GRANDORDER: TETANURAE GAUTHIER, 1986
MICRORDER: COELUROSAURIA HUENE, 1914
ORDER: MANIRAPTORFORMES HOLTZ, 1996
AVETHEROPODA PAUL, 2001
SUBORDER: COELURIA MARSH, 1881
Superfamily: TYRANNOSAUROIDEA Osborn, 1906 (sensu Bonaparte et. al., 1990)
Incertae sedis
*Timurlengia euotica
AVEPECTORA PAUL, 2001
SUBORDER: MANIRAPTORA GAUTHIER, 1986
INFRAORDER: PARVES SERENO, 1997
PARVORDER: DEINONYCHOSAURIA COLBERT & D. A. RUSSELL, 1969
Family: DROMAEOSAURIDAE Russell, 1969
CLADE NAME: EUDROMAEOSAURIA LONGRIC & CURRIE, 2009
Incerate sediS
Gen. sp indet
COHORT: ORNITHISCHIFORMES COOPER, 1985
MAGNORDER: ORNITHISCHIA SEELEY, 1888
SUPERORDER: NEORNITHISCHIA COOPER, 1985
GRANDORDER: ORNITHOPODA MARSH, 1871
MICRORDER: EUORNITHOPODA SERENO, 1986
ORDER: IGUANODONTIA DOLLO, 1888
SUBORDER: ANKYLOPOLLEXIA SERENO, 1986
INFRAORDER: STYRACOSTERNA SERENO, 1986
SUPERFAMILY: HADROSAUROIDEA COPE, 1869 (SENSU SERENO, 1986)
Incertae sedis
*Levnesovia transoxiana
_____________________________________________________________________________________
UZBEKISTAN
CENTRAL KYZYLKUM
A RELATIVELY FLAT AREA WITH SHALLOW RAVINES NORTH-NORTH-EAST OF THE DZHARAKUDUK SETTLEMENT
DZHARAKUDUK I
UCHKUDUK FORMATION
LOWER PART OF THE EARLY TURONIAN STAGE
UPPERMOST GALLIC SUBEPOCH
LOWERMOST GULF EPOCH
EARLY LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
Incertae sedis
Gen sp indet.
COHORT: THEROPODA EMEND MARSH, 1881
Incertae sedis
Gen. sp indet.
COHORT: ORNITHISCHIFORMES COOPER, 1985
MAGNORDER: ORNITHISCHIA SEELEY, 1888
SUPERORDER: NEORNITHISCHIA COOPER, 1985
GRANDORDER: ORNITHOPODA MARSH, 1871
MICRORDER: EUORNITHOPODA SERENO, 1986
ORDER: IGUANODONTIA DOLLO, 1888
INFRAORDER: STYRACOSTERNA SERENO, 1986
PARVORDER: HADROSAURIFORMES SERENO, 1997
SUPERFAMILY: HADROSAUROIDEA COPE, 1869 (SENSU SERENO, 1986)
Family: HADROSAURIDAE COPE, 1869
Gen. sp indet.
_____________________________________________________________________________________
UZBEKISTAN
KIZYLKUM DESERT
DZHYRAKUUK
LOCALITY CBI-?
BISSEKTY FORMATION
CONIACIAN STAGE
LOWER SENONIAN SUBEPOCH
MIDDLE GULF EPOCH
EARLY CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: AVEMETATARSALIA BENTON, 1999
COHORT: PROLACERTIFORMES CAMP, 1945 NOVA
SUPERORDER: TAPINOPLATIA PETERS, 2000
GRANDORDER: CHARACIOPODA PETERS, 2000
MICRORDER: FENESTRASAURIA PETERS, 2000
ORDER: PTEROSAURIA KAUP, 1834
SUBORDER: EUPTERODACTYLOIDEA BENNETT, 1994
Superfamily: AZHDARCHIDEA Nesov, 1984
Family: AZHDARCHIDAE Nesov, 1984 emend Padian, 1986
Azhdarcho lancicollis
CLASS: MAMMALIA LINNAEUS, 1758
SUBCLASS: PROTOTHERIA GILL, 1872
INFRACLASS: TRECHNOTHERIA McKENNA, 1975
LEGION: CLADOTHERIA McKENNA, 1975
SUBLEGION: ZATHERIA MCKENNA, 1975
INFRALEGION: TRIBOSPHENIDA (McKENNA, 1975) McKENNA & BELL, 1997
SUPERCOHORT: THERIA (PARKER & HASWELL, 1987) McKENNA & BELL, 1997
COHORT: PLACENTALIA (OWEN, 1837) McKENNA & BELL, 1997
MAGNORDER: EPITHERIA (McKENNA, 1975) McKENNA & BELL, 1997
ORDER: KENNALESTIDA McKENNA, 1975
Incertae sedis
Bulaklestes kezbe
SUPERORDER: PREPTOTHERIA McKENNA, 1975
GRANDORDER: UNGULATOMORPHA ARCHIBALD, 1996
Family: ZHELESTIDAE Nesov, 1990
Gen. sp indet.
_____________________________________________________________________________________
UZBEKISTAN
KIZYLKUM DESERT
DZHYRAKUUK
LOCALITY CBI-
BISSEKTY FORMATION
TURONIAN STAGE
UPPERMOST GALLIC SUBEPOCH
LOWER GULF EPOCH
EARLY CRETACEOUS
TEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
AVEPODA PAUL, 2002
SUPERORDER: NEOTHEROPODA BAKKER, 1986
GRANDORDER: TETANURAE GAUTHIER, 1986
MICRORDER: COELUROSAURIA HUENE, 1914
ORDER: SEGNOSAURIA BARSBOLD & PERLE, 1980
Superfamily: THERIZINOSAUROIDEA Maleev, 1954
Incertae sedis
Gen. sp indet.
ORDER: MANIRAPTORFORMES HOLTZ, 1996
AVETHEROPODA PAUL, 200
AVEPECTORA PAUL, 2001
SUBORDER: ORNITHOMIMOSAURIA BARSBOLD, 1976
Family: ORNITHOMIMIDAE Marsh, 1890
Incertae sedis
Gen. sp indet.
SUBORDER: MANIRAPTORA GAUTHIER, 1986
INFRAORDER: PARVES SERENO, 1997
PARVORDER: DEINONYCHOSAURIA COLBERT & D. A. RUSSELL, 1969
Family: TROODONTIDAE Gilmore, 1924
Urbacodon sp
_____________________________________________________________________________________
UZBEKISTAN
KIZYLKUM DESERT
DZHYRAKUUK
LOCALITY CBI-1
BISSEKTY FORMATION
CONIACIAN STAGE
LOWER SENONIAN SUBEPOCH
MIDDLE GULF EPOCH
EARLY CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 194
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
AVEPODA PAUL, 2002
SUPERORDER: NEOTHEROPODA BAKKER, 1986
GRANDORDER: TETANURAE GAUTHIER, 1986
MICRORDER: COELUROSAURIA HUENE, 1914
ORDER: MANIRAPTORFORMES HOLTZ, 1996
AVETHEROPODA PAUL, 2001
SUBORDER: COELURIA MARSH, 1881
Superfamily: TYRANNOSAUROIDEA Osborn, 1906 (sensu Bonaparte et. al., 1990)
Family: ALIORAMIDAE Olshevsky, Ford & Yamamoto, 1995
Alectrosaurus sp
_____________________________________________________________________________________
UZBEKISTAN
KYZYLKUM DESERT
CBI-2
DZHYRAKUDUK LOCALITY
BISSEKTY FORMATION
CONIACIAN STAGE
LOWER SENONIAN SUBEPOCH
MIDDLE GULF EPOCH
EARLY CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 184
COHORT: ORNITHISCHIFORMES COOPER, 1985
MAGNORDER: ORNITHISCHIA SEELEY, 1888
COHORT: ORNITHISCHIFORMES COOPER, 1985
MAGNORDER: ORNITHISCHIA SEELEY, 1888
SUPERORDER: NEORNITHISCHIA COOPER, 1985
GRANDORDER: ORNITHOPODA MARSH, 1871
MICRORDER: EUORNITHOPODA SERENO, 1986
ORDER: IGUANODONTIA DOLLO, 1888
INFRAORDER: STYRACOSTERNA SERENO, 1986
PARVORDER: HADROSAURIFORMES SERENO, 1997
SUPERFAMILY: HADROSAUROIDEA COPE, 1869 (SENSU SERENO, 1986)
Family: HADROSAURIDAE COPE, 1869
Incertae sedis
Gilmoreosaurus arkhangelskyi
_____________________________________________________________________________________
UZBEKISTAN
KIZYLKUM DESERT
DZHYRAKUUK
LOCALITY CBI-4
BISSEKTY FORMATION
CONIACIAN STAGE
LOWER SENONIAN SUBEPOCH
MIDDLE GULF EPOCH
EARLY CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: AMPHIBIA LINNAEUS, 1758
SUBCLASS: LISSAMPHIBIA, HAECKEL, 1866?
SUPERORDER: BATRACHIA BRONGNIART, 1800
ORDER: CAUDATA SCOPOLI, 1777
Incertae sedis
*Bissektia nana
SUBORDER: PORTEOIDEA (NOBEL 1931) EMEND EDWARDS, 1976
Family: BATRACHOSAUROIDIDAE Auffenberg, 1958
*Mynbulakia surgayi
Gen. sp indet.
SUBORDER: AMBYSTOMATOIDEA NOBLE, 1931
Family: SCAPHERPETONTIDAE Auffenberg & Goin, 1959
*Eoscapherpeton asiaticum
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: CHELONOMORPHA KUHN, 1960
SUPERCOHORT: TESTUDINES LINNAEUS, 1758
COHORT: CASICHELYDIA emend GAFFNEY & MEYLAN, 1988
MAGANORDER: CRYPTODIRA GAFFNEY & MEYLAN, 1988
SUPERORDER: SELMACRYPTODIRA GAFFNEY, HUTCHISON, JENKINS & MEEKR, 1987
GRANDORDER: DAIOCRYPTODIRA GAFFNEY & MEYLAN, 1988
MICRORDER: CENTROCRYPTODIRA GAFFNEY & MEYLAN, 1988
ORDER: POLYCRPTODIRA GAFFNEY & MEYLAN, 1988
SUBORDER: PROCOELOCRYPTODIRA GAFFNEY & MEYLAN, 1988
INFRAORDER: CHELOMACRYPTODIRA GAFFNEY & MEYLAN, 1988
PARVORDER: TRIONYCHIDA GRAY, 1870
Superfamily: TRIONYCHIDEA Bell, 1828 (sensu Gray, 1870)
Family: ADOCIDAE Cope, 1870
Gen. sp indet.
SUBCLASS: DIAPSIDA OSBORN, 1903 OSBORN, 1903
INFRACLASS: NEODIAPSIDA BENTON, 1985
LEGION: LEPIDOSAUROMORPHA DUMERIL & BIBRON, 1839
SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)
MAGNORDER: SQUAMATA OPPEL, 1811
SUPERORDER: LACERTILIA OWEN, 1842
GRANDORDER: ASCALABOTA GILMORE, 1928
ORDER: IGUANIA LATREILLE, 1825
SUBORDER: ACRODONTIA COPE, 1864
INFRAORDER: CHAMAELEOMORPHA FURBRINGER, 1900
PARVORDER: RHIPTOGLOSSA WEIGMANN, 1834
Superfamily: MONGOLOCHAMPOIDEA ALIFANOV, 1993
Family: MONGOLOCHAMPOIDAE Alifanov, 1993
*Buckantaus crassidens
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: CRUROTARSI SERENO & ARCUCCI, 1990
COHORT: CROCODYLOTARSI BENTON & CLARK, 1988
MAGNORDER: SUCHIA KREBS, 1974
SUPERORDER: CROCODYLOMORPHA WALKER, 1968
GRANDORDER: CROCODILIFORMES HAY, 1930
Incertae sedis
Gen. sp indet.
SUPERCOHORT: AVEMETATARSALIA BENTON, 1999
COHORT: PROLACERTIFORMES CAMP, 1945 NOVA
SUPERORDER: TAPINOPLATIA PETERS, 2000
GRANDORDER: CHARACIOPODA PETERS, 2000
MICRORDER: FENESTRASAURIA PETERS, 2000
ORDER: PTEROSAURIA KAUP, 1834
SUBORDER: EUPTERODACTYLOIDEA BENNETT, 1994
Superfamily: AZHDARCHIDEA Nesov, 1984
Family: AZHDARCHIDAE Nesov, 1984 emend Padian, 1986
Azhdarcho lancicollis
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
AVEPODA PAUL, 2002
SUPERORDER: NEOTHEROPODA BAKKER, 1986
GRANDORDER: TETANURAE GAUTHIER, 1986
MICRORDER: COELUROSAURIA HUENE, 1914
ORDER: SEGNOSAURIA BARSBOLD & PERLE, 1980
Superfamily: THERIZINOSAUROIDEA Maleev, 1954
Incertae sedis
Gen. sp indet.
ORDER: MANIRAPTORFORMES HOLTZ, 1996
AVETHEROPODA PAUL, 2001
AVEPECTORA PAUL, 2001
SUBORDER: ORNITHOMIMOSAURIA BARSBOLD, 1976
Family: ORNITHOMIMIDAE Marsh, 1890
Incertae sedis
Gen. sp indet.
SUBORDER: MANIRAPTORA GAUTHIER, 1986
INFRAORDER: PARVES SERENO, 1997
PARVORDER: DEINONYCHOSAURIA COLBERT & D. A. RUSSELL, 1969
Family: DROMAEOSAURIDAE Russell, 1969
Gen. sp indet.
Family: TROODONTIDAE Gilmore, 1924
Urbacodon sp
COHORT: ORNITHISCHIFORMES COOPER, 1985
MAGNORDER: ORNITHISCHIA SEELEY, 1888
SUPERORDER: NEORNITHISCHIA COOPER, 1985
GRANDORDER: ORNITHOPODA MARSH, 1871
MICRORDER: MARGINOCEPHALIA SERENO, 1986
ORDER: CERATOPIA MARSH, 1890
SUBORDER: NEOCERATOPIA SERENO, 1986
INFRAORDER: CORONOSAURIA SERENO, 1986
PARVORDER: CERATOPOMORPHA Wolfe & Kirkland, 1998 (emend)
Superfamily: CERATOPOIDEA Hay, 1902 (emend)
Incertae sedis
Turanoceratops tardabilis
SUPERORDER: EURYOPDA SERENO, 1986
GRANDORDER: ARMATOSAURIA ZHAO, 1983 (emend)
ORDER: ANKYLOSAURIA OSBORN, 1923
INFRAORDER: ANKYLOSAURA OSBORN, 1923
Family: ANKYLOSAURIDAE Brown, 1908
Incertae sedis
Gen. sp indet
CLASS: MAMMALIA LINNAEUS, 1758
Incertae sedis
Gen. sp indet.
SUBCLASS: PROTOTHERIA GILL, 1872
INFRACLASS: TRECHNOTHERIA McKENNA, 1975
LEGION: CLADOTHERIA McKENNA, 1975
SUBLEGION: ZATHERIA MCKENNA, 1975
INFRALEGION: TRIBOSPHENIDA (McKENNA, 1975) McKENNA & BELL, 1997
SUPERCOHORT: THERIA (PARKER & HASWELL, 1987) McKENNA & BELL, 1997
ORDER: DELTATHEROIDA KIELAN-JAWOROWSKA, 1982
Family: DELTATHERIDIIDAE Gregory & Simpson, 1926
Sulestes sp
Gen. sp. indet.
COHORT: PLACENTALIA (OWEN, 1837) McKENNA & BELL, 1997
Incertae sedis
*Cretasorex arkhanselksi
MAGNORDER: EPITHERIA (McKENNA, 1975) McKENNA & BELL, 1997
ORDER: KENNALESTIDA McKENNA, 1975
Incertae sedis
Daulestes kulbeckensis
= Kumlestes olzha
= Daulestes olzhaMAGNORDER: EPITHERIA (McKENNA, 1975) McKENNA & BELL, 1997
SUPERORDER: PREPTOTHERIA McKENNA, 1975
Incertae sedis
Gen. sp indet.
GRANDORDER: UNGULATOMORPHA ARCHIBALD, 1996
Family: ZHELESTIDAE Nesov, 1990
Aspanlestes cf. A. aptap
*Kumsuperus avus
Parazhelestes minor
Genus nova
Family: GYPSONICTIPIDAE Van Valen, 1967
Kennalestes sp nov ?
Gen. sp indet.
_____________________________________________________________________________________
UZEBEKISTAN
CENTRAL KYZULKUM DESERT
SITE CBI-4a
MIDDLE BISSEKTY FORMATION
CONIACIAN STAGE
LOWER SENONIAN SUBEPOCH
LOWER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: AVEMETATARSALIA BENTON, 1999
COHORT: PROLACERTIFORMES CAMP, 1945 NOVA
SUPERORDER: TAPINOPLATIA PETERS, 2000
GRANDORDER: CHARACIOPODA PETERS, 2000
MICRORDER: FENESTRASAURIA PETERS, 2000
ORDER: PTEROSAURIA KAUP, 1834
SUBORDER: EUPTERODACTYLOIDEA BENNETT, 1994
Superfamily: AZHDARCHIDEA Nesov, 1984
Family: AZHDARCHIDAE Nesov, 1984 emend Padian, 1986
Azhdarcho lancicollis
CLASS: MAMMALIA LINNAEUS, 1758
SUBCLASS: PROTOTHERIA GILL, 1872
INFRACLASS: TRECHNOTHERIA McKENNA, 1975
LEGION: CLADOTHERIA McKENNA, 1975
SUBLEGION: ZATHERIA MCKENNA, 1975
INFRALEGION: TRIBOSPHENIDA (McKENNA, 1975) McKENNA & BELL, 1997
SUPERCOHORT: THERIA (PARKER & HASWELL, 1987) McKENNA & BELL, 1997
COHORT: PLACENTALIA (OWEN, 1837) McKENNA & BELL, 1997
MAGNORDER: EPITHERIA (McKENNA, 1975) McKENNA & BELL, 1997
ORDER: KENNALESTIDA McKENNA, 1975
Incertae sedis
Bulaklestes kezbe
SUPERORDER: LEPTICTIDAE McKENNA, 1975
Incertae sedis
*Paranyctoides quadrans
= *Sailestes quadrans
_____________________________________________________________________________________
UZBEKISTAN
CENTRAL KIZYLKKUM DESERT
DZHYRAKUDUK
CBI-4b
MIDDLE OF THE BISSEKTY FORMATION
CONIACIAN STAGE
LOWER SENONIAN SUBEPOCH
MIDDLE GULF EPOCH
EARLY CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: AMPHIBIA LINNAEUS, 1758
SUBCLASS: LISSAMPHIBIA, HAECKEL, 1866?
SUPERORDER: SALIENTIA LAURENTI, 1768
ORDER: ANURA RAFINESQUE, 1815
SUBORDER: DISCOGLOSSOIDEI SOKOL, 1977
Family: DISCOGLOSSIDAE Guenther, 1859
Subfamily: GOBIATINAE Rocek & Nesov, 1993
*Gobiates bogatchovi
*Gobiates fritschi
*Gobiates spinari
*Gobiates furcatusSubfamily: DISCOGLOSSINAE Gunther, 1859
*Estesina elegans
CLASS: MAMMALIA LINNAEUS, 1758
SUBCLASS: PROTOTHERIA GILL, 1872
INFRACLASS: TRECHNOTHERIA McKENNA, 1975
LEGION: CLADOTHERIA McKENNA, 1975
SUBLEGION: ZATHERIA MCKENNA, 1975
INFRALEGION: TRIBOSPHENIDA (McKENNA, 1975) McKENNA & BELL, 1997
SUPERCOHORT: THERIA (PARKER & HASWELL, 1987) McKENNA & BELL, 1997
ORDER: DELTATHEROIDA KIELAN-JAWOROWSKA, 1982
Family: DELTATHERIDIIDAE Gregory & Simpson, 1926
*Sulestes karakshi
= Marsasia aenigmaCOHORT: PLACENTALIA (OWEN, 1837) McKENNA & BELL, 1997
MAGNORDER: EPITHERIA (McKENNA, 1975) McKENNA & BELL, 1997
ORDER: KENNALESTIDA McKENNA, 1975
Incertae sedis
Daulestes kulbeckensis
= Daulestes olzha
= *Kumlestes olzhaSUPERORDER: PREPTOTHERIA McKENNA, 1975
GRANDORDER: UNGULATOMORPHA ARCHIBALD, 1996
Family: ZHELESTIDAE Nesov, 1990
Subfamily: ZHELESTINAE Nesov, 1985
Eoungulatum kudukensis
= Eoungulatum kudukensisZhelestes temirkazyk = Kumsuperus avus
Family: KULBECKIIDAE NESOV, 1993
*Kulbeckia kulbecke
_____________________________________________________________________________________
UZBEKISTAN
CENTRAL KIZYLKKUM DESERT
DZHYRANKUDUK
CBI-4c
UPPER BISSEKTY FORMATION
CONIACIAN STAGE
LOWER SENONIAN SUBEPOCH
MIDDLE GULF EPOCH
EARLY CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
INFRACLASS: NEODIAPSIDA BENTON, 1985
LEGION: LEPIDOSAUROMORPHA DUMERIL & BIBRON, 1839
SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)
MAGNORDER: SQUAMATA OPPEL, 1811
SUPERORDER: LACERTILIA OWEN, 1842
GRANDORDER: ASCALABOTA GILMORE, 1928
ORDER: IGUANIA LATREILLE, 1825
SUBORDER: ACRODONTIA COPE, 1864
INFRAORDER: CHAMAELEOMORPHA FURBRINGER, 1900
PARVORDER: RHIPTOGLOSSA WEIGMANN, 1834
Superfamily: PRISCAGAMOIDEA Borsuk-Bialynicka & Moody, 1984
Family: PRISCAGAMIDAE Borsuk-Bialynicka & Moody, 1984
Subfamily: PRISCAGAMINAE Borsuk-Bialynicka & Moody, 1984
Tribe: PRISCAGAMINI Borsuk-Bialynicka & Moody, 1984 (sensu Alifanov, 1996)
*Sheikhia priscagama
___________________________________________________________________________________
UZBEKISTAN
KIZYLKUM DESERT
DZHYRAKUUK
LOCALITY CBI-4e
BISSEKTY FORMATION
CONIACIAN STAGE
LOWER SENONIAN SUBEPOCH
MIDDLE GULF EPOCH
EARLY CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: AVEMETATARSALIA BENTON, 1999
COHORT: PROLACERTIFORMES CAMP, 1945 NOVA
SUPERORDER: TAPINOPLATIA PETERS, 2000
GRANDORDER: CHARACIOPODA PETERS, 2000
MICRORDER: FENESTRASAURIA PETERS, 2000
ORDER: PTEROSAURIA KAUP, 1834
SUBORDER: EUPTERODACTYLOIDEA BENNETT, 1994
Superfamily: AZHDARCHIDEA Nesov, 1984
Family: AZHDARCHIDAE Nesov, 1984 emend Padian, 1986
Azhdarcho lancicollis
CLASS: MAMMALIA LINNAEUS, 1758
SUBCLASS: PROTOTHERIA GILL, 1872
INFRACLASS: TRECHNOTHERIA McKENNA, 1975
LEGION: CLADOTHERIA McKENNA, 1975
SUBLEGION: ZATHERIA MCKENNA, 1975
INFRALEGION: TRIBOSPHENIDA (McKENNA, 1975) McKENNA & BELL, 1997
SUPERCOHORT: THERIA (PARKER & HASWELL, 1987) McKENNA & BELL, 1997
ORDER: DELTATHEROIDA KIELAN-JAWOROWSKA, 1982
Family: DELTATHERIDIIDAE Gregory & Simpson, 1926
Sulestes karakshi
COHORT: PLACENTALIA (OWEN, 1837) McKENNA & BELL, 1997
MAGNORDER: EPITHERIA (McKENNA, 1975) McKENNA & BELL, 1997
ORDER: KENNALESTIDA McKENNA, 1975
Incertae sedis
Bulaklestes kezbe
Daulestes inobservabilis = Taslestes inobservabilis
= *Kennalestes? uzbekistanensisUchkudukodon nessovi
= Daulestes nessoviSUPERORDER: PREPTOTHERIA McKENNA, 1975
GRANDORDER: UNGULATOMORPHA ARCHIBALD, 1996
Family: ZHELESTIDAE Nesov, 1990
Gen. sp indet.
Subfamily: ZHELESTINAE Nesov, 1985
Aspanlestes aptap
Eoungulatum kudukensis
Parazhelestes mynbulakensi
Parazhelestes robustusZhelestes temirkazyk
Family: KULBECKIIDAE NESOV, 1993
Kulbeckia kulbecke
_____________________________________________________________________________________
UZBEKISTAN
CENTRAL KIZYLKKUM DESERT
DZHYRAKUDUK
CBI-4g
MIDDLE OF THE BISSEKTY FORMATION
CONIACIAN STAGE
LOWER SENONIAN SUBEPOCH
MIDDLE GULF EPOCH
EARLY CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: AVEMETATARSALIA BENTON, 1999
COHORT: PROLACERTIFORMES CAMP, 1945 NOVA
SUPERORDER: TAPINOPLATIA PETERS, 2000
GRANDORDER: CHARACIOPODA PETERS, 2000
MICRORDER: FENESTRASAURIA PETERS, 2000
ORDER: PTEROSAURIA KAUP, 1834
SUBORDER: EUPTERODACTYLOIDEA BENNETT, 1994
Superfamily: AZHDARCHIDEA Nesov, 1984
Family: AZHDARCHIDAE Nesov, 1984 emend Padian, 1986
Azhdarcho lancicollis
_____________________________________________________________________________________
UZBEKISTAN
CENTRAL KIZYLKKUM DESERT
DZHYRAKUDUK
CBI-4V
MIDDLE OF THE BISSEKTY FORMATION
CONIACIAN STAGE
LOWER SENONIAN SUBEPOCH
MIDDLE GULF EPOCH
EARLY CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: AMPHIBIA LINNAEUS, 1758
SUBCLASS: LISSAMPHIBIA, HAECKEL, 1866?
SUPERORDER: BATRACHIA BRONGNIART, 1800
ORDER: CAUDATA SCOPOLI, 1777
Incertae sedis
Gen. sp indet.
SUPERORDER: SALIENTIA LAURENTI, 1768
ORDER: ANURA RAFINESQUE, 1815
SUBORDER: DISCOGLOSSOIDEI SOKOL, 1977
Family: DISCOGLOSSIDAE Guenther, 1859
Subfamily: GOBIATINAE Rocek & Nesov, 1993
*Gobiates sosedkoi
= Eopelobates sosedkoi
Gobiates kizylkumensisCLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
INFRACLASS: NEODIAPSIDA BENTON, 1985
LEGION: LEPIDOSAUROMORPHA DUMERIL & BIBRON, 1839
SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)
MAGNORDER: SQUAMATA OPPEL, 1811
SUPERORDER: LACERTILIA OWEN, 1842
Incertae sedis
Gen. sp indet.
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: AVEMETATARSALIA BENTON, 1999
COHORT: PROLACERTIFORMES CAMP, 1945 NOVA
SUPERORDER: TAPINOPLATIA PETERS, 2000
GRANDORDER: CHARACIOPODA PETERS, 2000
MICRORDER: FENESTRASAURIA PETERS, 2000
ORDER: PTEROSAURIA KAUP, 1834
SUBORDER: EUPTERODACTYLOIDEA BENNETT, 1994
Superfamily: AZHDARCHIDEA Nesov, 1984
Family: AZHDARCHIDAE Nesov, 1984 emend Padian, 1986
Azhdarcho lancicollis
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
AVEPODA PAUL, 2002
SUPERORDER: NEOTHEROPODA BAKKER, 1986
GRANDORDER: TETANURAE GAUTHIER, 1986
MICRORDER: COELUROSAURIA HUENE, 1914
MICRORDER: COELUROSAURIA HUENE, 1914
ORDER: MANIRAPTORFORMES HOLTZ, 1996
AVETHEROPODA PAUL, 2002
AVEPECTORA PAUL, 2002
SUBORDER: MANIRAPTORA GAUTHIER, 1986
INFRAORDER: PARVES SERENO, 1997
PARVORDER: DEINONYCHOSAURIA COLBERT & D. A. RUSSELL, 1969
Family: TROODONTIDAE Gilmore, 1924
Urbacodon sp
CLASS: MAMMALIA LINNAEUS, 1758
SUBCLASS: PROTOTHERIA GILL, 1872
INFRACLASS: TRECHNOTHERIA McKENNA, 1975
LEGION: CLADOTHERIA McKENNA, 1975
SUBLEGION: ZATHERIA MCKENNA, 1975
INFRALEGION: TRIBOSPHENIDA (McKENNA, 1975) McKENNA & BELL, 1997
SUPERCOHORT: THERIA (PARKER & HASWELL, 1987) McKENNA & BELL, 1997
Incertae sedis
Gen. sp indet.
ORDER: DELTATHEROIDA KIELAN-JAWOROWSKA, 1982
Family: DELTATHERIDIIDAE Gregory & Simpson, 1926
Sulestes karakshi
Gen. sp indet.
COHORT: PLACENTALIA (OWEN, 1837) McKENNA & BELL, 1997
MAGNORDER: EPITHERIA (McKENNA, 1975) McKENNA & BELL, 1997
ORDER: KENNALESTIDA McKENNA, 1975
Incertae sedis
*Bulaklestes kezbe
SUPERORDER: PREPTOTHERIA McKENNA, 1975
GRANDORDER: UNGULATOMORPHA ARCHIBALD, 1996
Family: ZHELESTIDAE Nesov, 1990
Subfamily: ZHELESTINAE Nesov, 1985
Aspanlestes aptap
Eoungulatum kudukensis
Parazhelestes robustus
Zhelestes temirkazyk = Kumsuperus avus
= Sorlestes budan
= Zalambdalestes? mynbulakensisGen. sp indet.
_____________________________________________________________________________________
UZBEKISTAN
UZBEKISTAN
CENTRAL KYZYLKUM DESERT
NORTH OF
SITE CBI-4V
DZHYRAKUDUK LOCALITY
LOWER PART OF BISSEKTY FORMATION
UPPER TURONIAN STAGE
UPPERMOST GALLIC SUBEPOCH
LOWER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: AMPHIBIA LINNAEUS, 1758
SUBCLASS: LISSAMPHIBIA, HAECKEL, 1866?
ORDER: CAUDATA SCOPOLI, 1777
Incertae sedis
Nesovtriton mynbulakensis
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
AVEPODA PAUL, 2002
SUPERORDER: NEOTHEROPODA BAKKER, 1986
GRANDORDER: TETANURAE GAUTHIER, 1986
MICRORDER: COELUROSAURIA HUENE, 1914
ORDER: MANIRAPTORFORMES HOLTZ, 1996
AVETHEROPODA PAUL, 2001
SUBORDER: COELURIA MARSH, 1881
Superfamily: TYRANNOSAUROIDEA Osborn, 1906 (sensu Bonaparte et. al., 1990)
Family: ALIORAMIDAE Olshevsky, Ford & Yamamoto, 1995
Alectrosaurus sp
COHORT: ORNITHISCHIFORMES COOPER, 1985
MAGNORDER: ORNITHISCHIA SEELEY, 1888
COHORT: ORNITHISCHIFORMES COOPER, 1985
MAGNORDER: ORNITHISCHIA SEELEY, 1888
SUPERORDER: NEORNITHISCHIA COOPER, 1985
GRANDORDER: ORNITHOPODA MARSH, 1871
MICRORDER: EUORNITHOPODA SERENO, 1986
ORDER: HYPSILOPHODONTIA COOPER, 1983
Family: HYPSILOPHODONTIDAE Dollo, 1882
Incertae sedis? Or new families? (pending further research)
Gen. sp indet
_____________________________________________________________________________________
UZBEKISTAN
KYZYLKUM DESERT
CBI-5
DZHYRAKUDUK LOCALITY
BISSEKTY FORMATION
CONIACIAN STAGE
LOWER SENONIAN SUBEPOCH
MIDDLE GULF EPOCH
EARLY CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: AVEMETATARSALIA BENTON, 1999
COHORT: PROLACERTIFORMES CAMP, 1945 NOVA
SUPERORDER: TAPINOPLATIA PETERS, 2000
GRANDORDER: CHARACIOPODA PETERS, 2000
MICRORDER: FENESTRASAURIA PETERS, 2000
ORDER: PTEROSAURIA KAUP, 1834
SUBORDER: EUPTERODACTYLOIDEA BENNETT, 1994
Superfamily: AZHDARCHIDEA Nesov, 1984
Family: AZHDARCHIDAE Nesov, 1984 emend Padian, 1986
Azhdarcho lancicollis
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
AVEPODA PAUL, 2002
SUPERORDER: NEOTHEROPODA BAKKER, 1986
GRANDORDER: TETANURAE GAUTHIER, 1986
MICRORDER: COELUROSAURIA HUENE, 1914
ORDER: SEGNOSAURIA BARSBOLD & PERLE, 1980
Family: SEGNOSAURIDAE Perle, 1979
Gen. sp indet.
ORDER: MANIRAPTORFORMES HOLTZ, 1996
AVETHEROPODA PAUL, 2001
SUBORDER: COELURIA MARSH, 1881
Superfamily: TYRANNOSAUROIDEA Osborn, 1906 (sensu Bonaparte et. al., 1990)
Incertae sedis
Timurlengia euotica
AVEPECTORA PAUL, 2001
SUBORDER: ORNITHOMIMOSAURIA BARSBOLD, 1976
Family: ORNITHOMIMIDAE Marsh, 1890
Incertae sedis
Gen. sp indet.
COHORT: ORNITHISCHIFORMES COOPER, 1985
MAGNORDER: ORNITHISCHIA SEELEY, 1888
COHORT: ORNITHISCHIFORMES COOPER, 1985
MAGNORDER: ORNITHISCHIA SEELEY, 1888
SUPERORDER: NEORNITHISCHIA COOPER, 1985
GRANDORDER: ORNITHOPODA MARSH, 1871
MICRORDER: EUORNITHOPODA SERENO, 1986
ORDER: IGUANODONTIA DOLLO, 1888
INFRAORDER: STYRACOSTERNA SERENO, 1986
PARVORDER: HADROSAURIFORMES SERENO, 1997
SUPERFAMILY: HADROSAUROIDEA COPE, 1869 (SENSU SERENO, 1986)
Family: HADROSAURIDAE COPE, 1869
Incertae sedis
Gilmoreosaurus arkhangelskyi
CLASS: MAMMALIA LINNAEUS, 1758
SUBCLASS: PROTOTHERIA GILL, 1872
INFRACLASS: TRECHNOTHERIA McKENNA, 1975
LEGION: CLADOTHERIA McKENNA, 1975
SUBLEGION: ZATHERIA MCKENNA, 1975
INFRALEGION: TRIBOSPHENIDA (McKENNA, 1975) McKENNA & BELL, 1997
SUPERCOHORT: THERIA (PARKER & HASWELL, 1987) McKENNA & BELL, 1997
COHORT: PLACENTALIA (OWEN, 1837) McKENNA & BELL, 1997
MAGNORDER: EPITHERIA (McKENNA, 1975) McKENNA & BELL, 1997
SUPERORDER: PREPTOTHERIA McKENNA, 1975
GRANDORDER: UNGULATOMORPHA ARCHIBALD, 1996
Family: ZHELESTIDAE Nesov, 1990
Subfamily: ZHELESTINAE Nesov, 1985
Parazhelestes mynbulakensi
_____________________________________________________________________________________
UZBEKISTAN
CENTRAL KYZYLKUM DESERT
NORTH OF
CBI-5A
DZHYRAKUDUK LOCALITY
LOWER PART OF BISSEKTY FORMATION
UPPER TURONIAN STAGE
UPPERMOST GALLIC SUBEPOCH
LOWER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: AVEMETATARSALIA BENTON, 1999
COHORT: PROLACERTIFORMES CAMP, 1945 NOVA
SUPERORDER: TAPINOPLATIA PETERS, 2000
GRANDORDER: CHARACIOPODA PETERS, 2000
MICRORDER: FENESTRASAURIA PETERS, 2000
ORDER: PTEROSAURIA KAUP, 1834
SUBORDER: EUPTERODACTYLOIDEA BENNETT, 1994
Superfamily: AZHDARCHIDEA Nesov, 1984
Family: AZHDARCHIDAE Nesov, 1984 emend Padian, 1986
Azhdarcho lancicollis
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
Incertae sedis
Gen. sp indet.
AVEPODA PAUL, 2002
SUPERORDER: NEOTHEROPODA BAKKER, 1986
GRANDORDER: TETANURAE GAUTHIER, 1986
MICRORDER: COELUROSAURIA HUENE, 1914
ORDER: SEGNOSAURIA BARSBOLD & PERLE, 1980
Superfamily: THERIZINOSAUROIDEA Maleev, 1954
Incertae sedis
Gen. sp indet.
ORDER: MANIRAPTORFORMES HOLTZ, 1996
AVETHEROPODA PAUL, 2001
SUBORDER: COELURIA MARSH, 1881
Superfamily: TYRANNOSAUROIDEA Osborn, 1906 (sensu Bonaparte et. al., 1990)
Incertae sedis
Timurlengia euotica
Family: TYRANNOSAURIDAE Osborn, 1906
Gen. sp indet.
Family: ALIORAMIDAE Olshevsky, Ford & Yamamoto, 1995
Alectrosaurus sp
AVEPECTORA PAUL, 2001
SUBORDER: ORNITHOMIMOSAURIA BARSBOLD, 1976
Family: ORNITHOMIMIDAE Marsh, 1890
Incertae sedis
Gen. sp indet.
SUBORDER: MANIRAPTORA GAUTHIER, 1986
INFRAORDER: PARVES SERENO, 1997
PARVORDER: DEINONYCHOSAURIA COLBERT & D. A. RUSSELL, 1969
Family: TROODONTIDAE Gilmore, 1924
Urbacodon sp
CLASS: AVES LINNAEUS, 1758
SUBCLASS: SAURORNITHES NICHOLSON, 1879
AVEBREVICAUDA PAUL, 2001
INFRACLASS: ORNITHOTHORACES NOVA
LEGION: ORNITHURAE NOVA
COHORT: ENANTIORNITHES NOVA
SUPERORDER: EUENANTIORNITHES CHIAPPE, 2002
ORDER: ALEXORNITHIFORMES BRODKORB, 1976
Family: ALEXORNITHIDAE Brodkorb, 1976
*Kizylkumavis cretacea
Family: ZHYRAORNITHIDAE Nesov, 1984
*Zhyraornis longunovi
CLASS: MAMMALIA LINNAEUS, 1758
SUBCLASS: PROTOTHERIA GILL, 1872
INFRACLASS: TRECHNOTHERIA McKENNA, 1975
LEGION: CLADOTHERIA McKENNA, 1975
SUBLEGION: ZATHERIA MCKENNA, 1975
INFRALEGION: TRIBOSPHENIDA (McKENNA, 1975) McKENNA & BELL, 1997
SUPERCOHORT: THERIA (PARKER & HASWELL, 1987) McKENNA & BELL, 1997
ORDER: DELTATHEROIDA KIELAN-JAWOROWSKA, 1982
Family: DELTATHERIDIIDAE Gregory & Simpson, 1926
Sulestes karakshi
COHORT: PLACENTALIA (OWEN, 1837) McKENNA & BELL, 1997
MAGNORDER: EPITHERIA (McKENNA, 1975) McKENNA & BELL, 1997
ORDER: KENNALESTIDA McKENNA, 1975
Incertae sedis
Daulestes inobservabilis
= Taslestes inobservabilisUchkudukodon nessovi
= *Daulestes nessoviSUPERORDER: PREPTOTHERIA McKENNA, 1975
GRANDORDER: UNGULATOMORPHA ARCHIBALD, 1996
Family: ZHELESTIDAE Nesov, 1990
Subfamily: ZHELESTINAE Nesov, 1985
Aspanlestes aptap
Parazhelestes mynbulakensi
Family: KULBECKIIDAE NESOV, 1993
*Kulbeckia kulbecke
SUPERORDER: LEPTICTIDAE McKENNA, 1975
Incertae sedis
Genus: Nova
_____________________________________________________________________________________
KIZYLKUM DESERT
DZHYRAKUUK
LOCALITY CBI-5a
BISSEKTY FORMATION
CONIACIAN STAGE
LOWER SENONIAN SUBEPOCH
MIDDLE GULF EPOCH
EARLY CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
Incertae sedis
Gen. sp indet.
AVEPODA PAUL, 2002
SUPERORDER: NEOTHEROPODA BAKKER, 1986
GRANDORDER: TETANURAE GAUTHIER, 1986
MICRORDER: COELUROSAURIA HUENE, 1914
ORDER: MANIRAPTORFORMES HOLTZ, 1996
AVETHEROPODA PAUL, 2001
SUBORDER: COELURIA MARSH, 1881
Superfamily: TYRANNOSAUROIDEA Osborn, 1906 (sensu Bonaparte et. al., 1990)
Family: ALIORAMIDAE Olshevsky, Ford & Yamamoto, 1995
Alectrosaurus sp
CLASS: AVES LINNAEUS, 1758
SUBCLASS: SAURORNITHES NICHOLSON, 1879
AVEBREVICAUDA PAUL, 2001
INFRACLASS: ORNITHOTHORACES NOVA
LEGION: ORNITHURAE NOVA
COHORT: ENANTIORNITHES NOVA
SUPERORDER: EUENANTIORNITHES CHIAPPE, 2002
ORDER: ALEXORNITHIFORMES BRODKORB, 1976
Family: ALEXORNITHIDAE Brodkorb, 1976
*Kizylkumavis cretacea
Family: ZHYRAORNITHIDAE Nesov, 1984
*Zhyraornis longunovi
CLASS: MAMMALIA LINNAEUS, 1758
SUBCLASS: PROTOTHERIA GILL, 1872
INFRACLASS: TRECHNOTHERIA McKENNA, 1975
LEGION: CLADOTHERIA McKENNA, 1975
SUBLEGION: ZATHERIA MCKENNA, 1975
INFRALEGION: TRIBOSPHENIDA (McKENNA, 1975) McKENNA & BELL, 1997
SUPERCOHORT: THERIA (PARKER & HASWELL, 1987) McKENNA & BELL, 1997
COHORT: PLACENTALIA (OWEN, 1837) McKENNA & BELL, 1997
MAGNORDER: EPITHERIA (McKENNA, 1975) McKENNA & BELL, 1997
ORDER: KENNALESTIDA McKENNA, 1975
Incertae sedis
Daulestes inobservabilis
= Taslestes inobservabilisUchkudukodon nessovi
= *Daulestes nessoviSUPERORDER: PREPTOTHERIA McKENNA, 1975
GRANDORDER: UNGULATOMORPHA ARCHIBALD, 1996
Family: ZHELESTIDAE Nesov, 1990
Aspanlestes aptap
Family: KULBECKIIDAE NESOV, 1993
*Kulbeckia kulbecke
SUPERORDER: LEPTICTIDAE McKENNA, 1975
Incertae sedis
Genus: Nova
_____________________________________________________________________________________
UZBEKISTAN
KIZYLKUM DESERT
DZHYRAKUUK
LOCALITY CBI-6
BISSEKTY FORMATION
TURONIAN STAGE
UPPERMOST GALLIC SUBEPOCH
LOWER GULF EPOCH
EARLY CRETACEOUS
TEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
AVEPODA PAUL, 2002
SUPERORDER: NEOTHEROPODA BAKKER, 1986
GRANDORDER: TETANURAE GAUTHIER, 1986
MICRORDER: COELUROSAURIA HUENE, 1914
ORDER: MANIRAPTORFORMES HOLTZ, 1996
AVETHEROPODA PAUL, 200
AVEPECTORA PAUL, 2001
SUBORDER: ORNITHOMIMOSAURIA BARSBOLD, 1976
Family: ORNITHOMIMIDAE Marsh, 1890
Incertae sedis
Gen. sp indet.
_____________________________________________________________________________________
UZBEKISTAN
DZHYRAKUDUK LOCALITY
SITE CBI-6a
CONIACIAN STAGE
LOWER SENONIAN SUBEPOCH
LOWER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
AVEPODA PAUL, 2002
SUPERORDER: NEOTHEROPODA BAKKER, 1986
GRANDORDER: TETANURAE GAUTHIER, 1986
MICRORDER: COELUROSAURIA HUENE, 1914
ORDER: MANIRAPTORFORMES HOLTZ, 1996
AVETHEROPODA PAUL, 2001
SUBORDER: COELURIA MARSH, 1881
Superfamily: TYRANNOSAUROIDEA Osborn, 1906 (sensu Bonaparte et. al., 1990)
Family: TYRANNOSAURIDAE Osborn, 1906
Gen. sp indet.
_____________________________________________________________________________________
UZBEKISTAN
KIZYLKUM DESERT
DZHYRAKUUK
LOCALITY CBI-7
BISSEKTY FORMATION
TURONIAN STAGE
UPPERMOST GALLIC SUBEPOCH
LOWER GULF EPOCH
EARLY CRETACEOUS
TEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
AVEPODA PAUL, 2002
SUPERORDER: NEOTHEROPODA BAKKER, 1986
GRANDORDER: TETANURAE GAUTHIER, 1986
MICRORDER: COELUROSAURIA HUENE, 1914
ORDER: MANIRAPTORFORMES HOLTZ, 1996
AVETHEROPODA PAUL, 200
AVEPECTORA PAUL, 2001
SUBORDER: ORNITHOMIMOSAURIA BARSBOLD, 1976
Family: ORNITHOMIMIDAE Marsh, 1890
Incertae sedis
Gen. sp indet.
_____________________________________________________________________________________
UZBEKISTAN
DZHYRAKUDUK LOCALITY
SITE CBI-7a
CONIACIAN STAGE
LOWER SENONIAN SUBEPOCH
LOWER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: AVEMETATARSALIA BENTON, 1999
COHORT: PROLACERTIFORMES CAMP, 1945 NOVA
SUPERORDER: TAPINOPLATIA PETERS, 2000
GRANDORDER: CHARACIOPODA PETERS, 2000
MICRORDER: FENESTRASAURIA PETERS, 2000
ORDER: PTEROSAURIA KAUP, 1834
SUBORDER: EUPTERODACTYLOIDEA BENNETT, 1994
Superfamily: AZHDARCHIDEA Nesov, 1984
Family: AZHDARCHIDAE Nesov, 1984 emend Padian, 1986
Azhdarcho lancicollis
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
Incertae sedis
Gen. sp indet.
COHORT: THEROPODA EMEND MARSH, 1881
AVEPODA PAUL, 2002
SUPERORDER: NEOTHEROPODA BAKKER, 1986
GRANDORDER: TETANURAE GAUTHIER, 1986
MICRORDER: COELUROSAURIA HUENE, 1914
ORDER: MANIRAPTORFORMES HOLTZ, 1996
AVETHEROPODA PAUL, 2001
SUBORDER: COELURIA MARSH, 1881
Superfamily: TYRANNOSAUROIDEA Osborn, 1906 (sensu Bonaparte et. al., 1990)
Family: ALIORAMIDAE Olshevsky, Ford & Yamamoto, 1995
Alectrosaurus sp
COHORT: ORNITHISCHIFORMES COOPER, 1985
MAGNORDER: ORNITHISCHIA SEELEY, 1888
SUPERORDER: NEORNITHISCHIA COOPER, 1985
GRANDORDER: ORNITHOPODA MARSH, 1871
MICRORDER: EUORNITHOPODA SERENO, 1986
ORDER: IGUANODONTIA DOLLO, 1888
INFRAORDER: STYRACOSTERNA SERENO, 1986
PARVORDER: HADROSAURIFORMES SERENO, 1997
SUPERFAMILY: HADROSAUROIDEA COPE, 1869 (SENSU SERENO, 1986)
Family: HADROSAURIDAE COPE, 1869
Incertae sedis
Gen. sp indet.
CLASS: MAMMALIA LINNAEUS, 1758
SUBCLASS: PROTOTHERIA GILL, 1872
INFRACLASS: TRECHNOTHERIA McKENNA, 1975
LEGION: CLADOTHERIA McKENNA, 1975
SUBLEGION: ZATHERIA MCKENNA, 1975
INFRALEGION: TRIBOSPHENIDA (McKENNA, 1975) McKENNA & BELL, 1997
SUPERCOHORT: THERIA (PARKER & HASWELL, 1987) McKENNA & BELL, 1997
COHORT: PLACENTALIA (OWEN, 1837) McKENNA & BELL, 1997
MAGNORDER: EPITHERIA (McKENNA, 1975) McKENNA & BELL, 1997
SUPERORDER: PREPTOTHERIA McKENNA, 1975
GRANDORDER: UNGULATA LINNAEUS, 1766
MICRORDER: EPARCTOCYONA McKENNA, 1975
ORDER: CONDYLARTHRA COPE, 1881
Family: PERIPTYCHIDAE Cope, 1882Gen. Sp indet.
Genus nova?
_____________________________________________________________________________________
UZBEKISTAN
CENTRAL KYZYLKUM DESERT
CBI-7A
AITYM FORMATION
SANTONIAN STAGE
LOWER SENONIAN SUBEPOCH
MIDDLE GULF EPOCH
LATE CRETACEOUS
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: ORNITHISCHIFORMES COOPER, 1985
MAGNORDER: ORNITHISCHIA SEELEY, 1888
SUPERORDER: NEORNITHISCHIA COOPER, 1985
GRANDORDER: ORNITHOPODA MARSH, 1871
MICRORDER: EUORNITHOPODA SERENO, 1986
ORDER: IGUANODONTIA DOLLO, 1888
INFRAORDER: STYRACOSTERNA SERENO, 1986
PARVORDER: HADROSAURIFORMES SERENO, 1997
SUPERFAMILY: HADROSAUROIDEA COPE, 1869 (SENSU SERENO, 1986)
Family: HADROSAURIDAE COPE, 1869
Incertae sedis
Gen. sp indet.
_____________________________________________________________________________________
UZBEKISTAN
CENTRAL KYZYLKUM DESERT
CBI-7A
DZHETYMTAU BED
CAMPANIAN STAGE
SENONIAN SUBEPOCH
MIDDLE GULF EPOCH
LATE CRETACEOUS
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
Incertae sedis
Gen. sp indet.
_____________________________________________________________________________________
UZBEKISTAN
KIZYLKUM DESERT
DZHYRAKUUK
LOCALITY CBI-14
BISSEKTY FORMATION
CONIACIAN STAGE
LOWER SENONIAN SUBEPOCH
MIDDLE GULF EPOCH
EARLY CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: AMPHIBIA LINNAEUS, 1758
SUBCLASS: LISSAMPHIBIA, HAECKEL, 1866?
SUPERORDER: BATRACHIA BRONGNIART, 1800
ORDER: CAUDATA SCOPOLI, 1777
Incertae sedis
*Nesovtriton mynbulakensis
SUBORDER: PROSIRENOIDEA ESTES, 1981
Family: PROSIRENIDAE Estes, 1969
Gen. sp indet.
SUBORDER: AMBYSTOMATOIDEA NOBLE, 1931
Family: SCAPHERPETONTIDAE Auffenberg & Goin, 1959
Gen sp indet.
SUPERORDER: SALIENTIA LAURENTI, 1768
ORDER: ANURA RAFINESQUE, 1815
SUBORDER: DISCOGLOSSOIDEI SOKOL, 1977
Family: DISCOGLOSSIDAE Guenther, 1859
Incertae sedis
Gen. sp indet.
Subfamily: GOBIATINAE Rocek & Nesov, 1993
*Gobiates dzhyrakudukensis
*Gobiates tatarinovi
*Gobiates asiaticus
*Gobiates kizylkumensis*Gobiatoides parvus
Gobiatoides spSubfamily: ALYTINAE Fitzinger, 1943
*Kizylkuma antiqua
Subfamily: DISCOGLOSSINAE Gunther, 1859
*Procerobatrachus paulus
*Saevesoederberghia egredia
SUBORDER: MESOBATRACHIA LAURENT, 1979
Superfamily: PELOBATOIDEA Stanius, 1856 , nom. corr. Spinar, 1971 (sensu Bolkay, 1919)
Family: PELOBATIDAE Stanius, 1856 , nom. corr. Spinar, 1971
Gen. sp indet.
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: CHELONOMORPHA KUHN, 1960
SUPERCOHORT: TESTUDINES LINNAEUS, 1758
COHORT: CASICHELYDIA emend GAFFNEY & MEYLAN, 1988
MAGANORDER: CRYPTODIRA GAFFNEY & MEYLAN, 1988
SUPERORDER: SELMACRYPTODIRA GAFFNEY, HUTCHISON, JENKINS & MEEKR, 1987
GRANDORDER: DAIOCRYPTODIRA GAFFNEY & MEYLAN, 1988
MICRORDER: CENTROCRYPTODIRA GAFFNEY & MEYLAN, 1988
ORDER: POLYCRPTODIRA GAFFNEY & MEYLAN, 1988
SUBORDER: PROCOELOCRYPTODIRA GAFFNEY & MEYLAN, 1988
INFRAORDER: CHELOMACRYPTODIRA GAFFNEY & MEYLAN, 1988
PARVORDER: TRIONYCHIDA GRAY, 1870
Superfamily: TRIONYCHIDEA Bell, 1828 (sensu Gray, 1870)
Family: TRIONYCHIDAE Bell, 1828
Khunnuchelys kizlkumensis
Family: ADOCIDAE Cope, 1870
Subfamily: SHACHEMYDINAE Khosatzky in Nesov & Khozatzkii, 1977
Shachemys sp
SUBCLASS: DIAPSIDA OSBORN, 1903
INFRACLASS: NEODIAPSIDA BENTON, 1985
LEGION: LEPIDOSAUROMORPHA DUMERIL & BIBRON, 1839
SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)
MAGNORDER: SQUAMATA OPPEL, 1811
SUPERORDER: LACERTILIA OWEN, 1842
GRANDORDER: ASCALABOTA GILMORE, 1928
ORDER: IGUANIA LATREILLE, 1825
SUBORDER: ACRODONTIA COPE, 1864
INFRAORDER: CHAMAELEOMORPHA FURBRINGER, 1900
PARVORDER: RHIPTOGLOSSA WEIGMANN, 1834
Superfamily: PRISCAGAMOIDEA Borsuk-Bialynicka & Moody, 1984
Family: PRISCAGAMIDAE Borsuk-Bialynicka & Moody, 1984
Subfamily: PRISCAGAMINAE Borsuk-Bialynicka & Moody, 1984
Tribe: PRISCAGAMINI Borsuk-Bialynicka & Moody, 1984 (sensu Alifanov, 1996)
Sheikhia sp
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: CRUROTARSI SERENO & ARCUCCI, 1990
COHORT: CROCODYLOTARSI BENTON & CLARK, 1988
MAGNORDER: SUCHIA KREBS, 1974
SUPERORDER: CROCODYLOMORPHA WALKER, 1968
GRANDORDER: CROCODILIFORMES HAY, 1930
MICRORDER: MESOEUROCROCODYLIA WHETSTONE & WHYBROW, 1983
ORDER: MESOSUCHIA EMEND HUXLEY, 1875
PARVORDER: METASUCHIA BENTON & CLARK, 1988
INFRAORDER: METAMESOSUCHIA HULKE, 1878
NEOSUCHIA CLARK, IN BENTON & CLARK, 1988
Family: PARALLIGAOTRIDAE Konjukova, 1954
*Shamosuchus borealis
= Kangajsuchus borealisORDER: EUSUCHIA emend HUXLEY, 1875
INFRAORDER: CROCODYLIA emend GMELIN, 1877
PARVORDER: BREVIROSTRES
Superfamily: CROCODILOIDEA Cuvier, 1807 (sensu Fitzinger, 1826)
Family: CROCODYLIDAE Cuvier, 1807
Subfamily: CROCODYLIDINAE Cuvier, 1807
*Tadzhikoesuchus kizylkumensis
SUPERCOHORT: AVEMETATARSALIA BENTON, 1999
COHORT: PROLACERTIFORMES CAMP, 1945 NOVA
SUPERORDER: TAPINOPLATIA PETERS, 2000
GRANDORDER: CHARACIOPODA PETERS, 2000
MICRORDER: FENESTRASAURIA PETERS, 2000
ORDER: PTEROSAURIA KAUP, 1834
SUBORDER: EUPTERODACTYLOIDEA BENNETT, 1994
Superfamily: AZHDARCHIDEA Nesov, 1984
Family: AZHDARCHIDAE Nesov, 1984 emend Padian, 1986
Azhdarcho lancicollis
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
Incertae sedis
Gen sp indet.
AVEPODA PAUL, 2002
SUPERORDER: NEOTHEROPODA BAKKER, 1986
GRANDORDER: TETANURAE GAUTHIER, 1986
ORDER: SPINOSAURIA OLSHEVSKY, 1991
Incertae sedis
*Asiamericana asiatica
MICRORDER: COELUROSAURIA HUENE, 1914
ORDER: SEGNOSAURIA BARSBOLD & PERLE, 1980
Superfamily: THERIZINOSAUROIDEA Maleev, 1954
Incertae sedis
Gen. sp indet.
ORDER: MANIRAPTORFORMES HOLTZ, 1996
AVETHEROPODA PAUL, 2001
SUBORDER: COELURIA MARSH, 1881
Superfamily: TYRANNOSAUROIDEA Osborn, 1906 (sensu Bonaparte et. al., 1990)
Incertae sedis
Timurlengia euotica
Family: TYRANNOSAURIDAE Osborn, 1906
Gen. sp indet.
ORDER: MANIRAPTORFORMES HOLTZ, 1996
AVETHEROPODA PAUL, 2001
AVEPECTORA PAUL, 2001
SUBORDER: ORNITHOMIMOSAURIA BARSBOLD, 1976
Family: ORNITHOMIMIDAE Marsh, 1890
Incertae sedis
Gen. sp indet.
SUBORDER: MANIRAPTORA GAUTHIER, 1986
INFRAORDER: PARVES SERENO, 1997
PARVORDER: DEINONYCHOSAURIA COLBERT & D. A. RUSSELL, 1969
Family: DROMAEOSAURIDAE Russell, 1969
*Paronychodon asiaticus
Gen. sp indet.
Family: TROODONTIDAE Gilmore, 1924
Urbacodon sp
COHORT: ORNITHISCHIFORMES COOPER, 1985
SUPERORDER: SAUROPODOMORPHA HUENE, 1932
GRANDORDER: BRONTOSAURIA OLSHEVSKY, 1991 (emend)
MICRORDER: SAUROPODA MARSH, 1878
ORDER: EUSAUROPODA
SUBORDER: NEOSAUROPODA BONAPARTE, 1986
INFRAORDER: MACRONARIA WILSON & SERENO, 1998
PARVORDER: TITANOSAURIFORMES SALGADO ET AL, 1997
Superfamily: TITANOSAUROIDEA Lydekker, 1885 (sensu Upchurch, 1999)
Family: TITANOSAURIDAE Lydekker, 1885
Incertae sedis
Gen. Sp indet.
CLASS: AVES LINNAEUS, 1758
Genus nova
Incertae sedis
*Kuszholia mengi
Kuzholia sp*Platanavis nana
Gen. sp indet.
SUBCLASS: SAURORNITHES NICHOLSON, 1879
AVEBREVICAUDA PAUL, 2001
INFRACLASS: ORNITHOTHORACES NOVA
LEGION: ORNITHURAE NOVA
COHORT: ENANTIORNITHES NOVA
Incertae sedis
*Catenoleimus anachoretus
Gen. sp indet.
ORDER: ALEXORNITHIFORMES BRODKORB, 1976
Family: ALEXORNITHIDAE Brodkorb, 1976
*Abavornis bonaparti
Abavornis sp*Explorornis nessovi
*Explorornis walkeri
= Enantiornis walkeri
Explorornis sp 1 & 2*Incolornis silvae
*Incolornis martini
= Enantiornis martini*Kizylkumavis cretacea
*Lenesornis maltschevskyi
= Ichthyornis maltschevskyi*Sazavis prisca
Family: ZHYRAORNITHIDAE Nesov, 1984
*Zhyraornis kashkarovi
Zhyraornis novaSUBCLASS: ODONTORNITHES MARSH, 1873
INFRACLASS: ODONTOTORMAE MARSH, 1880
ORDER: ICHTHYORNTHIFORMES MARSH, 1873
Incertae sedis
Genus nova
Family: ICHTHYORNITHIDAE Marsh, 1873
*Ichthyornis maltshevskyii
CLASS: MAMMALIA LINNAEUS, 1758
Incertae sedis
Gen. sp indet.
SUBCLASS: THERIIFORMES (ROWE, 1988) McKENNA & BELL, 1997
INFRACLASS: ALLOTHERIA (MARSH, 1880) McKENNA & BELL, 1997
ORDER: MULTITUBERCULATA COPE, 1884
SUBORDER: CIMOLODONTA McKENNA, 1975
Incertae sedis
*Uzbekbaatar kizylkumensis
SUBCLASS: PROTOTHERIA GILL, 1872
INFRACLASS: TRECHNOTHERIA McKENNA, 1975
LEGION: CLADOTHERIA McKENNA, 1975
SUBLEGION: ZATHERIA MCKENNA, 1975
INFRALEGION: TRIBOSPHENIDA (McKENNA, 1975) McKENNA & BELL, 1997
SUPERCOHORT: THERIA (PARKER & HASWELL, 1987) McKENNA & BELL, 1997
Incertae sedis
Gen sp indet.
ORDER: DELTATHEROIDA KIELAN-JAWOROWSKA, 1982
Family: DELTATHERIDIIDAE Gregory & Simpson, 1926
Sulestes karakshi
= *Deltatherus kizylkumensis
= Deltatheroides kizylkumensis
Sulestes sp
= Marsasia spCOHORT: PLACENTALIA (OWEN, 1837) McKENNA & BELL, 1997
MAGNORDER: EPITHERIA (McKENNA, 1975) McKENNA & BELL, 1997
ORDER: KENNALESTIDA McKENNA, 1975
Incertae sedis
Bulaklestes kezbe
Daulestes kulbeckensis
Daulestes inobservabilis
= Taslestes inobservabilisUchkudukodon nessovi
= Daulestes nessoviSUPERORDER: LEPTICTIDAE McKENNA, 1975
Incertae sedis
Paranyctoides quadrans
= *Paranyctoides aralensisFamily: genus and species indeterminate
( = Suborder: Mixotheridia)SUPERORDER: PREPTOTHERIA McKENNA, 1975
GRANDORDER: UNGULATOMORPHA ARCHIBALD, 1996
Family: ZHELESTIDAE Nesov, 1990
Aspanlestes aptap
= *Zhelestes bezelgern
Aspanlestes sp*Eoungulatum kudukensis
= Eoungulatum kudukensis*Parazhelestes robustus
*Parazhelestes minor
= Parazhelestes minor*Sorlestes budan
Zhelestes sp. cf. Z. temirkazyk
Gen. sp indet.
GRANDORDER: UNGULATA LINNAEUS, 1766
MICRORDER: EPARCTOCYONA McKENNA, 1975
ORDER: CONDYLARTHRA COPE, 1881
Family: PERIPTYCHIDAE Cope, 1882Gen. Sp indet.
Gen. sp indet.
Family: KULBECKIIDAE NESOV, 1993
Kulbeckia kulbecke
_____________________________________________________________________________________
UZBEKISTAN
KIZYLKUM DESERT
DZHYRAKUUK
LOCALITY CBI-14
BISSEKTY FORMATION
TURONIAN STAGE
UPPERMOST GALLIC SUBEPOCH
LOWER GULF EPOCH
EARLY CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
AVEPODA PAUL, 2002
SUPERORDER: NEOTHEROPODA BAKKER, 1986
GRANDORDER: TETANURAE GAUTHIER, 1986
MICRORDER: COELUROSAURIA HUENE, 1914
ORDER: MANIRAPTORFORMES HOLTZ, 1996
AVETHEROPODA PAUL, 200
AVEPECTORA PAUL, 2001
SUBORDER: ORNITHOMIMOSAURIA BARSBOLD, 1976
Family: ORNITHOMIMIDAE Marsh, 1890
Incertae sedis
Gen. sp indet.
_____________________________________________________________________________________
UZBEKISTAN
CENTRAL KIZYLKKUM DESERT
DZHYRAKUDUK
CBI-14a
MIDDLE OF THE BISSEKTY FORMATION
CONIACIAN STAGE
LOWER SENONIAN SUBEPOCH
MIDDLE GULF EPOCH
EARLY CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
AVEPODA PAUL, 2002
SUPERORDER: NEOTHEROPODA BAKKER, 1986
GRANDORDER: TETANURAE GAUTHIER, 1986
MICRORDER: COELUROSAURIA HUENE, 1914
ORDER: SEGNOSAURIA BARSBOLD & PERLE, 1980
Superfamily: THERIZINOSAUROIDEA Maleev, 1954
Incertae sedis
Gen. sp indet.
_____________________________________________________________________________________
UZBEKISTAN
CENTRAL KIZYLKKUM DESERT
DZHYRAKUDUK
CBI-14v
MIDDLE OF THE BISSEKTY FORMATION
CONIACIAN STAGE
LOWER SENONIAN SUBEPOCH
MIDDLE GULF EPOCH
EARLY CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: AVEMETATARSALIA BENTON, 1999
COHORT: PROLACERTIFORMES CAMP, 1945 NOVA
SUPERORDER: TAPINOPLATIA PETERS, 2000
GRANDORDER: CHARACIOPODA PETERS, 2000
MICRORDER: FENESTRASAURIA PETERS, 2000
ORDER: PTEROSAURIA KAUP, 1834
SUBORDER: EUPTERODACTYLOIDEA BENNETT, 1994
Superfamily: AZHDARCHIDEA Nesov, 1984
Family: AZHDARCHIDAE Nesov, 1984 emend Padian, 1986
Azhdarcho lancicollis
_____________________________________________________________________________________
UZBEKISTAN
CENTRAL KIZYLKKUM DESERT
DZHYRANKUDUK
CBI-15
UPPER BISSEKTY FORMATION
CONIACIAN STAGE
LOWER SENONIAN SUBEPOCH
MIDDLE GULF EPOCH
EARLY CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: MAMMALIA LINNAEUS, 1758
SUBCLASS: PROTOTHERIA GILL, 1872
INFRACLASS: TRECHNOTHERIA McKENNA, 1975
LEGION: CLADOTHERIA McKENNA, 1975
SUBLEGION: ZATHERIA MCKENNA, 1975
INFRALEGION: TRIBOSPHENIDA (McKENNA, 1975) McKENNA & BELL, 1997
SUPERCOHORT: THERIA (PARKER & HASWELL, 1987) McKENNA & BELL, 1997
COHORT: PLACENTALIA (OWEN, 1837) McKENNA & BELL, 1997
MAGNORDER: EPITHERIA (McKENNA, 1975) McKENNA & BELL, 1997
ORDER: KENNALESTIDA McKENNA, 1975
Incertae sedis
Bulaklestes kezbe
_____________________________________________________________________________________
UZBEKISTAN
CENTRAL KIZYLKKUM DESERT
DZHYRAKUDUK
CBI-15a
MIDDLE OF THE BISSEKTY FORMATION
CONIACIAN STAGE
LOWER SENONIAN SUBEPOCH
MIDDLE GULF EPOCH
EARLY CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
AVEPODA PAUL, 2002
SUPERORDER: NEOTHEROPODA BAKKER, 1986
GRANDORDER: TETANURAE GAUTHIER, 1986
MICRORDER: COELUROSAURIA HUENE, 1914
ORDER: SEGNOSAURIA BARSBOLD & PERLE, 1980
Superfamily: THERIZINOSAUROIDEA Maleev, 1954
Incertae sedis
Gen. sp indet.
_____________________________________________________________________________________
UZBEKISTAN
CENTRAL KIZYLKKUM DESERT
DZHYRANKUDUK
CBI-16
UPPER BISSEKTY FORMATION
CONIACIAN STAGE
LOWER SENONIAN SUBEPOCH
MIDDLE GULF EPOCH
EARLY CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: AVEMETATARSALIA BENTON, 1999
COHORT: PROLACERTIFORMES CAMP, 1945 NOVA
SUPERORDER: TAPINOPLATIA PETERS, 2000
GRANDORDER: CHARACIOPODA PETERS, 2000
MICRORDER: FENESTRASAURIA PETERS, 2000
ORDER: PTEROSAURIA KAUP, 1834
SUBORDER: EUPTERODACTYLOIDEA BENNETT, 1994
Superfamily: AZHDARCHIDEA Nesov, 1984
Family: AZHDARCHIDAE Nesov, 1984 emend Padian, 1986
Azhdarcho lancicollis
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
Incertae sedis
Gen. sp indet.
AVEPODA PAUL, 2002
SUPERORDER: NEOTHEROPODA BAKKER, 1986
GRANDORDER: TETANURAE GAUTHIER, 1986
MICRORDER: COELUROSAURIA HUENE, 1914
ORDER: SEGNOSAURIA BARSBOLD & PERLE, 1980
Family: SEGNOSAURIDAE Perle, 1979
Gen. sp indet.
ORDER: MANIRAPTORFORMES HOLTZ, 1996
AVETHEROPODA PAUL, 200
AVEPECTORA PAUL, 2001
SUBORDER: ORNITHOMIMOSAURIA BARSBOLD, 1976
Family: ORNITHOMIMIDAE Marsh, 1890
Incertae sedis
Gen. sp indet.
___________________________________________________________________________________
UZBEKISTAN
CENTRAL KIZYLKKUM DESERT
DZHYRANKUDUK
CBI-17
UPPER BISSEKTY FORMATION
CONIACIAN STAGE
LOWER SENONIAN SUBEPOCH
MIDDLE GULF EPOCH
EARLY CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: AMPHIBIA LINNAEUS, 1758
SUBCLASS: LISSAMPHIBIA, HAECKEL, 1866?
ORDER: ALLOCAUDATA FOX & NAYLOR, 1982
Family: ALBANERPETONTIDAE Fox & Naylor, 1982
*Nukusurus sodalis
ORDER: CAUDATA SCOPOLI, 1777
Incertae sedis
Nesovtriton mynbulakensis
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: AVEMETATARSALIA BENTON, 1999
COHORT: PROLACERTIFORMES CAMP, 1945 NOVA
SUPERORDER: TAPINOPLATIA PETERS, 2000
GRANDORDER: CHARACIOPODA PETERS, 2000
MICRORDER: FENESTRASAURIA PETERS, 2000
ORDER: PTEROSAURIA KAUP, 1834
SUBORDER: EUPTERODACTYLOIDEA BENNETT, 1994
Superfamily: AZHDARCHIDEA Nesov, 1984
Family: AZHDARCHIDAE Nesov, 1984 emend Padian, 1986
Azhdarcho lancicollis
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
MICRORDER: COELUROSAURIA HUENE, 1914
ORDER: MANIRAPTORFORMES HOLTZ, 1996
AVETHEROPODA PAUL, 200
AVEPECTORA PAUL, 2001
SUBORDER: ORNITHOMIMOSAURIA BARSBOLD, 1976
Family: ORNITHOMIMIDAE Marsh, 1890
Incertae sedis
Gen. sp indet.
SUBORDER: MANIRAPTORA GAUTHIER, 1986
INFRAORDER: PARVES SERENO, 1997
PARVORDER: DEINONYCHOSAURIA COLBERT & D. A. RUSSELL, 1969
Family: TROODONTIDAE Gilmore, 1924
Urbacodon sp
COHORT: ORNITHISCHIFORMES COOPER, 1985
MAGNORDER: ORNITHISCHIA SEELEY, 1888
COHORT: ORNITHISCHIFORMES COOPER, 1985
MAGNORDER: ORNITHISCHIA SEELEY, 1888
SUPERORDER: EURYOPDA SERENO, 1986
GRANDORDER: ARMATOSAURIA ZHAO, 1983 (emend)
ORDER: ANKYLOSAURIA OSBORN, 1923
INFRAORDER: ANKYLOSAURA OSBORN, 1923
Family: ANKYLOSAURIDAE Brown, 1908
Incertae sedis
Gen. sp indet.
CLASS: MAMMALIA LINNAEUS, 1758
Incertae sedis
Gen. sp indet.
SUBCLASS: THERIIFORMES (ROWE, 1988) McKENNA & BELL, 1997
INFRACLASS: ALLOTHERIA (MARSH, 1880) McKENNA & BELL, 1997
ORDER: MULTITUBERCULATA COPE, 1884
SUBORDER: CIMOLODONTA McKENNA, 1975
Incertae sedis
Uzbekbaatar kizylkumensis
SUBCLASS: PROTOTHERIA GILL, 1872
INFRACLASS: TRECHNOTHERIA McKENNA, 1975
LEGION: CLADOTHERIA McKENNA, 1975
SUBLEGION: ZATHERIA MCKENNA, 1975
INFRALEGION: TRIBOSPHENIDA (McKENNA, 1975) McKENNA & BELL, 1997
SUPERCOHORT: THERIA (PARKER & HASWELL, 1987) McKENNA & BELL, 1997
Incertae sedis
Genus nova.
ORDER: DELTATHEROIDA KIELAN-JAWOROWSKA, 1982
Family: DELTATHERIDIIDAE Gregory & Simpson, 1926
Sulestes karakshi
COHORT: PLACENTALIA (OWEN, 1837) McKENNA & BELL, 1997
MAGNORDER: EPITHERIA (McKENNA, 1975) McKENNA & BELL, 1997
SUPERORDER: PREPTOTHERIA McKENNA, 1975
GRANDORDER: UNGULATOMORPHA ARCHIBALD, 1996
Family: ZHELESTIDAE Nesov, 1990
*Zhelestes temirkazyk
_____________________________________________________________________________________
UZBEKISTAN
KIZYLKUM DESERT
DZHYRAKUUK
LOCALITY CBI-17a
BISSEKTY FORMATION
CONIACIAN STAGE
LOWER SENONIAN SUBEPOCH
MIDDLE GULF EPOCH
EARLY CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: AVEMETATARSALIA BENTON, 1999
COHORT: PROLACERTIFORMES CAMP, 1945 NOVA
SUPERORDER: TAPINOPLATIA PETERS, 2000
GRANDORDER: CHARACIOPODA PETERS, 2000
MICRORDER: FENESTRASAURIA PETERS, 2000
ORDER: PTEROSAURIA KAUP, 1834
SUBORDER: EUPTERODACTYLOIDEA BENNETT, 1994
Superfamily: AZHDARCHIDEA Nesov, 1984
Family: AZHDARCHIDAE Nesov, 1984 emend Padian, 1986
Azhdarcho lancicollis
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
MICRORDER: COELUROSAURIA HUENE, 1914
ORDER: SEGNOSAURIA BARSBOLD & PERLE, 1980
Superfamily: THERIZINOSAUROIDEA Maleev, 1954
Incertae sedis
Gen. sp indet.
ORDER: MANIRAPTORFORMES HOLTZ, 1996
AVETHEROPODA PAUL, 200
AVEPECTORA PAUL, 2001
SUBORDER: ORNITHOMIMOSAURIA BARSBOLD, 1976
Family: ORNITHOMIMIDAE Marsh, 1890
Incertae sedis
Gen. sp indet.
CLASS: MAMMALIA LINNAEUS, 1758
SUBCLASS: PROTOTHERIA GILL, 1872
INFRACLASS: TRECHNOTHERIA McKENNA, 1975
LEGION: CLADOTHERIA McKENNA, 1975
SUBLEGION: ZATHERIA MCKENNA, 1975
INFRALEGION: TRIBOSPHENIDA (McKENNA, 1975) McKENNA & BELL, 1997
SUPERCOHORT: THERIA (PARKER & HASWELL, 1987) McKENNA & BELL, 1997
ORDER: DELTATHEROIDA KIELAN-JAWOROWSKA, 1982
Family: DELTATHERIDIIDAE Gregory & Simpson, 1926
Sulestes karakshi
_____________________________________________________________________________________
UZBEKISTAN
KIZYLKUM DESERT
DZHYRAKUUK
LOCALITY CBI-17g
BISSEKTY FORMATION
CONIACIAN STAGE
LOWER SENONIAN SUBEPOCH
MIDDLE GULF EPOCH
EARLY CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 194
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
AVEPODA PAUL, 2002
SUPERORDER: NEOTHEROPODA BAKKER, 1986
GRANDORDER: TETANURAE GAUTHIER, 1986
MICRORDER: COELUROSAURIA HUENE, 1914
ORDER: MANIRAPTORFORMES HOLTZ, 1996
AVETHEROPODA PAUL, 2001
SUBORDER: COELURIA MARSH, 1881
Superfamily: TYRANNOSAUROIDEA Osborn, 1906 (sensu Bonaparte et. al., 1990)
Family: ALIORAMIDAE Olshevsky, Ford & Yamamoto, 1995
Alectrosaurus sp
_____________________________________________________________________________________
UZBEKISTAN
KYZYLKUM DESERT
CBI-19
DZHYRAKUDUK LOCALITY
BISSEKTY FORMATION
CONIACIAN STAGE
LOWER SENONIAN SUBEPOCH
MIDDLE GULF EPOCH
EARLY CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 184
COHORT: ORNITHISCHIFORMES COOPER, 1985
MAGNORDER: ORNITHISCHIA SEELEY, 1888
SUPERORDER: NEORNITHISCHIA COOPER, 1985
GRANDORDER: ORNITHOPODA MARSH, 1871
MICRORDER: EUORNITHOPODA SERENO, 1986
ORDER: IGUANODONTIA DOLLO, 1888
INFRAORDER: STYRACOSTERNA SERENO, 1986
PARVORDER: HADROSAURIFORMES SERENO, 1997
SUPERFAMILY: HADROSAUROIDEA COPE, 1869 (SENSU SERENO, 1986)
Family: HADROSAURIDAE COPE, 1869
Incertae sedis
Gilmoreosaurus arkhangelskyi
_____________________________________________________________________________________
UZBEKISTAN
CENTRAL KIZYLKKUM DESERT
DZHYRAKUDUK
CBI-20
MIDDLE OF THE BISSEKTY FORMATION
CONIACIAN STAGE
LOWER SENONIAN SUBEPOCH
MIDDLE GULF EPOCH
EARLY CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
AVEPODA PAUL, 2002
SUPERORDER: NEOTHEROPODA BAKKER, 1986
GRANDORDER: TETANURAE GAUTHIER, 1986
MICRORDER: COELUROSAURIA HUENE, 1914
ORDER: SEGNOSAURIA BARSBOLD & PERLE, 1980
Superfamily: THERIZINOSAUROIDEA Maleev, 1954
Incertae sedis
Gen. sp indet.
_____________________________________________________________________________________
UZBEKISTAN
KYZYLKUM DESERT
CBI-23
DZHYRAKUDUK LOCALITY
BISSEKTY FORMATION
TURONIAN STAGE
UPPERMOST GALLIC SUBEPOCH
LOWER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 184
COHORT: ORNITHISCHIFORMES COOPER, 1985
MAGNORDER: ORNITHISCHIA SEELEY, 1888
COHORT: ORNITHISCHIFORMES COOPER, 1985
MAGNORDER: ORNITHISCHIA SEELEY, 1888
SUPERORDER: NEORNITHISCHIA COOPER, 1985
GRANDORDER: ORNITHOPODA MARSH, 1871
MICRORDER: EUORNITHOPODA SERENO, 1986
ORDER: IGUANODONTIA DOLLO, 1888
INFRAORDER: STYRACOSTERNA SERENO, 1986
PARVORDER: HADROSAURIFORMES SERENO, 1997
SUPERFAMILY: HADROSAUROIDEA COPE, 1869 (SENSU SERENO, 1986)
Family: HADROSAURIDAE COPE, 1869
Incertae sedis
Gilmoreosaurus arkhangelskyi
_____________________________________________________________________________________
UZBEKISTAN
KYZYLKUM DESERT
CBI-24
DZHYRAKUDUK LOCALITY
BISSEKTY FORMATION
TURONIAN STAGE
UPPERMOST GALLIC SUBEPOCH
LOWER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 184
COHORT: ORNITHISCHIFORMES COOPER, 1985
MAGNORDER: ORNITHISCHIA SEELEY, 1888
COHORT: ORNITHISCHIFORMES COOPER, 1985
MAGNORDER: ORNITHISCHIA SEELEY, 1888
SUPERORDER: NEORNITHISCHIA COOPER, 1985
GRANDORDER: ORNITHOPODA MARSH, 1871
MICRORDER: EUORNITHOPODA SERENO, 1986
ORDER: IGUANODONTIA DOLLO, 1888
INFRAORDER: STYRACOSTERNA SERENO, 1986
PARVORDER: HADROSAURIFORMES SERENO, 1997
SUPERFAMILY: HADROSAUROIDEA COPE, 1869 (SENSU SERENO, 1986)
Family: HADROSAURIDAE COPE, 1869
Incertae sedis
Gilmoreosaurus arkhangelskyi
_____________________________________________________________________________________
UZEBEKISTAN
DZHARAKUDUK
CBI-27
LOWER SECTION OF THE BISSEKT FORMATION
(BISSEKTIYSKAYA SVITA)
UPPER TURONIAN STAGE
UPPERMOST GALLIC SUBEPOCH
OR CONIACIAN STAGE
LOWERMOST SENONIAN SUBEPOCH
LOWER GULF EPOCH
EARLY LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
AVEPODA PAUL, 2002
SUPERORDER: NEOTHEROPODA BAKKER, 1986
GRANDORDER: TETANURAE GAUTHIER, 1986
MICRORDER: COELUROSAURIA HUENE, 1914
ORDER: SEGNOSAURIA BARSBOLD & PERLE, 1980
Superfamily: THERIZINOSAUROIDEA Maleev, 1954
Incertae sedis
Gen. sp indet.
COHORT: ORNITHISCHIFORMES COOPER, 1985
MAGNORDER: ORNITHISCHIA SEELEY, 1888
SUPERORDER: NEORNITHISCHIA COOPER, 1985
GRANDORDER: ORNITHOPODA MARSH, 1871
MICRORDER: MARGINOCEPHALIA SERENO, 1986
ORDER: CERATOPIA MARSH, 1890
SUBORDER: NEOCERATOPIA SERENO, 1986
INFRAORDER: CORONOSAURIA SERENO, 1986
PARVORDER: CERATOPOMORPHA Wolfe & Kirkland, 1998 (emend)
Superfamily: CERATOPOIDEA Hay, 1902 (emend)
Incertae sedis
*Turanoceratops tardabilis
_____________________________________________________________________________________
UZBEKISTAN
KYZYLKUM DESERT
CBI-27
DZHYRAKUDUK LOCALITY
BISSEKTY FORMATION
TURONIAN STAGE
UPPERMOST GALLIC SUBEPOCH
LOWER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 184
COHORT: THEROPODA EMEND MARSH, 1881
AVEPODA PAUL, 2002
SUPERORDER: NEOTHEROPODA BAKKER, 1986
GRANDORDER: TETANURAE GAUTHIER, 1986
MICRORDER: COELUROSAURIA HUENE, 1914
ORDER: MANIRAPTORFORMES HOLTZ, 1996
AVETHEROPODA PAUL, 2001
SUBORDER: COELURIA MARSH, 1881
Superfamily: TYRANNOSAUROIDEA Osborn, 1906 (sensu Bonaparte et. al., 1990)
Incertae sedis
Timurlengia euotica
COHORT: ORNITHISCHIFORMES COOPER, 1985
MAGNORDER: ORNITHISCHIA SEELEY, 1888
SUPERORDER: NEORNITHISCHIA COOPER, 1985
GRANDORDER: ORNITHOPODA MARSH, 1871
MICRORDER: MARGINOCEPHALIA SERENO, 1986
ORDER: CERATOPIA MARSH, 1890
SUBORDER: NEOCERATOPIA SERENO, 1986
INFRAORDER: CORONOSAURIA SERENO, 1986
PARVORDER: CERATOPOMORPHA Wolfe & Kirkland, 1998 (emend)
Superfamily: CERATOPOIDEA Hay, 1902 (emend)
Incertae sedis
Turanoceratops tardabilis
_____________________________________________________________________________________
UZBEKISTAN
KIZYLKUM DESERT
DZHYRAKUDUK LOCALITY
CBI-28
BISSEKTY FORMATION
LATE TURONIAN STAGE
UPPER GALLIC SUBEPOCH
LOWER GULF EPOCH
EARLY LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: CHELONOMORPHA KUHN, 1960
SUPERCOHORT: TESTUDINES LINNAEUS, 1758
COHORT: CASICHELYDIA emend GAFFNEY & MEYLAN, 1988
MAGANORDER: CRYPTODIRA GAFFNEY & MEYLAN, 1988
SUPERORDER: SELMACRYPTODIRA GAFFNEY, HUTCHISON, JENKINS & MEEKR, 1987
GRANDORDER: DAIOCRYPTODIRA GAFFNEY & MEYLAN, 1988
MICRORDER: CENTROCRYPTODIRA GAFFNEY & MEYLAN, 1988
ORDER: POLYCRPTODIRA GAFFNEY & MEYLAN, 1988
SUBORDER: PROCOELOCRYPTODIRA GAFFNEY & MEYLAN, 1988
INFRAORDER: CHELOMACRYPTODIRA GAFFNEY & MEYLAN, 1988
PARVORDER: TRIONYCHIDA GRAY, 1870
Superfamily: TRIONYCHIDEA Bell, 1828 (sensu Gray, 1870)
Family: TRIONYCHIDAE Bell, 1828
*Khunnuchelys kizlkumensis
SUBCLASS: DIAPSIDA OSBORN, 1903
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: CRUROTARSI SERENO & ARCUCCI, 1990
COHORT: CROCODYLOTARSI BENTON & CLARK, 1988
MAGNORDER: SUCHIA KREBS, 1974
SUPERORDER: CROCODYLOMORPHA WALKER, 1968
GRANDORDER: CROCODILIFORMES HAY, 1930
Incertae sedis
Gen. sp indet.
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
AVEPODA PAUL, 2002
SUPERORDER: NEOTHEROPODA BAKKER, 1986
GRANDORDER: TETANURAE GAUTHIER, 1986
MICRORDER: COELUROSAURIA HUENE, 1914
ORDER: SEGNOSAURIA BARSBOLD & PERLE, 1980
Superfamily: THERIZINOSAUROIDEA Maleev, 1954
Incertae sedis
Gen. sp indet.
ORDER: MANIRAPTORFORMES HOLTZ, 1996
AVETHEROPODA PAUL, 2001
SUBORDER: COELURIA MARSH, 1881
Superfamily: TYRANNOSAUROIDEA Osborn, 1906 (sensu Bonaparte et. al., 1990)
Incertae sedis
Timurlengia euotica
Family: ALIORAMIDAE Olshevsky, Ford & Yamamoto, 1995
Alectrosaurus sp
AVEPECTORA PAUL, 2001
SUBORDER: ORNITHOMIMOSAURIA BARSBOLD, 1976
Family: ORNITHOMIMIDAE Marsh, 1890
Incertae sedis
Gen. sp indet.
COHORT: ORNITHISCHIFORMES COOPER, 1985
MAGNORDER: ORNITHISCHIA SEELEY, 1888
COHORT: ORNITHISCHIFORMES COOPER, 1985
MAGNORDER: ORNITHISCHIA SEELEY, 1888
SUPERORDER: NEORNITHISCHIA COOPER, 1985
GRANDORDER: ORNITHOPODA MARSH, 1871
MICRORDER: EUORNITHOPODA SERENO, 1986
ORDER: IGUANODONTIA DOLLO, 1888
INFRAORDER: STYRACOSTERNA SERENO, 1986
PARVORDER: HADROSAURIFORMES SERENO, 1997
SUPERFAMILY: HADROSAUROIDEA COPE, 1869 (SENSU SERENO, 1986)
Family: HADROSAURIDAE COPE, 1869
Incertae sedis
Gilmoreosaurus arkhangelskyi
MICRORDER: MARGINOCEPHALIA SERENO, 1986
ORDER: CERATOPIA MARSH, 1890
SUBORDER: NEOCERATOPIA SERENO, 1986
INFRAORDER: CORONOSAURIA SERENO, 1986
PARVORDER: CERATOPOMORPHA Wolfe & Kirkland, 1998 (emend)
Superfamily: CERATOPOIDEA Hay, 1902 (emend)
Incertae sedis
Turanoceratops tardabilis
SUPERORDER: EURYOPDA SERENO, 1986
GRANDORDER: ARMATOSAURIA ZHAO, 1983 (emend)
ORDER: ANKYLOSAURIA OSBORN, 1923
INFRAORDER: ANKYLOSAURA OSBORN, 1923
Family: ANKYLOSAURIDAE Brown, 1908
Incertae sedis
Gen. sp indet.
_____________________________________________________________________________________
UZBEKISTAN
CENTRAL KIZYLKKUM DESERT
DZHYRAKUDUK
CBI-41
MIDDLE OF THE BISSEKTY FORMATION
CONIACIAN STAGE
LOWER SENONIAN SUBEPOCH
MIDDLE GULF EPOCH
EARLY CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: AVEMETATARSALIA BENTON, 1999
COHORT: PROLACERTIFORMES CAMP, 1945 NOVA
SUPERORDER: TAPINOPLATIA PETERS, 2000
GRANDORDER: CHARACIOPODA PETERS, 2000
MICRORDER: FENESTRASAURIA PETERS, 2000
ORDER: PTEROSAURIA KAUP, 1834
SUBORDER: EUPTERODACTYLOIDEA BENNETT, 1994
Superfamily: AZHDARCHIDEA Nesov, 1984
Family: AZHDARCHIDAE Nesov, 1984 emend Padian, 1986
Azhdarcho lancicollis
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
AVEPODA PAUL, 2002
SUPERORDER: NEOTHEROPODA BAKKER, 1986
GRANDORDER: TETANURAE GAUTHIER, 1986
MICRORDER: COELUROSAURIA HUENE, 1914
ORDER: SEGNOSAURIA BARSBOLD & PERLE, 1980
Superfamily: THERIZINOSAUROIDEA Maleev, 1954
Incertae sedis
Gen. sp indet.
_____________________________________________________________________________________
UZBEKISTAN
CENTRAL KIZYLKKUM DESERT
DZHYRAKUDUK
CBI-50
MIDDLE OF THE BISSEKTY FORMATION
CONIACIAN STAGE
LOWER SENONIAN SUBEPOCH
MIDDLE GULF EPOCH
EARLY CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: AVEMETATARSALIA BENTON, 1999
COHORT: PROLACERTIFORMES CAMP, 1945 NOVA
SUPERORDER: TAPINOPLATIA PETERS, 2000
GRANDORDER: CHARACIOPODA PETERS, 2000
MICRORDER: FENESTRASAURIA PETERS, 2000
ORDER: PTEROSAURIA KAUP, 1834
SUBORDER: EUPTERODACTYLOIDEA BENNETT, 1994
Superfamily: AZHDARCHIDEA Nesov, 1984
Family: AZHDARCHIDAE Nesov, 1984 emend Padian, 1986
Azhdarcho lancicollis
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
AVEPODA PAUL, 2002
SUPERORDER: NEOTHEROPODA BAKKER, 1986
GRANDORDER: TETANURAE GAUTHIER, 1986
MICRORDER: COELUROSAURIA HUENE, 1914
ORDER: MANIRAPTORFORMES HOLTZ, 1996
AVETHEROPODA PAUL, 2001
SUBORDER: COELURIA MARSH, 1881
Superfamily: TYRANNOSAUROIDEA Osborn, 1906 (sensu Bonaparte et. al., 1990)
Family: TYRANNOSAURIDAE Osborn, 1906
Gen. sp indet.
_____________________________________________________________________________________
_____________________________________________________________________________________
UZBEKISTAN
DZHYRAKUDUK
CBI-51
UPPER MIDDLE BISSEKTY FORMATION
CONIACIAN STAGE
LOWER SENONIAN SUBEPOCH
LOWER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
AVEPODA PAUL, 2002
SUPERORDER: NEOTHEROPODA BAKKER, 1986
GRANDORDER: TETANURAE GAUTHIER, 1986
MICRORDER: COELUROSAURIA HUENE, 1914
ORDER: MANIRAPTORFORMES HOLTZ, 1996
AVETHEROPODA PAUL, 200
AVEPECTORA PAUL, 2001
SUBORDER: ORNITHOMIMOSAURIA BARSBOLD, 1976
Family: ORNITHOMIMIDAE Marsh, 1890
Incertae sedis
Gen. sp indet.
CLASS: MAMMALIA LINNAEUS, 1758
SUBCLASS: PROTOTHERIA GILL, 1872
INFRACLASS: TRECHNOTHERIA McKENNA, 1975
LEGION: CLADOTHERIA McKENNA, 1975
SUBLEGION: ZATHERIA MCKENNA, 1975
INFRALEGION: TRIBOSPHENIDA (McKENNA, 1975) McKENNA & BELL, 1997
SUPERCOHORT: THERIA (PARKER & HASWELL, 1987) McKENNA & BELL, 1997
ORDER: DELTATHEROIDA KIELAN-JAWOROWSKA, 1982
Family: DELTATHERIDIIDAE Gregory & Simpson, 1926
Sulestes karakshi
= *Marsasia aenigma
_____________________________________________________________________________________
UZBEKISTAN
CENTRAL KIZYLKKUM DESERT
DZHYRAKUDUK
CBI-52
MIDDLE OF THE BISSEKTY FORMATION
CONIACIAN STAGE
LOWER SENONIAN SUBEPOCH
MIDDLE GULF EPOCH
EARLY CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: AVEMETATARSALIA BENTON, 1999
COHORT: PROLACERTIFORMES CAMP, 1945 NOVA
SUPERORDER: TAPINOPLATIA PETERS, 2000
GRANDORDER: CHARACIOPODA PETERS, 2000
MICRORDER: FENESTRASAURIA PETERS, 2000
ORDER: PTEROSAURIA KAUP, 1834
SUBORDER: EUPTERODACTYLOIDEA BENNETT, 1994
Superfamily: AZHDARCHIDEA Nesov, 1984
Family: AZHDARCHIDAE Nesov, 1984 emend Padian, 1986
Azhdarcho lancicollis
_____________________________________________________________________________________
UZBEKISTAN
CENTRAL KIZYLKKUM DESERT
DZHYRAKUDUK
CBI-56
MIDDLE OF THE BISSEKTY FORMATION
CONIACIAN STAGE
LOWER SENONIAN SUBEPOCH
MIDDLE GULF EPOCH
EARLY CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: AVEMETATARSALIA BENTON, 1999
COHORT: PROLACERTIFORMES CAMP, 1945 NOVA
SUPERORDER: TAPINOPLATIA PETERS, 2000
GRANDORDER: CHARACIOPODA PETERS, 2000
MICRORDER: FENESTRASAURIA PETERS, 2000
ORDER: PTEROSAURIA KAUP, 1834
SUBORDER: EUPTERODACTYLOIDEA BENNETT, 1994
Superfamily: AZHDARCHIDEA Nesov, 1984
Family: AZHDARCHIDAE Nesov, 1984 emend Padian, 1986
Azhdarcho lancicollis
_____________________________________________________________________________________
UZBEKISTAN
KYCYLKUM DESERT
DZHARAKUDUK
CBI-117
AITYM FORMATION
?SANTONIAN STAGE
SENONIAN SUBEPOCH
UPPER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: MAMMALIA LINNAEUS, 1758
SUBCLASS: THERIIFORMES (ROWE, 1988) McKENNA & BELL, 1997
INFRACLASS: ALLOTHERIA (MARSH, 1880) McKENNA & BELL, 1997
ORDER: MULTITUBERCULATA COPE, 1884
SUBORDER: CIMOLODONTA McKENNA, 1975
Incertae sedis
Uzbekbaatar wardi
SUBCLASS: PROTOTHERIA GILL, 1872
INFRACLASS: HOLOTHERIA (WIBEL, ROUGIER, NOVACEK, MCKENNA & DASHZEVEG, 1995) MCKENNA & BELL, 1997
SUPERLEGION: SYMMETRODONTA (SIMPSON, 1925) McKENNA & BELL, 1997
LEGION: SYMMETRODONTA SIMPSON, 1925
ORDER: SPALACOTHERIIDA PROTHERO, 1981
Family: SPALACOTHERIIDAE Marsh, 1887
Shalbaatar sp
= cf. Shalbaatar spINFRACLASS: TRECHNOTHERIA McKENNA, 1975
LEGION: CLADOTHERIA McKENNA, 1975
SUBLEGION: ZATHERIA MCKENNA, 1975
INFRALEGION: TRIBOSPHENIDA (McKENNA, 1975) McKENNA & BELL, 1997
SUPERCOHORT: THERIA (PARKER & HASWELL, 1987) McKENNA & BELL, 1997
ORDER: DELTATHEROIDA KIELAN-JAWOROWSKA, 1982
Family: DELTATHERIDIIDAE Gregory & Simpson, 1926
Sulestes sp
= Deltatherus sp
= cf. Deltatherus spCOHORT: MARSUPIALIA (ILLIGER, 1811) McKENNA & BELL, 1997
COHORT: PLACENTALIA (OWEN, 1837) McKENNA & BELL, 1997
MAGNORDER: EPITHERIA (McKENNA, 1975) McKENNA & BELL, 1997
ORDER: KENNALESTIDA McKENNA, 1975
Incertae sedis
Daulestes sp
= aff. Daulestes spSUPERORDER: LEPTICTIDAE McKENNA, 1975
Incertae sedis
Paranyctoides aralensis
= Paranyctoides sp. cf. P. aralensis
Paranyctoides spSUPERORDER: PREPTOTHERIA McKENNA, 1975
GRANDORDER: UNGULATOMORPHA ARCHIBALD, 1996
Family: ZHELESTIDAE Nesov, 1990
Subfamily: ZHELESTINAE Nesov, 1985
Aspanlestes sp
Eoungulatum kudukensis
= Eoungulatum sp. cf. E. kudukensisParazhelestes robustus
= Parazhelestes sp. cf. P. robustus
*Parazhelestes minor
= Parazhelestes sp. aff minorGRANDORDER: UNGULATA LINNAEUS, 1766
MICRORDER: EPARCTOCYONA McKENNA, 1975
ORDER: CONDYLARTHRA COPE, 1881
Family: KULBECKIIDAE NESOV, 1993
Kulbeckia kulbecke
= Kulbeckia sp. cf. K. kulbecke
K. sp
= aff. Kulbeckia sp
_____________________________________________________________________________________
UZBEKISTAN
DZHYRAKUDUK
CBI-116-3
UPPER MIDDLE BISSEKTY FORMATION
TURONIAN STAGE
UPPERMOST GALLIC SUBEPOCH
LOWER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
AVEPODA PAUL, 2002
SUPERORDER: NEOTHEROPODA BAKKER, 1986
GRANDORDER: TETANURAE GAUTHIER, 1986
MICRORDER: COELUROSAURIA HUENE, 1914
ORDER: MANIRAPTORFORMES HOLTZ, 1996
AVETHEROPODA PAUL, 200
AVEPECTORA PAUL, 2001
SUBORDER: ORNITHOMIMOSAURIA BARSBOLD, 1976
Family: ORNITHOMIMIDAE Marsh, 1890
Incertae sedis
Gen. sp indet.
_____________________________________________________________________________________
UZBEKISTAN
KYZYL-KUM
UPPER TURONIAN-CONIACIAN STAGE
LOWER SENONIAN SUBEPOCH
LOWER GULF EPOCH
EARLY LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
AVEPODA PAUL, 2002
SUPERORDER: NEOTHEROPODA BAKKER, 1986
GRANDORDER: TETANURAE GAUTHIER, 1986
MICRORDER: COELUROSAURIA HUENE, 1914
ORDER: MANIRAPTORFORMES HOLTZ, 1996
AVETHEROPODA PAUL, 2001
SUBORDER: COELURIA MARSH, 1881
Superfamily: TYRANNOSAUROIDEA Osborn, 1906 (sensu Bonaparte et. al., 1990)
Family: ALIORAMIDAE Olshevsky, Ford & Yamamoto, 1995
Alectrosaurus sp
_____________________________________________________________________________________
UZBEKISTAN
CENTRAL KIZYLKKUM DESERT
DZHYRANKUDUK
CDZ-17A
LOWER BISSEKTY FORMATION
CONIACIAN STAGE
LOWER SENONIAN SUBEPOCH
MIDDLE GULF EPOCH
EARLY CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: AMPHIBIA LINNAEUS, 1758
SUBCLASS: LISSAMPHIBIA, HAECKEL, 1866?
SUPERORDER: SALIENTIA LAURENTI, 1768
ORDER: ANURA RAFINESQUE, 1815
Incertae sedis
Aralobatrachus robustus
_____________________________________________________________________________________
UZBEKISTAN
CENTRAL KYZYLKUM
A SMALL ISOLATED RAVINE 9 KM SOUTH OF THE SETTLEMENT DAUGZTAU
DAUGYZTAU I
LATE TURONIAN STAGE
UPPERMOST GALLIC SUBEPOCH- SANTONIAN STAGE
LOWER SENONIAN SUBEPOCH
LOWER GULF EPOCH
EARLY LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
Incertae sedis
Gen. sp indet.
COHORT: ORNITHISCHIFORMES COOPER, 1985
MAGNORDER: ORNITHISCHIA SEELEY, 1888
SUPERORDER: NEORNITHISCHIA COOPER, 1985
GRANDORDER: ORNITHOPODA MARSH, 1871
MICRORDER: EUORNITHOPODA SERENO, 1986
ORDER: IGUANODONTIA DOLLO, 1888
INFRAORDER: STYRACOSTERNA SERENO, 1986
PARVORDER: HADROSAURIFORMES SERENO, 1997
SUPERFAMILY: HADROSAUROIDEA COPE, 1869 (SENSU SERENO, 1986)
Family: HADROSAURIDAE COPE, 1869
Gen. sp indet.
_____________________________________________________________________________________
UZBEKISTAN
CENTRAL PART OF KIZYLKUM DESERT
DZHARAK HUDUK LOCALITY
LOWER PART OF THE TAIKARSHIN BEDS
UPPER TURONIAN STAGE
UPPER GALLIC SUBEPOCH
LOWER GULF EPOCH
EARLY LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: AMPHIBIA LINNAEUS, 1758
SUBCLASS: LISSAMPHIBIA, HAECKEL, 1866?
SUPERORDER: BATRACHIA BRONGNIART, 1800
ORDER: CAUDATA SCOPOLI, 1777
SUBORDER: AMBYSTOMATOIDEA NOBLE, 1931
Family: SCAPHERPETONTIDAE Auffenberg & Goin, 1959
Eoscapherpeton asiaticum
SUPERORDER: SALIENTIA LAURENTI, 1768
Incertae sedis
Gen. sp indet.
ORDER: ANURA RAFINESQUE, 1815
Incertae sedis
*Aralobatrachus robustus
*Itemirella cretacea
SUBORDER: DISCOGLOSSOIDEI SOKOL, 1977
Family: DISCOGLOSSIDAE Guenther, 1859
Subfamily: ALYTINAE Fitzinger, 1943
Kizylkuma antiqua
SUBORDER: MESOBATRACHIA LAURENT, 1979
Superfamily: PELOBATOIDEA Stanius, 1856 , nom. corr. Spinar, 1971 (sensu Bolkay, 1919)
Family: PELOBATIDAE Stanius, 1856 , nom. corr. Spinar, 1971
Gen. sp indet.
_____________________________________________________________________________________
UZBEKISTAN
NAVOI VILOYAT
CENTRAL KYZYLKUM DESERT
ITEMIR-DZHARAKUDUK DEPRESSION
ITEMIR LOCALITY
SITE IT-01
DZHARAKUDUK FORMATION
CENOMANIAN STAGE
UPPER MIDDLE GALLIC SUBEPOCH
LOWERMOST GULF EPOCH
MIDDLE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
AVEPODA PAUL, 2002
SUPERORDER: NEOTHEROPODA BAKKER, 1986
GRANDORDER: TETANURAE GAUTHIER, 1986
MICRORDER: COELUROSAURIA HUENE, 1914
ORDER: MANIRAPTORFORMES HOLTZ, 1996
AVETHEROPODA PAUL, 2002
AVEPECTORA PAUL, 2002
SUBORDER: MANIRAPTORA GAUTHIER, 1986
INFRAORDER: PARVES SERENO, 1997
PARVORDER: DEINONYCHOSAURIA COLBERT & D. A. RUSSELL, 1969
Family: TROODONTIDAE Gilmore, 1924
*Urbacodon itemirensis
_____________________________________________________________________________________
UZEBEKISTAN
ITEMIR-DZHARAKUDUK DEPRESSION
DZHARAKUDUK
BISSEKT FORMATION (BISSEKTIYSKAYA SVITA)
UPPER TURONIAN STAGE
UPPERMOST GALLIC SUBEPOCH
LOWER GULF EPOCH
EARLY LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
AVEPODA PAUL, 2002
SUPERORDER: NEOTHEROPODA BAKKER, 1986
GRANDORDER: TETANURAE GAUTHIER, 1986
MICRORDER: COELUROSAURIA HUENE, 1914
ORDER: MANIRAPTORFORMES HOLTZ, 1996
AVETHEROPODA PAUL, 2002
AVEPECTORA PAUL, 2002
SUBORDER: MANIRAPTORA GAUTHIER, 1986
INFRAORDER: PARVES SERENO, 1997
PARVORDER: DEINONYCHOSAURIA COLBERT & D. A. RUSSELL, 1969
Family: TROODONTIDAE Gilmore, 1924
Urbacodon sp
_____________________________________________________________________________________
UZBEKISTAN
DZHARA-KHUDUK
UPPER TURONIAN STAGE
UPPER GALLIC SUBEPOCH
LOWER GULF EPOCH
EARLY LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: CRUROTARSI SERENO & ARCUCCI, 1990
COHORT: CROCODYLOTARSI BENTON & CLARK, 1988
MAGNORDER: SUCHIA KREBS, 1974
SUPERORDER: CROCODYLOMORPHA WALKER, 1968
GRANDORDER: CROCODILIFORMES HAY, 1930
Incertae sedis
Zholsuchus procerus
ORDER: PROTOSUCHIA emend MOOK, 1934
Superfamily: PROTOSUCHOIDEA Brown, 1933 (sensu Deraniyagala, 1939)
Family: PROTOSUCHIDAE Brown, 1933
Gen. sp indet.
MICRORDER: MESOEUROCROCODYLIA WHETSTONE & WHYBROW, 1983
ORDER: MESOSUCHIA EMEND HUXLEY, 1875
PARVORDER: METASUCHIA BENTON & CLARK, 1988
INFRAORDER: METAMESOSUCHIA HULKE, 1878
NEOSUCHIA CLARK, IN BENTON & CLARK, 1988
Family: PARALLIGAOTRIDAE Konjukova, 1954
Kanajusuchus borealis
*Shamosuchus occidentalis
ORDER: EUSUCHIA emend HUXLEY, 1875
INFRAORDER: CROCODYLIA emend GMELIN, 1877
PARVORDER: BREVIROSTRES
Superfamily: CROCODILOIDEA Cuvier, 1807 (sensu Fitzinger, 1826)
Family: CROCODYLIDAE Cuvier, 1807
Subfamily: CROCODYLIDINAE Cuvier, 1807
Tadzhikoesuchus kizylkumensis
_____________________________________________________________________________________
UZBEKISTAN
DZHARAKHUDUK (= BISSEKTY)
UPPER TURONIAN STAGE- LOWER CONIACIAN STAGE
UPPER GALLIC SUBEPOCH
LOWER GULF EPOCH
EARLY LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: CHELONOMORPHA KUHN, 1960
SUPERCOHORT: TESTUDINES LINNAEUS, 1758
COHORT: CASICHELYDIA emend GAFFNEY & MEYLAN, 1988
MAGANORDER: CRYPTODIRA GAFFNEY & MEYLAN, 1988
SUPERORDER: SELMACRYPTODIRA GAFFNEY, HUTCHISON, JENKINS & MEEKR, 1987
GRANDORDER: DAIOCRYPTODIRA GAFFNEY & MEYLAN, 1988
MICRORDER: CENTROCRYPTODIRA GAFFNEY & MEYLAN, 1988
ORDER: POLYCRPTODIRA GAFFNEY & MEYLAN, 1988
SUBORDER: PROCOELOCRYPTODIRA GAFFNEY & MEYLAN, 1988
INFRAORDER: CHELOMACRYPTODIRA GAFFNEY & MEYLAN, 1988
PARVORDER: TRIONYCHIDA GRAY, 1870
Superfamily: TRIONYCHIDEA Bell, 1828 (sensu Gray, 1870)
Family: DERMATEMYDIDAE Gray, 1870
Subfamily: LINDHOLMEMIDINAE Ckhikvadze, 1975 (in SHUVALOV & CKHIKVADZE, 1975)
Lindholmemys elegans
Family: ADOCIDAE Cope, 1870
Subfamily: SHACHEMYDINAE Khosatzky in Nesov & Khozatzkii, 1977
Shachemys baibolatica ancestralis
_____________________________________________________________________________________
UZBEKISTAN
KYZYLKUM DESERT
CBI-23
DZHYRAKUDUK LOCALITY
BISSEKTY FORMATION
TURONIAN STAGE
UPPERMOST GALLIC SUBEPOCH
LOWER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 184
COHORT: ORNITHISCHIFORMES COOPER, 1985
MAGNORDER: ORNITHISCHIA SEELEY, 1888
COHORT: ORNITHISCHIFORMES COOPER, 1985
MAGNORDER: ORNITHISCHIA SEELEY, 1888
SUPERORDER: NEORNITHISCHIA COOPER, 1985
GRANDORDER: ORNITHOPODA MARSH, 1871
MICRORDER: EUORNITHOPODA SERENO, 1986
ORDER: IGUANODONTIA DOLLO, 1888
INFRAORDER: STYRACOSTERNA SERENO, 1986
PARVORDER: HADROSAURIFORMES SERENO, 1997
SUPERFAMILY: HADROSAUROIDEA COPE, 1869 (SENSU SERENO, 1986)
Family: HADROSAURIDAE COPE, 1869
Incertae sedis
Gilmoreosaurus arkhangelskyi
_____________________________________________________________________________________
UZBEKISTAN
KYZYLKUM DESERT
CBI-24
DZHYRAKUDUK LOCALITY
BISSEKTY FORMATION
TURONIAN STAGE
UPPERMOST GALLIC SUBEPOCH
LOWER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 184
COHORT: ORNITHISCHIFORMES COOPER, 1985
MAGNORDER: ORNITHISCHIA SEELEY, 1888
COHORT: ORNITHISCHIFORMES COOPER, 1985
MAGNORDER: ORNITHISCHIA SEELEY, 1888
SUPERORDER: NEORNITHISCHIA COOPER, 1985
GRANDORDER: ORNITHOPODA MARSH, 1871
MICRORDER: EUORNITHOPODA SERENO, 1986
ORDER: IGUANODONTIA DOLLO, 1888
INFRAORDER: STYRACOSTERNA SERENO, 1986
PARVORDER: HADROSAURIFORMES SERENO, 1997
SUPERFAMILY: HADROSAUROIDEA COPE, 1869 (SENSU SERENO, 1986)
Family: HADROSAURIDAE COPE, 1869
Incertae sedis
Gilmoreosaurus arkhangelskyi
_____________________________________________________________________________________
UZBEKISTAN
KYZYLKUM DESERT
CBI-27
DZHYRAKUDUK LOCALITY
BISSEKTY FORMATION
TURONIAN STAGE
UPPERMOST GALLIC SUBEPOCH
LOWER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 184
COHORT: ORNITHISCHIFORMES COOPER, 1985
MAGNORDER: ORNITHISCHIA SEELEY, 1888
COHORT: ORNITHISCHIFORMES COOPER, 1985
MAGNORDER: ORNITHISCHIA SEELEY, 1888
SUPERORDER: NEORNITHISCHIA COOPER, 1985
GRANDORDER: ORNITHOPODA MARSH, 1871
MICRORDER: MARGINOCEPHALIA SERENO, 1986
ORDER: CERATOPIA MARSH, 1890
SUBORDER: NEOCERATOPIA SERENO, 1986
INFRAORDER: CORONOSAURIA SERENO, 1986
PARVORDER: CERATOPOMORPHA Wolfe & Kirkland, 1998 (emend)
Superfamily: CERATOPOIDEA Hay, 1902 (emend)
Incertae sedis
Turanoceratops tardabilis
_____________________________________________________________________________________
UZBEKISTAN
KIZYLKUM DESERT
DZHYRAKUDUK LOCALITY
CBI-28
BISSEKTY FORMATION
LATE TURONIAN STAGE
UPPER GALLIC SUBEPOCH
LOWER GULF EPOCH
EARLY LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: CHELONOMORPHA KUHN, 1960
SUPERCOHORT: TESTUDINES LINNAEUS, 1758
COHORT: CASICHELYDIA emend GAFFNEY & MEYLAN, 1988
MAGANORDER: CRYPTODIRA GAFFNEY & MEYLAN, 1988
SUPERORDER: SELMACRYPTODIRA GAFFNEY, HUTCHISON, JENKINS & MEEKR, 1987
GRANDORDER: DAIOCRYPTODIRA GAFFNEY & MEYLAN, 1988
MICRORDER: CENTROCRYPTODIRA GAFFNEY & MEYLAN, 1988
ORDER: POLYCRPTODIRA GAFFNEY & MEYLAN, 1988
SUBORDER: PROCOELOCRYPTODIRA GAFFNEY & MEYLAN, 1988
INFRAORDER: CHELOMACRYPTODIRA GAFFNEY & MEYLAN, 1988
PARVORDER: TRIONYCHIDA GRAY, 1870
Superfamily: TRIONYCHIDEA Bell, 1828 (sensu Gray, 1870)
Family: TRIONYCHIDAE Bell, 1828
*Khunnuchelys kizlkumensis
SUBCLASS: DIAPSIDA OSBORN, 1903
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
AVEPODA PAUL, 2002
SUPERORDER: NEOTHEROPODA BAKKER, 1986
GRANDORDER: TETANURAE GAUTHIER, 1986
MICRORDER: COELUROSAURIA HUENE, 1914
ORDER: MANIRAPTORFORMES HOLTZ, 1996
AVETHEROPODA PAUL, 2001
SUBORDER: COELURIA MARSH, 1881
Superfamily: TYRANNOSAUROIDEA Osborn, 1906 (sensu Bonaparte et. al., 1990)
Family: ALIORAMIDAE Olshevsky, Ford & Yamamoto, 1995
Alectrosaurus sp
COHORT: ORNITHISCHIFORMES COOPER, 1985
MAGNORDER: ORNITHISCHIA SEELEY, 1888
COHORT: ORNITHISCHIFORMES COOPER, 1985
MAGNORDER: ORNITHISCHIA SEELEY, 1888
SUPERORDER: NEORNITHISCHIA COOPER, 1985
GRANDORDER: ORNITHOPODA MARSH, 1871
MICRORDER: EUORNITHOPODA SERENO, 1986
ORDER: IGUANODONTIA DOLLO, 1888
INFRAORDER: STYRACOSTERNA SERENO, 1986
PARVORDER: HADROSAURIFORMES SERENO, 1997
SUPERFAMILY: HADROSAUROIDEA COPE, 1869 (SENSU SERENO, 1986)
Family: HADROSAURIDAE COPE, 1869
Incertae sedis
Gilmoreosaurus arkhangelskyi
MICRORDER: MARGINOCEPHALIA SERENO, 1986
ORDER: CERATOPIA MARSH, 1890
SUBORDER: NEOCERATOPIA SERENO, 1986
INFRAORDER: CORONOSAURIA SERENO, 1986
PARVORDER: CERATOPOMORPHA Wolfe & Kirkland, 1998 (emend)
Superfamily: CERATOPOIDEA Hay, 1902 (emend)
Incertae sedis
Turanoceratops tardabilis
SUPERORDER: EURYOPDA SERENO, 1986
GRANDORDER: ARMATOSAURIA ZHAO, 1983 (emend)
ORDER: ANKYLOSAURIA OSBORN, 1923
INFRAORDER: ANKYLOSAURA OSBORN, 1923
Family: ANKYLOSAURIDAE Brown, 1908
Incertae sedis
Gen. sp indet.
_____________________________________________________________________________________
UZBEKISTAN
CENTRAL KYZYLKUM DESERT
NORTH OF
SITE CBI-4V
DZHYRAKUDUK LOCALITY
LOWER PART OF BISSEKTY FORMATION
UPPER TURONIAN STAGE
UPPERMOST GALLIC SUBEPOCH
LOWER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
AVEPODA PAUL, 2002
SUPERORDER: NEOTHEROPODA BAKKER, 1986
GRANDORDER: TETANURAE GAUTHIER, 1986
MICRORDER: COELUROSAURIA HUENE, 1914
ORDER: MANIRAPTORFORMES HOLTZ, 1996
AVETHEROPODA PAUL, 2001
SUBORDER: COELURIA MARSH, 1881
Superfamily: TYRANNOSAUROIDEA Osborn, 1906 (sensu Bonaparte et. al., 1990)
Family: ALIORAMIDAE Olshevsky, Ford & Yamamoto, 1995
Alectrosaurus sp
COHORT: ORNITHISCHIFORMES COOPER, 1985
MAGNORDER: ORNITHISCHIA SEELEY, 1888
COHORT: ORNITHISCHIFORMES COOPER, 1985
MAGNORDER: ORNITHISCHIA SEELEY, 1888
SUPERORDER: NEORNITHISCHIA COOPER, 1985
GRANDORDER: ORNITHOPODA MARSH, 1871
MICRORDER: EUORNITHOPODA SERENO, 1986
ORDER: HYPSILOPHODONTIA COOPER, 1983
Family: HYPSILOPHODONTIDAE Dollo, 1882
Incertae sedis? Or new families? (pending further research)
Gen. sp indet
_____________________________________________________________________________________
UZBEKISTAN
CENTRAL KYZYLKUM DESERT
NORTH OF
CBI-5A
DZHYRAKUDUK LOCALITY
LOWER PART OF BISSEKTY FORMATION
UPPER TURONIAN STAGE
UPPERMOST GALLIC SUBEPOCH
LOWER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
Incertae sedis
Gen. sp indet.
AVEPODA PAUL, 2002
SUPERORDER: NEOTHEROPODA BAKKER, 1986
GRANDORDER: TETANURAE GAUTHIER, 1986
MICRORDER: COELUROSAURIA HUENE, 1914
ORDER: MANIRAPTORFORMES HOLTZ, 1996
AVETHEROPODA PAUL, 2001
SUBORDER: COELURIA MARSH, 1881
Superfamily: TYRANNOSAUROIDEA Osborn, 1906 (sensu Bonaparte et. al., 1990)
Family: ALIORAMIDAE Olshevsky, Ford & Yamamoto, 1995
Alectrosaurus sp
_____________________________________________________________________________________
UZBEKISTAN
KYZYLKUM DESERT
CDZH-23
DZHYRAKUDUK LOCALITY
BISSEKTY FORMATION
TURONIAN STAGE
UPPERMOST GALLIC SUBEPOCH
LOWER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 184
COHORT: ORNITHISCHIFORMES COOPER, 1985
MAGNORDER: ORNITHISCHIA SEELEY, 1888
COHORT: ORNITHISCHIFORMES COOPER, 1985
MAGNORDER: ORNITHISCHIA SEELEY, 1888
SUPERORDER: NEORNITHISCHIA COOPER, 1985
GRANDORDER: ORNITHOPODA MARSH, 1871
MICRORDER: EUORNITHOPODA SERENO, 1986
ORDER: IGUANODONTIA DOLLO, 1888
INFRAORDER: STYRACOSTERNA SERENO, 1986
PARVORDER: HADROSAURIFORMES SERENO, 1997
SUPERFAMILY: HADROSAUROIDEA COPE, 1869 (SENSU SERENO, 1986)
Family: HADROSAURIDAE COPE, 1869
Incertae sedis
Gilmoreosaurus arkhangelskyi
_____________________________________________________________________________________
UZBEKISTAN
KIZYLKUM DESERT
DZHYRAKUUK
LOCALITY CDZH-14A
BISSEKTY FORMATION
CONIACIAN STAGE
LOWER SENONIAN SUBEPOCH
MIDDLE GULF EPOCH
EARLY CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
AVEPODA PAUL, 2002
SUPERORDER: NEOTHEROPODA BAKKER, 1986
GRANDORDER: TETANURAE GAUTHIER, 1986
MICRORDER: COELUROSAURIA HUENE, 1914
ORDER: MANIRAPTORFORMES HOLTZ, 1996
AVETHEROPODA PAUL, 2001
SUBORDER: COELURIA MARSH, 1881
Superfamily: TYRANNOSAUROIDEA Osborn, 1906 (sensu Bonaparte et. al., 1990)
Family: ALIORAMIDAE Olshevsky, Ford & Yamamoto, 1995
Alectrosaurus sp
_____________________________________________________________________________________
UZBEKISTAN
KIZYLKUM DESERT
DZHYRAKUUK
LOCALITY CDZH-16
BISSEKTY FORMATION
TURONIAN STAGE
UPPERMOST GALLIC SUBEPOCH
LOWER GULF EPOCH
EARLY LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
AVEPODA PAUL, 2002
SUPERORDER: NEOTHEROPODA BAKKER, 1986
GRANDORDER: TETANURAE GAUTHIER, 1986
ORDER: MANIRAPTORFORMES HOLTZ, 1996
AVETHEROPODA PAUL, 2001
SUBORDER: COELURIA MARSH, 1881
Incertae sedis
Gen. sp indet.
Superfamily: TYRANNOSAUROIDEA Osborn, 1906 (sensu Bonaparte et. al., 1990)
Family: ALIORAMIDAE Olshevsky, Ford & Yamamoto, 1995
Alectrosaurus sp
SUBORDER: MANIRAPTORA GAUTHIER, 1986
AVEPECTORA PAUL, 2001
INFRAORDER: OVIRAPTOROSAURIA BARSBOLD, 1976
Family: OVIRAPTORIDAE Barsbold, 1976
Gen. sp indet.
INFRAORDER: PARVES SERENO, 1997
PARVORDER: DEINONYCHOSAURIA COLBERT & D. A. RUSSELL, 1969
Family: TROODONTIDAE Gilmore, 1924
Urbacodon sp
_____________________________________________________________________________________
UZBEKISTAN
KIZYLKUM DESERT
DZHYRAKUDUK LOCALITY
CBI-28
BISSEKTY FORMATION
LATE TURONIAN STAGE
UPPER GALLIC SUBEPOCH
LOWER GULF EPOCH
EARLY LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: CHELONOMORPHA KUHN, 1960
SUPERCOHORT: TESTUDINES LINNAEUS, 1758
COHORT: CASICHELYDIA emend GAFFNEY & MEYLAN, 1988
MAGANORDER: CRYPTODIRA GAFFNEY & MEYLAN, 1988
SUPERORDER: SELMACRYPTODIRA GAFFNEY, HUTCHISON, JENKINS & MEEKR, 1987
GRANDORDER: DAIOCRYPTODIRA GAFFNEY & MEYLAN, 1988
MICRORDER: CENTROCRYPTODIRA GAFFNEY & MEYLAN, 1988
ORDER: POLYCRPTODIRA GAFFNEY & MEYLAN, 1988
SUBORDER: PROCOELOCRYPTODIRA GAFFNEY & MEYLAN, 1988
INFRAORDER: CHELOMACRYPTODIRA GAFFNEY & MEYLAN, 1988
PARVORDER: TRIONYCHIDA GRAY, 1870
Superfamily: TRIONYCHIDEA Bell, 1828 (sensu Gray, 1870)
Family: TRIONYCHIDAE Bell, 1828
*Khunnuchelys kizlkumensis
SUBCLASS: DIAPSIDA OSBORN, 1903
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
AVEPODA PAUL, 2002
SUPERORDER: NEOTHEROPODA BAKKER, 1986
GRANDORDER: TETANURAE GAUTHIER, 1986
MICRORDER: COELUROSAURIA HUENE, 1914
ORDER: MANIRAPTORFORMES HOLTZ, 1996
AVETHEROPODA PAUL, 2001
SUBORDER: COELURIA MARSH, 1881
Superfamily: TYRANNOSAUROIDEA Osborn, 1906 (sensu Bonaparte et. al., 1990)
Family: ALIORAMIDAE Olshevsky, Ford & Yamamoto, 1995
Alectrosaurus sp
COHORT: ORNITHISCHIFORMES COOPER, 1985
MAGNORDER: ORNITHISCHIA SEELEY, 1888
SUPERORDER: EURYOPDA SERENO, 1986
GRANDORDER: ARMATOSAURIA ZHAO, 1983 (emend)
ORDER: ANKYLOSAURIA OSBORN, 1923
INFRAORDER: ANKYLOSAURA OSBORN, 1923
Family: ANKYLOSAURIDAE Brown, 1908
Incertae sedis
Gen. sp indet.
_____________________________________________________________________________________
UZBEKISTAN
KYZYLKUM DESERT
SITE CDZH-17
LOWER BISSEKTY FORMATION
CONIACIAN STAGE
LOWER SENONIAN SUBEPOCH
LOWER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
AVEPODA PAUL, 2002
SUPERORDER: NEOTHEROPODA BAKKER, 1986
GRANDORDER: TETANURAE GAUTHIER, 1986
MICRORDER: COELUROSAURIA HUENE, 1914
ORDER: MANIRAPTORFORMES HOLTZ, 1996
AVETHEROPODA PAUL, 2001
SUBORDER: COELURIA MARSH, 1881
Superfamily: TYRANNOSAUROIDEA Osborn, 1906 (sensu Bonaparte et. al., 1990)
Incertae sedis
Timurlengia euotica
Family: ALIORAMIDAE Olshevsky, Ford & Yamamoto, 1995
Alectrosaurus sp
AVEPECTORA PAUL, 2001
SUBORDER: ORNITHOMIMOSAURIA BARSBOLD, 1976
Family: ORNITHOMIMIDAE Marsh, 1890
Incertae sedis
Gen. sp indet.
SUBORDER: MANIRAPTORA GAUTHIER, 1986
INFRAORDER: OVIRAPTOROSAURIA BARSBOLD, 1976
Family: CAENAGNATHIDAE Sternberg, 1940
*Caenagnathasia martinsoni
Family: OVIRAPTORIDAE Barsbold, 1976
Gen. sp indet.
COHORT: ORNITHISCHIFORMES COOPER, 1985
SUPERORDER: SAUROPODOMORPHA HUENE, 1932
GRANDORDER: BRONTOSAURIA OLSHEVSKY, 1991 (emend)
MICRORDER: SAUROPODA MARSH, 1878
ORDER: EUSAUROPODA
SUBORDER: NEOSAUROPODA BONAPARTE, 1986
INFRAORDER: MACRONARIA WILSON & SERENO, 1998
PARVORDER: TITANOSAURIFORMES SALGADO ET AL, 1997
Superfamily: TITANOSAUROIDEA Lydekker, 1885 (sensu Upchurch, 1999)
Family: TITANOSAURIDAE Lydekker, 1885
Incertae sedis
Gen. Sp indet.
MAGNORDER: ORNITHISCHIA SEELEY, 1888
SUPERORDER: NEORNITHISCHIA COOPER, 1985
GRANDORDER: ORNITHOPODA MARSH, 1871
MICRORDER: EUORNITHOPODA SERENO, 1986
ORDER: IGUANODONTIA DOLLO, 1888
INFRAORDER: STYRACOSTERNA SERENO, 1986
PARVORDER: HADROSAURIFORMES SERENO, 1997
SUPERFAMILY: HADROSAUROIDEA COPE, 1869 (SENSU SERENO, 1986)
Family: HADROSAURIDAE COPE, 1869
Subfamily: LAMBEOSAURINAE Parks, 1923
Tribe: PARASAUROLOPHINI Brett-Surman vide Chapman & Brett-Surman, 1990
Bactrosaurus kysylkumensis = Cionodon? kysylkumense
CLASS: MAMMALIA LINNAEUS, 1758
Incertae sedis
Gen. sp indet.
_____________________________________________________________________________________
UZBEKISTAN
CENTRAL KYZYLKUM DESERT
CDZH-17a
LOWER BISSEKTY FORMATION
CONIACIAN STAGE
LOWER SENONIAN SUBEPOCH
LOWER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: AMPHIBIA LINNAEUS, 1758
SUBCLASS: LISSAMPHIBIA, HAECKEL, 1866?
SUPERORDER: BATRACHIA BRONGNIART, 1800
ORDER: CAUDATA SCOPOLI, 1777
SUBORDER: PORTEOIDEA (NOBEL 1931) EMEND EDWARDS, 1976
Family: BATRACHOSAUROIDIDAE Auffenberg, 1958
Mynbulakia surgayi
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
INFRACLASS: NEODIAPSIDA BENTON, 1985
LEGION: LEPIDOSAUROMORPHA DUMERIL & BIBRON, 1839
SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)
MAGNORDER: SQUAMATA OPPEL, 1811
SUPERORDER: LACERTILIA OWEN, 1842
GRANDORDER: ASCALABOTA GILMORE, 1928
ORDER: IGUANIA LATREILLE, 1825
SUBORDER: DRACOMORPHA NORTHCUTT, 1978
INFRAORDER: TEIUMORPHA ALIFANOV, 2000
Family: TEIIDAE Gray, 1827
Gen. sp indet.
GRANDORDER: SCINCOGEKKONOMORPHA SUKHANOV, 1961
ICRORDER: SCINCOMORPHA CAMP, 1923
ORDER: LACERTOGNATHA EMEND ALIFANOV, 2000
SUBORDER: GEKKOTA CUVIER, 1817
Incertae sedis
Gen. sp indet.
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
AVEPODA PAUL, 2002
SUPERORDER: NEOTHEROPODA BAKKER, 1986
GRANDORDER: TETANURAE GAUTHIER, 1986
MICRORDER: COELUROSAURIA HUENE, 1914
ORDER: SEGNOSAURIA BARSBOLD & PERLE, 1980
Family: SEGNOSAURIDAE Perle, 1979
Gen. sp indet.
ORDER: MANIRAPTORFORMES HOLTZ, 1996
AVETHEROPODA PAUL, 2001
SUBORDER: COELURIA MARSH, 1881
Superfamily: TYRANNOSAUROIDEA Osborn, 1906 (sensu Bonaparte et. al., 1990)
Incertae sedis
Timurlengia euotica
COHORT: ORNITHISCHIFORMES COOPER, 1985
MAGNORDER: ORNITHISCHIA SEELEY, 1888
SUPERORDER: NEORNITHISCHIA COOPER, 1985
GRANDORDER: ORNITHOPODA MARSH, 1871
MICRORDER: EUORNITHOPODA SERENO, 1986
ORDER: HYPSILOPHODONTIA COOPER, 1983
Family: HYPSILOPHODONTIDAE Dollo, 1882
Incertae sedis? Or new families? (pending further research)
Gen. sp indet
ORDER: IGUANODONTIA DOLLO, 1888
INFRAORDER: STYRACOSTERNA SERENO, 1986
PARVORDER: HADROSAURIFORMES SERENO, 1997
SUPERFAMILY: HADROSAUROIDEA COPE, 1869 (SENSU SERENO, 1986)
Family: HADROSAURIDAE COPE, 1869
Incertae sedis
Gilmoreosaurus arkhangelskyi
Subfamily: LAMBEOSAURINAE Parks, 1923
Tribe: PARASAUROLOPHINI Brett-Surman vide Chapman & Brett-Surman, 1990
Bactrosaurus kysylkumensis = Cionodon? kysylkumense
CLASS: MAMMALIA LINNAEUS, 1758
SUBCLASS: PROTOTHERIA GILL, 1872
INFRACLASS: TRECHNOTHERIA McKENNA, 1975
LEGION: CLADOTHERIA McKENNA, 1975
SUBLEGION: ZATHERIA MCKENNA, 1975
INFRALEGION: TRIBOSPHENIDA (McKENNA, 1975) McKENNA & BELL, 1997
SUPERCOHORT: THERIA (PARKER & HASWELL, 1987) McKENNA & BELL, 1997
ORDER: DELTATHEROIDA KIELAN-JAWOROWSKA, 1982
Family: DELTATHERIDIIDAE Gregory & Simpson, 1926
Gen. sp indet.
COHORT: PLACENTALIA (OWEN, 1837) McKENNA & BELL, 1997
MAGNORDER: EPITHERIA (McKENNA, 1975) McKENNA & BELL, 1997
ORDER: KENNALESTIDA McKENNA, 1975
Incertae sedis
*Daulestes kulbeckensis
Daulestes inobservabilis
= *Taslestes inobservabilisSUPERORDER: PREPTOTHERIA McKENNA, 1975
GRANDORDER: UNGULATOMORPHA ARCHIBALD, 1996
Family: ZHELESTIDAE Nesov, 1990
*Aspanlestes aptap
Sorlestes budan
cf. Zhelestes sp
Gen. sp indet.
Family: KULBECKIIDAE NESOV, 1993
Kulbeckia kulbecke
= *Kulbeckia rara
_____________________________________________________________________________________
UZBEKISTAN
KYZYLKUM DESERT
SITE CDZH-17g
DZHYRAKUDUK LOCALITY
BISSEKTY FORMATION
LATE TURONIAN STAGE
UPPERMOST GALLIC SUBEPOCH
LOWER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 184
COHORT: THEROPODA EMEND MARSH, 1881
AVEPODA PAUL, 2002
SUPERORDER: NEOTHEROPODA BAKKER, 1986
GRANDORDER: TETANURAE GAUTHIER, 1986
MICRORDER: COELUROSAURIA HUENE, 1914
ORDER: MANIRAPTORFORMES HOLTZ, 1996
AVETHEROPODA PAUL, 2001
SUBORDER: COELURIA MARSH, 1881
Superfamily: TYRANNOSAUROIDEA Osborn, 1906 (sensu Bonaparte et. al., 1990)
Family: TYRANNOSAURIDAE Osborn, 1906
Gen. sp indet.
COHORT: ORNITHISCHIFORMES COOPER, 1985
MAGNORDER: ORNITHISCHIA SEELEY, 1888
COHORT: ORNITHISCHIFORMES COOPER, 1985
MAGNORDER: ORNITHISCHIA SEELEY, 1888
SUPERORDER: NEORNITHISCHIA COOPER, 1985
GRANDORDER: ORNITHOPODA MARSH, 1871
MICRORDER: EUORNITHOPODA SERENO, 1986
ORDER: HYPSILOPHODONTIA COOPER, 1983
Family: HYPSILOPHODONTIDAE Dollo, 1882
Incertae sedis? Or new families? (pending further research)
Gen. sp indet
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UZBEKISTAN
DZHYRAKUDUK LOCALITY
CDZH-17v
TURONIAN STAGE
UPPERMOST GALLIC SUBEPOCH
LOWER SENONIAN SUBEPOCH
LOWER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
AVEPODA PAUL, 2002
SUPERORDER: NEOTHEROPODA BAKKER, 1986
GRANDORDER: TETANURAE GAUTHIER, 1986
MICRORDER: COELUROSAURIA HUENE, 1914
ORDER: MANIRAPTORFORMES HOLTZ, 1996
AVETHEROPODA PAUL, 2001
SUBORDER: COELURIA MARSH, 1881
Superfamily: TYRANNOSAUROIDEA Osborn, 1906 (sensu Bonaparte et. al., 1990)
Family: TYRANNOSAURIDAE Osborn, 1906
Gen. sp indet.
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UZBEKISTAN
CENTRAL KIZYLKKUM DESERT
DZHYRAKUDUK
VBI-14
MIDDLE OF THE BISSEKTY FORMATION
CONIACIAN STAGE
LOWER SENONIAN SUBEPOCH
MIDDLE GULF EPOCH
EARLY CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: AVEMETATARSALIA BENTON, 1999
COHORT: PROLACERTIFORMES CAMP, 1945 NOVA
SUPERORDER: TAPINOPLATIA PETERS, 2000
GRANDORDER: CHARACIOPODA PETERS, 2000
MICRORDER: FENESTRASAURIA PETERS, 2000
ORDER: PTEROSAURIA KAUP, 1834
SUBORDER: EUPTERODACTYLOIDEA BENNETT, 1994
Superfamily: AZHDARCHIDEA Nesov, 1984
Family: AZHDARCHIDAE Nesov, 1984 emend Padian, 1986
Azhdarcho lancicollis
_____________________________________________________________________________________
UZBEKISTAN
CENTRAL KYZYLKUM
RAVINE NEAR SAND-PITS 7 KM NORTH-NORTH-WEST OF THE SETTLEMENT OF DAUGYZTAU
BELTAU (DAUGYZTAU III)
RED SANDSTONE.
TURONIAN STAGE
UPPERMOST GALLIC SUBEPOCH-SANTONIAN STAGE
LOWER SENONIAN SUBEPOCH
LOWER GULF EPOCH
LATE CRETACEOUS
VETEROVATA
OOSUPERCOHORT: DINOSAURIOVUM VIALOV, 1975
Incertae sedis
Gen. sp indet.
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