SOUTH DAKOTA
BUTTE COUNTY
BELLE FOURCHE RESERVOIR
SHELL/VERTEBRATE LAG BED
AT THE GREENHORN/CARLILE FORMATIONAL CONTACT
COLLIGONICERAS WOLLGARI ZONE
EARLY MIDDLE TURONIAN
EARLY LATE CRETACEOUS
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: CHELONOMORPHA KUHN, 1960
SUPERCOHORT: TESTUDINES LINNAEUS, 1758
COHORT: CASICHELYDIA emend GAFFNEY & MEYLAN, 1988
MAGANORDER: CRYPTODIRA GAFFNEY & MEYLAN, 1988
SUPERORDER: SELMACRYPTODIRA GAFFNEY, HUTCHISON, JENKINS & MEEKR, 1987
GRANDORDER: DAIOCRYPTODIRA GAFFNEY & MEYLAN, 1988
MICRORDER: CENTROCRYPTODIRA GAFFNEY & MEYLAN, 1988
ORDER: POLYCRPTODIRA GAFFNEY & MEYLAN, 1988
SUBORDER: PROCOELOCRYPTODIRA GAFFNEY & MEYLAN, 1988
INFRAORDER: CHELONOIDEA GAFFNEY & MEYLAN, 1988
Superfamily: EUCHELONIOIDEA Gaffney & Meylan, 1988
Family: CHELONIIDAE Oppel, 1811
Gen. sp indet.
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SOUTH DAKOTA
BUTTE COUNTY
BELLE FOURCHE RESERVOIR
GREENHORN FORMATION
CALCARENITE BED
UPPERMOST FOURTH MEMBER
COLLIGNONICERAS WOLLGARI ZONE
EARLY MIDDLE TURONIAN STAGE
UPPERMOST GALLIC SUBEPOCH
LOWER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
INFRACLASS: NEODIAPSIDA BENTON, 1985
LEGION: LEPIDOSAUROMORPHA BENTON, 1983
SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)
MAGNORDER: SQUAMATA, OPPEL, 1811
SUPERORDER: LACERTILIA OWEN, 1842
GRANDORDER: SCINCOGEKKONOMORPHA SUKHANOV, 1961
MICRORDER: SCINCOMORPHA CAMP, 1923
ORDER: ANGUIMORPHA FURBRINGER, 1900
SUBORDER: DIPLOGLOSSA COPE, 1864
INFRAORDER: THECAGLOSSA COPE, 1890
PARVORDER: MONSTERSAURIA NORELL & GAO, 1977
Superfamily: VARANOIDEA Gray, 1827 (sensu Camp, 1923)
Family: MOSASAURIDAE Gervais, 1853
Incertae sedis
Gen. sp indet.
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SOUTH DAKOTA
BUTTE COUNTY
BELLE FOURCHE RESERVOIR
GREENHORN FORMATION
VERTEBRATE LAG ON TOP OF OYSTER BED ABOUT 6 “ BELOW GREENHORN/CARLILE BOUNDARY
UPPERMOST FORUTH MEMBER
COLLIGNONICERAS WOLLGARI ZONE
EARLY MIDDLE TURONIAN STAGE
UPPERMOST GALLIC SUBEPOCH
LOWER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
INFRACLASS: NEODIAPSIDA BENTON, 1985
LEGION: LEPIDOSAUROMORPHA BENTON, 1983
SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)
MAGNORDER: SQUAMATA, OPPEL, 1811
SUPERORDER: LACERTILIA OWEN, 1842
GRANDORDER: SCINCOGEKKONOMORPHA SUKHANOV, 1961
MICRORDER: SCINCOMORPHA CAMP, 1923
ORDER: ANGUIMORPHA FURBRINGER, 1900
SUBORDER: DIPLOGLOSSA COPE, 1864
INFRAORDER: THECAGLOSSA COPE, 1890
PARVORDER: MONSTERSAURIA NORELL & GAO, 1977
Superfamily: VARANOIDEA Gray, 1827 (sensu Camp, 1923)
Family: MOSASAURIDAE Gervais, 1853
Incertae sedis
Gen. sp indet.
INFRACLASS: EURYAPSIDA COLBERT, 1945
SUPERORDER: SAUROPTERYGIA OWEN 1860
GRANDORDER: EOSAUROPTERYGIA emend RIEPPEL, 1994
ORDER: PLESIOSAURIA OWEN, 1840 NON BLAINVILLE, 1835
SUBORDER: PLESIOSAURIA OWEN, 1840 NON BLAINVILLE, 1835
INFRAORDER: PLESIOSAUROMORPHA NOVA
EUPLESIOSAURIA O’KEEFE, 2001
Superfamily: CRYPTOCLEIDOIDEA Williston, 1925
TRICLEDIA O’KEEFE , 2001
Family: POLYCOTYLIDAE Cope, 1869
Gen. sp indet.
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SOUTH DAKOTA
BUTTE COUNTY
BELLE FOURCHE RESERVOIR
GREENHORN FORMATION
BASAL ORMAN LAKE MEMBER
DUNVEGANOCERAS PONDI ZONE
LOWER UPPER CENOMANIAN STAGE
UPPER GALLIC SUBEPOCH
LOWER GULF EPOCH
MIDDLE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
INFRACLASS: NEODIAPSIDA BENTON, 1985
LEGION: LEPIDOSAUROMORPHA BENTON, 1983
SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)
MAGNORDER: SQUAMATA, OPPEL, 1811
SUPERORDER: LACERTILIA OWEN, 1842
GRANDORDER: SCINCOGEKKONOMORPHA SUKHANOV, 1961
MICRORDER: SCINCOMORPHA CAMP, 1923
ORDER: ANGUIMORPHA FURBRINGER, 1900
SUBORDER: DIPLOGLOSSA COPE, 1864
INFRAORDER: THECAGLOSSA COPE, 1890
PARVORDER: MONSTERSAURIA NORELL & GAO, 1977
Superfamily: VARANOIDEA Gray, 1827 (sensu Camp, 1923)
Family: MOSASAURIDAE Gervais, 1853
Subfamily: DOLICHOSAURIDAE Gervais, 1852
Tribe: DOLICHOSAURINI Gervais, 1852
Coniasaurus crassidens
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SOUTH DAKOTA
BUTTE COUNTY
BELLE FOURCHE RESERVOIR
GREENHORN FORMATION
BASAL CALCARENITE
ORMAN LAKE MEMBER
DUNVEGANOCERAS PONDI ZONE
LOWER UPPER CENOMANIAN STAGE
UPPER GALLIC SUBEPOCH
LOWER GULF EPOCH
MIDDLE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
INFRACLASS: NEODIAPSIDA BENTON, 1985
LEGION: LEPIDOSAUROMORPHA BENTON, 1983
SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)
MAGNORDER: SQUAMATA, OPPEL, 1811
SUPERORDER: LACERTILIA OWEN, 1842
GRANDORDER: SCINCOGEKKONOMORPHA SUKHANOV, 1961
MICRORDER: SCINCOMORPHA CAMP, 1923
ORDER: ANGUIMORPHA FURBRINGER, 1900
SUBORDER: DIPLOGLOSSA COPE, 1864
INFRAORDER: THECAGLOSSA COPE, 1890
PARVORDER: MONSTERSAURIA NORELL & GAO, 1977
Superfamily: VARANOIDEA Gray, 1827 (sensu Camp, 1923)
Family: MOSASAURIDAE Gervais, 1853
Subfamily: DOLICHOSAURIDAE Gervais, 1852
Tribe: DOLICHOSAURINI Gervais, 1852
Coniasaurus crassidens
INFRACLASS: EURYAPSIDA COLBERT, 1945
SUPERORDER: SAUROPTERYGIA OWEN, 1860
GRANDORDER: EOSAUROPTERYGIA emend RIEPPEL, 1994
ORDER: PLESIOSAURIA OWEN, 1840 NON BLAINVILLE, 1835
SUBORDER: PLESIOSAURIA OWEN, 1840 NON BLAINVILLE, 1835
INFRAORDER: PLIOSAUROMORPHA NOVA
Superfamily: PLIOSAUROIDEA Seeley, 1874
Family: PLIOSAURIDAE Seeley, 1874
Polyptychodon interruptus
Family: Nova?
Gen sp nova?
(cross between pliosaurs and polycotylids)
Plesiosauria gen sp indet
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