EUROPEAN RUSSIA
ARKHANGELSK REGION
MEZENSKII DISTRICT
GLYADNAYA SHCHEL’YA LOCALITY
KRASNOSHCHEL’YA FORMATION
URZHUMIAN HORIZON
MEZEN FAUNAL ASSEMBLAGE
LOWER TATARIAN AGE
CAPITANIAN STAGE
MIDDLE ZECHSTEIN EPOCH
LATE MIDDLE PERMIAN
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: AMPHIBIA LINNAEUS, 1758
NEOTETRAPODA
SUBCLASS: REPTILIOMORPHA SAVE-SODERBERGH, 1934
ORDER: SEYMOURIAMORPHA WATSON, 1917
SUBORDER: NYCTEROLETERIDAE TATARINOV, 1972
Family: NYCTEROLETERIDAE Romer, 1956
Subfamily: NYCTEROLECTERINAE Romer, 1956
Macroleter poezicus
Subfamily: BASHKYROLETERINAE Ivachnenko, 1997
*Bashkyroleter mesensis
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: PARAREPTILIA OLSON 1947
INFRACLASS: SAUROPSIDA HUXLEY, 1864
ORDER: PROCOLOPHONOMORPHA ROMER, 1964
SUBORDER: PROCOLPHOMORPHA SEELEY, 1888
Family: NYCTIPHRURETIDAE Efremov, 1938
*Nyctiphruretus acudens
SUBCLASS: DIAPSIDA OSBORN, 1904
Incertae sedis
*Lanthanolania ivakhenennkoi
CLADE NAME: NEOVARANOPSIA SPINDLER, WERNEBURG, SCHNEIDER, LUTHARDT, ANNACKER & RÖßLER, 2019
Family: VARANOPSEIDAE Romer & Price, 1940
Subfamily: MESENOSAURINAE Romer 1956
Mesenosaurus romeri
SUBCLASS: SYNAPSIDA OSBORN, 1903
SUPERORDER: THERASPIDA BROOM, 1905
ORDER: EOTHERIODONTIA OLSON, 1962
Incertae sedis
Family: NIKKASAURIDAE Ivakhnenko, 2000
*Nikkasaurus tatarinovi
*Reiszia gubini
INFRAORDER: EOTITANOSUCHIA BOONSTRA, 1963
Family: NIIAFTASUCHIDAE Ivakhenko, 1990
Niaftasuchus zekkeli
Family: ALRAUSUCHIDAE Ivakhenko, 2008
Alrausuchus tagax
_____________________________________________________________________________________
EUROPEAN RUSSIA
ARKHANGELSK REGION
MEZENSKII DISTRICT
DOROGAYA GORA LOCALITY
BIARMIAN SERIES
URZHUMIAN STAGE
URZHUMIAN HORIZON
MEZEN FAUNAL ASSEMBLAGE
LOWER TATARIAN AGE
CAPITANIAN STAGE
MIDDLE ZECHSTEIN EPOCH
LATE MIDDLE PERMIAN
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: AMPHIBIA LINNAEUS, 1758
NEOTETRAPODA
SUBCLASS: REPTILIOMORPHA SAVE-SODERBERGH, 1934
ORDER: SEYMOURIAMORPHA WATSON, 1917
SUBORDER: NYCTEROLETERIDAE TATARINOV, 1972
Family: NYCTEROLETERIDAE Romer, 1956
Subfamily: NYCTEROLECTERINAE Romer, 1956
Macroleter poezicus
Subfamily: BASHKYROLETERINAE Ivachnenko, 1997
Bashkyroleter mesensis
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: PARAREPTILIA OLSON 1947
INFRACLASS: SAUROPSIDA HUXLEY, 1864
ORDER: PROCOLOPHONOMORPHA ROMER, 1964
SUBORDER: PROCOLPHOMORPHA SEELEY, 1888
Family: NYCTIPHRURETIDAE Efremov, 1938
Nyctiphruretus acudens
SUBCLASS: DIAPSIDA OSBORN, 1904
CLADE NAME: NEOVARANOPSIA SPINDLER, WERNEBURG, SCHNEIDER, LUTHARDT, ANNACKER & RÖßLER, 2019
Family: VARANOPSEIDAE Romer & Price, 1940
Subfamily: MESENOSAURINAE Romer 1956
Mesenosaurus romeri
SUBCLASS: SYNAPSIDA OSBORN, 1903
INFRACLASS DINOMORPHA IVAKNENKO, 2008
SUPERORDER: GORGONTIA IVAKNENKO, 2008
ORDER: EOTHERIODONTIA OLSON, 1962
INFRAORDER: EOTITANOSUCHIA BOONSTRA, 1963
Family: NIIAFTASUCHIDAE Ivakhenko, 1990
Niaftasuchus zekkeli
_____________________________________________________________________________________
EUROPEAN RUSSIA
ARKHANGELSK REGION
MEZENSKII DISTRICT
KOSMOGORODSKO'E LOCALITY
BIARMIAN SERIES
URZHUMIAN STAGE
URZHUMIAN HORIZON
MEZEN FAUNAL ASSEMBLAGE
LOWER TATARIAN AGE
CAPITANIAN STAGE
MIDDLE ZECHSTEIN EPOCH
LATE MIDDLE PERMIAN
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: AMPHIBIA LINNAEUS, 1758
NEOTETRAPODA
SUBCLASS: REPTILIOMORPHA SAVE-SODERBERGH, 1934
ORDER: SEYMOURIAMORPHA WATSON, 1917
SUBORDER: NYCTEROLETERIDAE TATARINOV, 1972
Family: NYCTEROLETERIDAE Romer, 1956
Subfamily: NYCTEROLECTERINAE Romer, 1956
Macroleter poezicus
Subfamily: BASHKYROLETERINAE Ivachnenko, 1997
Bashkyroleter mesensis
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: PARAREPTILIA OLSON 1947
INFRACLASS: SAUROPSIDA HUXLEY, 1864
ORDER: PROCOLOPHONOMORPHA ROMER, 1964
SUBORDER: PROCOLPHOMORPHA SEELEY, 1888
Family: NYCTIPHRURETIDAE Efremov, 1938
Nyctiphruretus acudens
SUBCLASS: DIAPSIDA OSBORN, 1904
CLADE NAME: NEOVARANOPSIA SPINDLER, WERNEBURG, SCHNEIDER, LUTHARDT, ANNACKER & RÖßLER, 2019
Family: VARANOPSEIDAE Romer & Price, 1940
Subfamily: MESENOSAURINAE Romer 1956
Mesenosaurus romeri
SUBCLASS: SYNAPSIDA OSBORN, 1903
SUPERORDER: THERASPIDA BROOM, 1905
ORDER: EOTHERIODONTIA OLSON, 1962
INFRAORDER: EOTITANOSUCHIA BOONSTRA, 1963
Family: ALRAUSUCHIDAE Ivakhenko, 2008
Alrausuchus tagax
_____________________________________________________________________________________
EUROPEAN RUSSIA
ARKHANGELSK REGION
MEZENSKII DISTRICT
UST’-PEZA LOCALITY
BIARMIN SERIES
URZHUMINA STAGE
URZHUMIAN HORIZON
UPPER KIMZHA BEDS
MEZEN FAUNAL ASSEMBLAGE
LOWER TATARIAN AGE
CAPITANIAN STAGE
MIDDLE ZECHSTEIN EPOCH
LATE MIDDLE PERMIAN
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: AMPHIBIA LINNAEUS, 1758
NEOTETRAPODA
SUBCLASS: REPTILIOMORPHA SAVE-SODERBERGH, 1934
ORDER: SEYMOURIAMORPHA WATSON, 1917
SUBORDER: NYCTEROLETERIDAE TATARINOV, 1972
Family: NYCTEROLETERIDAE Romer, 1956
Subfamily: NYCTEROLECTERINAE Romer, 1956
*Macroleter poezicus
Subfamily: BASHKYROLETERINAE Ivachnenko, 1997
Bashkyroleter mesensis
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: PARAREPTILIA OLSON 1947
INFRACLASS: SAUROPSIDA HUXLEY, 1864
ORDER: PROCOLOPHONOMORPHA ROMER, 1964
SUBORDER: PROCOLPHOMORPHA SEELEY, 1888
Family: NYCTIPHRURETIDAE Efremov, 1938
*Nyctiphruretus acudens
SUBCLASS: DIAPSIDA OSBORN, 1904
CLADE NAME: NEOVARANOPSIA SPINDLER, WERNEBURG, SCHNEIDER, LUTHARDT, ANNACKER & RÖßLER, 2019
Family: VARANOPSEIDAE Romer & Price, 1940
Subfamily: MESENOSAURINAE Romer 1956
Mesenosaurus romeri
SUBCLASS: SYNAPSIDA OSBORN, 1903
SUPERORDER: THERASPIDA BROOM, 1905
ORDER: EOTHERIODONTIA OLSON, 1962
INFRAORDER: EOTITANOSUCHIA BOONSTRA, 1963
Family: ALRAUSUCHIDAE Ivakhenko, 2008
Alrausuchus tagax
= Biarmosuchus tagax
_____________________________________________________________________________________
EUROPEAN RUSSIA
ARKHANGELSK REGION
MEZENSKII DISTRICT
BELOKUR’E LOCALITY
KRASNOSHCHEL’YA FORMATION
URZHUMIAN STAGE
URZHUMIAN HORIZON
MEZEN FAUNAL ASSEMBLAGE
LOWER TATARIAN AGE
CAPITANIAN STAGE
MIDDLE ZECHSTEIN EPOCH
LATE MIDDLE PERMIAN
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: AMPHIBIA LINNAEUS, 1758
NEOTETRAPODA
SUBCLASS: REPTILIOMORPHA SAVE-SODERBERGH, 1934
ORDER: SEYMOURIAMORPHA WATSON, 1917
SUBORDER: NYCTEROLETERIDAE TATARINOV, 1972
Family: NYCTEROLETERIDAE Romer, 1956
Subfamily: NYCTEROLECTERINAE Romer, 1956
Macroleter poezicus
Subfamily: BASHKYROLETERINAE Ivachnenko, 1997
Bashkyroleter mesensis
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: PARAREPTILIA OLSON 1947
INFRACLASS: SAUROPSIDA HUXLEY, 1864
ORDER: PROCOLOPHONOMORPHA ROMER, 1964SUBORDER: PROCOLPHOMORPHA SEELEY, 1888
Family: NYCTIPHRURETIDAE Efremov, 1938
Nyctiphruretus acudens
SUBCLASS: DIAPSIDA OSBORN, 1904
CLADE NAME: NEOVARANOPSIA SPINDLER, WERNEBURG, SCHNEIDER, LUTHARDT, ANNACKER & RÖßLER, 2019
Family: VARANOPSEIDAE Romer & Price, 1940
Subfamily: MESENOSAURINAE Romer 1956
Mesenosaurus romeri
SUBCLASS: SYNAPSIDA OSBORN, 1903
SUPERORDER: THERASPIDA BROOM, 1905
ORDER: EOTHERIODONTIA OLSON, 1962
Incertae sedis
Family: NIKKASAURIDAE Ivakhnenko, 2000
Nikkasaurus tatarinovi
_____________________________________________________________________________________
EUROPEAN RUSSIA
ARKHANGELSK REGION
MENZEN DISTRICT
UST-NYAFTA
BIARMIAN SERIES
UPPER KIMZHA BEDS
URZHUMIAN STAGE
UPPER URZHUMIAN HORIZON
MEZEN FAUNAL ASSEMBLAGE
LOWER TATARIAN AGE
CAPITANIAN STAGE
UPPER GUADELUPIAN STAGE
MIDDLE ZECHSTEIN EPOCH
LATE MIDDLE PERMIAN
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: AMPHIBIA LINNAEUS, 1758
NEOTETRAPODA
SUBCLASS: REPTILIOMORPHA SAVE-SODERBERGH, 1934
ORDER: SEYMOURIAMORPHA WATSON, 1917
SUBORDER: NYCTEROLETERIDAE TATARINOV, 1972
Family: NYCTEROLETERIDAE Romer, 1956
Subfamily: NYCTEROLECTERINAE Romer, 1956
Macroleter poezicus
Subfamily: BASHKYROLETERINAE Ivachnenko, 1997
Bashkyroleter mesensis
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: PARAREPTILIA OLSON 1947
INFRACLASS: SAUROPSIDA HUXLEY, 1864
ORDER: PROCOLOPHONOMORPHA ROMER, 1964
SUBORDER: PROCOLPHOMORPHA SEELEY, 1888
Family: NYCTIPHRURETIDAE Efremov, 1938
Nyctiphruretus acudens
SUBCLASS: DIAPSIDA OSBORN, 1904
CLADE NAME: NEOVARANOPSIA SPINDLER, WERNEBURG, SCHNEIDER, LUTHARDT, ANNACKER & RÖßLER, 2019
Family: VARANOPSEIDAE Romer & Price, 1940
Subfamily: MESENOSAURINAE Romer 1956
Mesenosaurus romeri
SUBCLASS: SYNAPSIDA OSBORN, 1903
ORDER: PELYCOSAURIA COPE, 1878
SUBORDER: EUPELYCOSAURIA KEMP, 1982
INFRAORDER: OPHIACHODONTIA NOVA
Family: VARANOPIDAE emend Romer & Price, 1940, REISZ & DILKES, 2003
*Pyozia mesenensis
INFRACLASS DINOMORPHA IVAKNENKO, 2008
SUPERORDER: GORGONTIA IVAKNENKO, 2008
ORDER: EOTHERIODONTIA OLSON, 1962
INFRAORDER: EOTITANOSUCHIA BOONSTRA, 1963
Family: NIIAFTASUCHIDAE Ivakhenko, 1990
*Niaftasuchus zekkeli
_____________________________________________________________________________________
EUROPEAN RUSSIA
ARKHANGELSK REGION
LESHUKONSKII DISTRICT
ON THE MEZEN RIVER
NEAR THE JUCTION WITH THE VASHKA RIVER
NYISAGORA LOCALITY
BIARMIAN SERIES
URZHUMIAN STAGE
URZHUMIAN HORIZON
ZONE II
MEZEN FAUNAL ASSEMBLAGE
EARLIEST EARLY TATARIAN AGE
GUADELUPIAN STAGE
MIDDLE ZEICHSTEIN EPOCH
MIDDLE PERMIAN
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: AMPHIBIA LINNAEUS, 1758
NEOTETRAPODA
SUBCLASS: REPTILIOMORPHA SAVE-SODERBERGH, 1934
ORDER: SEYMOURIAMORPHA WATSON, 1917
SUBORDER: NYCTEROLETERIDAE TATARINOV, 1972
Family: NYCTEROLETERIDAE Romer, 1956
Subfamily: NYCTEROLECTERINAE Romer, 1956
Macroleter poezicus
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: PARAREPTILIA OLSON 1947
INFRACLASS: SAUROPSIDA HUXLEY, 1864
ORDER: LANTHANOSUCHIDAE TATARINOV, 1971
Family: LANTHANOSUCHIDAE Efremov, 1946
Lanthaniscus efremovi
ORDER: PROCOLOPHONOMORPHA ROMER, 1964
SUBORDER: PROCOLPHOMORPHA SEELEY, 1888
Family: NYCTIPHRURETIDAE Efremov, 1938
Nyctiphruretus acudens
SUBCLASS: DIAPSIDA OSBORN, 1904
CLADE NAME: NEOVARANOPSIA SPINDLER, WERNEBURG, SCHNEIDER, LUTHARDT, ANNACKER & RÖßLER, 2019
Family: VARANOPSEIDAE Romer & Price, 1940
Subfamily: MESENOSAURINAE Romer 1956
Mesenosaurus romeri
SUBCLASS: SYNAPSIDA OSBORN, 1903
ORDER: PELYCOSAURIA COPE, 1878
SUBORDER: CASEASAURIA WILLISTON, 1912
Family: CASEIDAE Williston, 1912
Ennatosaurus tecton
INFRACLASS DINOMORPHA IVAKNENKO, 2008
SUPERORDER: GORGONTIA IVAKNENKO, 2008
ORDER: DEINOCEPHALIA OLSEN, 1962
SUBORDER: EUDEINOCEPHALIA APPLEBY, CHARIG, COX, KERMACK & TARLO, 1967
INFRAORDER: EOTITANOSUCHIA BOONSTRA, 1963
Family: NIIAFTASUCHIDAE Ivakhenko, 1990
Niaftasuchus zekkeli
_____________________________________________________________________________________
EUROPEAN RUSSIA
ARKHANGELSK REGION
LESHUKONSKII DISTRICT
UST-VASHKA LOCALITY
KRASNOSHCHEL’YA FORMATION
BIARMIAN SERIES
URZHUMIAN STAGE
URZHUMIAN HORIZON
MEZEN FAUNAL ASSEMBLAGE
LOWER TATARIAN AGE
CAPITANIAN STAGE
MIDDLE ZECHSTEIN EPOCH
LATE MIDDLE PERMIAN
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: AMPHIBIA LINNAEUS, 1758
NEOTETRAPODA
SUBCLASS: REPTILIOMORPHA SAVE-SODERBERGH, 1934
ORDER: SEYMOURIAMORPHA WATSON, 1917
SUBORDER: NYCTEROLETERIDAE TATARINOV, 1972
Family: NYCTEROLETERIDAE Romer, 1956
Subfamily: BASHKYROLETERINAE Ivachnenko, 1997
Bashkyroleter mesensis
Subfamily: NYCTEROLECTERINAE Romer, 1956
Macroleter poezicus
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: PARAREPTILIA OLSON 1947
INFRACLASS: SAUROPSIDA HUXLEY, 1864
ORDER: PROCOLOPHONOMORPHA ROMER, 1964
SUBORDER: PROCOLPHOMORPHA SEELEY, 1888
Family: NYCTIPHRURETIDAE Efremov, 1938
Nyctiphruretus acudens
SUBCLASS: DIAPSIDA OSBORN, 1904
CLADE NAME: NEOVARANOPSIA SPINDLER, WERNEBURG, SCHNEIDER, LUTHARDT, ANNACKER & RÖßLER, 2019
Family: VARANOPSEIDAE Romer & Price, 1940
Subfamily: MESENOSAURINAE Romer 1956
Mesenosaurus romeri
SUBCLASS: SYNAPSIDA OSBORN, 1903
INFRACLASS DINOMORPHA IVAKNENKO, 2008
SUPERORDER: NIKKASAURIA IVAKHNENKO, 2008
ORDER: NIKKASAURIDA IVAKHNENNKO, 2008
Family: NIKKASAURIDAE Ivakhnenko, 2000
Nikkasaurus tatarinovi
*Reiszia tippula
SUPERORDER: GORGONTIA IVAKNENKO, 2008
ORDER: DEINOCEPHALIA OLSEN, 1962
SUBORDER: EUDEINOCEPHALIA APPLEBY, CHARIG, COX, KERMACK & TARLO, 1967
INFRAORDER: EOTITANOSUCHIA BOONSTRA, 1963
Family: NIIAFTASUCHIDAE Ivakhenko, 1990
Niaftasuchus zekkeli
Family: ALRAUSUCHIDAE Ivakhenko, 2008
Alrausuchus tagax
_____________________________________________________________________________________
EUROPEAN RUSSIA
ARKHANGELSK REGION
PEZA-1
BIARMIAN SERIES
UPPER KIMZHA BEDS
URZHUMIAN STAGE
URZHUMIAN HORIZON
MEZEN FAUNAL ASSEMBLAGE
LOWER TATARIAN AGE
LOPINGIAN SUBEPOCH
UPPER ZECHSTEIN EPOCH
LATE PERMIAN
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: AMPHIBIA LINNAEUS, 1758
NEOTETRAPODA
SUBCLASS: REPTILIOMORPHA SAVE-SODERBERGH, 1934
ORDER: SEYMOURIAMORPHA WATSON, 1917
SUBORDER: NYCTEROLETERIDAE TATARINOV, 1972
Family: NYCTEROLETERIDAE Romer, 1956
Subfamily: NYCTEROLECTERINAE Romer, 1956
Macroleter poezicus
Nycteroleter ineptus
Subfamily: BASHKYROLETERINAE Ivachnenko, 1997
Bashkyroleter mesensis
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: PARAREPTILIA OLSON 1947
INFRACLASS: SAUROPSIDA HUXLEY, 1864
ORDER: LANTHANOSUCHIDAE TATARINOV, 1971
Family: LANTHANOSUCHIDAE Efremov, 1946
Lanthaniscus efremovi
ORDER: PROCOLOPHONOMORPHA ROMER, 1964
SUBORDER: PROCOLPHOMORPHA SEELEY, 1888
Family: NYCTIPHRURETIDAE Efremov, 1938
Nyctiphruretus acudens
SUBCLASS: DIAPSIDA OSBORN, 1904
CLADE NAME: NEOVARANOPSIA SPINDLER, WERNEBURG, SCHNEIDER, LUTHARDT, ANNACKER & RÖßLER, 2019
Family: VARANOPSEIDAE Romer & Price, 1940
Subfamily: MESENOSAURINAE Romer 1956
Mesenosaurus romeri
SUBCLASS: SYNAPSIDA OSBORN, 1903
SUPERORDER: THERASPIDA BROOM, 1905
ORDER: EOTHERIODONTIA OLSON, 1962
INFRAORDER: EOTITANOSUCHIA BOONSTRA, 1963
Family: NIIAFTASUCHIDAE Ivakhenko, 1990
Niaftasuchus zekkeli
Family: ALRAUSUCHIDAE Ivakhenko, 2008
*Alrausuchus tagax
= Biarmosuchus tagax
_____________________________________________________________________________________
ARKHANGELSK REGION
MEZENSKII DISTRICT
UST’-PEZA LOCALITY
UPPER KAZANIAN SUBSTAGE OR
LOWER TATARIAN AGE
LOPINGIAN SUBEPOCH
UPPER ZECHSTEIN EPOCH
LATE PERMIAN
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: SYNAPSIDA OSBORN, 1903
SUPERORDER: THERASPIDA BROOM, 1905
ORDER: EOTHERIODONTIA OLSON, 1962
SUBORDER: EOTITANOSUCHIA BOONSTRA, 1963
INFRAORDER: EOTITANOSUCHIA BOONSTRA, 1963
Family: EOTITANOSUCHIDAE CHUDINOV, 1960
Biarmosuchus tagax
_____________________________________________________________________________________
EUROPEAN RUSSIA
ARKHANGELSK REGION
MEZENSKII DISTRICT
UST’-VASHKA LOCALITY
UPPER KAZANIAN SUBSTAGE OR
LOWER TATARIAN AGE
LOPINGIAN SUBEPOCH
UPPER ZECHSTEIN EPOCH
LATE PERMIAN
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: SYNAPSIDA OSBORN, 1903
SUPERORDER: THERASPIDA BROOM, 1905
ORDER: EOTHERIODONTIA OLSON, 1962
SUBORDER: EOTITANOSUCHIA BOONSTRA, 1963
INFRAORDER: EOTITANOSUCHIA BOONSTRA, 1963
Family: EOTITANOSUCHIDAE CHUDINOV, 1960
Biarmosuchus tagax
_____________________________________________________________________________________
EUROPEAN RUSSIA
ARKHANGEL’SK REGION
LOWER MEZEN RIVER
ZONE III
LOWER TATARIAN AGE
LOPINGIAN SUBEPOCH
UPPER ZECHSTEIN EPOCH
LATE PERMIAN
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: AMPHIBIA LINNAEUS, 1758
NEOTETRAPODA
SUBCLASS: REPTILIOMORPHA SAVE-SODERBERGH, 1934
ORDER: SEYMOURIAMORPHA WATSON, 1917
SUBORDER: NYCTEROLETERIDAE TATARINOV, 1972
Family: NYCTEROLETERIDAE Romer, 1956
Subfamily: NYCTEROLECTERINAE Romer, 1956
*Nycteroleter ineptus
_____________________________________________________________________________________