Species: dilatus GILMORE, 1928
Etymology: Latin, dilatus, "expanded, spread."

Holotype: USNM 10749

Locality: Niobrara County, Wyoming.

Horizon: Lance Formation.

Biostratigraphy:

Age: Lancian age, upper Maastrichtian Stage, Upper Senonian subepoch, Upper Upper Gulf epoch, Late Cretaceous.

Material: Dentary.

Referred material:

USNM 10813, 10815-10816, 17018: Dentaries.

 

EBERTH, 1987

Locality: Dinosaur Provincial Park, Newell County, Alberta Province, Canada.

Horizon: Dinosaur Park Formation, upper Judith River Group.

Biostratigraphy:

Age: Judithian age, Middle Campanian Stage, middle Senonian subepoch, upper Gulf Epoch, Late Cretaceous.

Material:

Number: Not given:

 

BRAYNT, 1990

Locality: Northeastern Montana.

Horizon: Hell Creek Formation.

Biostratigraphy:

Age: Lancian age, upper Maastrichtian Stage, Upper Senonian subepoch, Upper Upper Gulf epoch, Late Cretaceous.

Material:

UCMP 130691: Dentary fragments.

UCMP 123294: Tooth plate.

UCMP 130750: Tooth plate.

UCMP 130693: Vertebra.

 

BREITHUPT, 1982

Locality: East flank of Rock Springs Uplift, Sweetwater County, Wyoming.

Horizon: Lance Formation.

Biostratigraphy:

Age: Lancian age, upper Maastrichtian Stage, Upper Senonian subepoch, Upper Upper Gulf epoch, Late Cretaceous.

Material:

UW 14268: Right dentary fragment.

UW 15034: Right dentary fragment.

UW 15035: ?left dentary fragment.

UW 15036: Maxillary fragment.

UW 15037: Vomer fragments.

UW 15038: Premaxillary fragment.

UW 15039: Pterygoplatine.

 

BREITHAUPT, 1985

Locality: Wyoming.

Horizon: "Mesaverde" Formation.

Biostratigraphy:

Age: Judithian age, Campanian, Senonian subepoch, Upper Gulf epoch, Late Cretaceous.

Material:

Number: Not given:

ESTES, 1964

Locality: V-5003, S 1/2, Sec. 20, T37N, R64W, Lance Creek, Powder River Basin, Niobara County, Wyoming.

Horizon: Lance Formation.

Biostratigraphy:

Age: Lancian age, upper Maastrichtian Stage, Upper Senonian subepoch, Upper Upper Gulf epoch, Late Cretaceous.

Material:

Number: Not given: 2 fused pathological vertebra.

 

Localities: UCMP V-5616, Altman Blowout, this large blowout, site of the UCMP field-party camps form 1956 to 1960, is just east of the crest of the divide separating Lance Creek and Hancock Creek and 100 yards east of a dirt road. Fossils were found in anthills, talus, and sandstone exposed in the blowout, which is in the NE 1/4, Sec. 9, T37N, R64W, Niobrara County (Formerly Converse County), Wyoming.

Horizon: Lance Formation.

Biostratigraphy:

Age: Lancian age, upper Maastrichtian Stage, Upper Senonian subepoch, Upper Upper Gulf epoch, Late Cretaceous.

Material:

Number: Not given: A few vertebrae, one first ceratobranchia-.

Locality: UC V-5618, Eastern Wyoming.

Horizon: Lance Formation.

Biostratigraphy:

Age: Lancian age, upper Maastrichtian Stage, Upper Senonian subepoch, Upper Upper Gulf epoch, Late Cretaceous.

Material:

Number: Not given: Vertebra.

 

Locality: UC V-5619, Eastern Wyoming.

Horizon: Lance Formation.

Biostratigraphy:

Age: Lancian age, upper Maastrichtian Stage, Upper Senonian subepoch, Upper Upper Gulf epoch, Late Cretaceous.

Material:

Number: Not given: A few jaw fragments and a first ceratobranchial.

 

Locality: UCMP V-5620, Lull Quarry, near the head of a tributary of Buck Creek, Powder River Basin, Niobrara County (Formerly Converse County), Wyoming.

Horizon: Lance Formation.

Biostratigraphy:

Age: Lancian age, upper Maastrichtian Stage, Upper Senonian subepoch, Upper Upper Gulf epoch, Late Cretaceous.

Material:

UC 46046, 46054, 46062, 46065, 46077, 46097, 46104, 46197, 46613, 49491-49492, 49495--49497, 46460: Dentaries.

UC 46043, 49493-49494, 49537, 55714-55717: Maxillae.

UC 46073, 46083, 46612, 4949-49500: Premaxillae.

UC 46013, 49498: Vomers.

UC 49505-49506: First ceratobranchials.

UC 49518-49519: Atlases.

UC 46627: Dorsal vertebrae.

Many unnumbered palatal tooth plates and specimens of the above bones are present.

 

Locality: UCMP V-5621, Cow Creek, site 1, anthills and talus found on and around outcrops of cross-stratified sandstone on the north side and near the upper end of a gully emptying into a northern tributary of Cow Creek, Niobrara County (Formerly Converse County), Wyoming.

Horizon: Lance Formation.

Biostratigraphy:

Age: Lancian age, upper Maastrichtian Stage, Upper Senonian subepoch, Upper Upper Gulf epoch, Late Cretaceous.

Material:

Number: Not given: Vertebra.

 

Locality: V-5623, Curiosity Point, anthills and talus found along the southern side of a twin-leaked erosional remnant of cross-stratified sandstone approximately 200 years north of a dirt road,
Wyoming.

Horizon: Lance Formation.

Biostratigraphy:

Age: Lancian age, upper Maastrichtian Stage, Upper Senonian subepoch, Upper Upper Gulf epoch, Late Cretaceous.

Material:

Number: Not given: A fragment of palatal tooth plate.

 

Locality: UCMP V-5711, Bushy Tailed Blowout, Quarry area includes the floor of a small blowout approximately 10 yards northwest of the highest point on the ridge forming the northeastern side of the blowouts of loc. V-5003, S 1/2, Sec. 20, T37N, R64W, Lance Creek, Powder River Basin, Niobrara County, Wyoming.

Horizon: Lance Formation.

Biostratigraphy:

Age: Lancian age, upper Maastrichtian Stage, Upper Senonian subepoch, Upper Upper Gulf epoch, Late Cretaceous.

Material:

Number: Not given: Many unnumbered dentaries, palatal tooth plates, atlases, vertebrae, and ceratobranchials.

GARDNER, 2003

UALVP 43910-43913: Premaxillae.

UALVP 43587: Maxilla.

UALVP 1614, 43873-43874, 43915-43918: Dentaries.

UALVP 43914: Vomer.

UALVP 43919: Atlas.

UALVP 43895-43899: Trunk vertebrae.

UALVP 43901: Caudal vertebra.

 

Locality: UC V-5713, Eastern Wyoming.

Horizon: Lance Formation.

Biostratigraphy:

Age: Lancian age, upper Maastrichtian Stage, Upper Senonian subepoch, Upper Upper Gulf epoch, Late Cretaceous.

Material:

Number: Not given: Dentary.

 

Locality: UCMP V-5815, Hatcher Number 2, western side of a cross-stratified sandstone erosional remnant near the center of a large blowout on the crest of a ridge, the distance form the nearest point on the road, approximately 150 yards south of the quarry, to the Johnson ranch is 0.4 miles, Niobrara County (Formerly Converse County), Eastern Wyoming.

Horizon: Lance Formation.

Biostratigraphy:

Age: Lancian age, upper Maastrichtian Stage, Upper Senonian subepoch, Upper Upper Gulf epoch, Late Cretaceous.

Material:

Number: Not given: A few vertebrae, Atlases, palatal tooth plates and jaw elements.

 

Locality: UCMP V-5817, False Disciple Point, cross-stratified sandstone and sandy shales cropping out on the top of the northern end of a long, northwestward-trending ridge that extends from the crest of the divide between the Hancock and Lance Creek drainages, Niobrara County (Formerly Converse County), Wyoming.

Horizon: Lance Formation.

Biostratigraphy:

Age: Lancian age, upper Maastrichtian Stage, Upper Senonian subepoch, Upper Upper Gulf epoch, Late Cretaceous.

Material:

Number: Not given: A few dentaries, palatal tooth plates and vertebrae.

 

Locality: UCMP V-5820, Kent 484, found on the surface near the top of the east side of a small draw on the south side of the prominent hill northeast of the J. Krejci ranch house, Niobrara County (Formerly Converse County), Wyoming.

Horizon: Lance Formation.

Biostratigraphy:

Age: Lancian age, upper Maastrichtian Stage, Upper Senonian subepoch, Upper Upper Gulf epoch, Late Cretaceous.

Material:

Number: Not given: Dentary.

 

ESTES, BERBERIAN & MESZOELY, 1969

Locality: Bug Creek Anthills (BCA), 80 feet below the Tullock Formation in the west half of Section 9, T22N, R43E, McCone County, Montana.

Horizon: Hell Creek Formation.

Biostratigraphy:

Age: Lancian age, upper Maastrichtian Stage, Upper Senonian subepoch, Upper Upper Gulf epoch, Late Cretaceous.

Material:

MCZ 3675: Vertebrae, Atlantes, and dentaries.

GARDNER, 2003

UALVP 43849-43854: Premaxillae.

UALVP 43855-43856: Maxillae.

UALVP 43858-43871: Dentaries.

UALVP 43872: Lot of seven fragmentary dentaries.

UALVP 43875-43876: Vomers.

UALVP 43877: Palatine.

UALVP 40046-40048, 43878-43882: Atlantes.

UALVP 40049-40050, 43883-43894: Trunk vertebrae.

UALVP 43900: Caudal vertebra.

 

GARDNER, 2003

Locality: Wounded Knee, Saskatchewan Province, Canada.

Horizon: Frenchman Formation.

Biostratigraphy:

Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.

Material:

UALVP 43921-43926: Premaxillae.

UALVP 43927: Maxilla.

UALVP 43930-43933: Dentaries.

UALVP 43928-43929: Vomers.

UALVP 43934: Atlas.

 

LANGSTON, STANDHARDT, & STEVENS, 1989

Number: Not given.

Locality: Big Bend, Brewster County, Texas.

Horizon: Lower Upper Tornillo.

Biostratigraphy:

Age: Lancian Age, Maastrichtian, Upper Senonian subepoch, Upper Upper Gulf Epoch, Late Cretaceous.

Material:

Number: Not given:

SAHNI, 1972

Locality: Clambank Hollow Locality, Montana.

Horizon: Judith River Formation.

Biostratigraphy:

Age: Judithian age, Campanian Stage, Senonian subepoch, Upper Gulf epoch, Late Cretaceous.

Material:

AMNH 8477: A fragmentary, edentulous, right dentary.

 

STENERSON & O’CONNER, 1994

Locality: Northwestern South Dakota.

Horizon: Hell Creek Formation.

Biostratigraphy:

Age: Lancian age, upper Maastrichtian Stage, Upper Senonian subepoch, Upper Upper Gulf epoch, Late Cretaceous.

Material:

Number: Not given:

 

WHITMORE, 1988

Locality: SDSM Locality V851, Zerbst Locality, east of the Cheyenne River, Niobrara County, Wyoming.

Horizon: Lance Formation.

Biostratigraphy:

Age: Lancian age, upper Maastrichtian Stage, Upper Senonian subepoch, Upper Upper Gulf epoch, Late Cretaceous.

Material:

SDSM 12377: 3 atlantes.

SDSM 12384: 19 vertebrae.

 

Locality: SDSM Locality V852, Hanson Locality, Niobrara County, Wyoming.

Horizon: Lance Formation.

Biostratigraphy:

Age: Lancian age, upper Maastrichtian Stage, Upper Senonian subepoch, Upper Upper Gulf epoch, Late Cretaceous.

Material:

SDSM 12376: 1 atlas.

 

Locality: SDSM Locality V685, Greasewood Creek Locality, northern side of the south fork of the Greasewood Creek, Niobrara County, Wyoming.

Horizon: Lance Formation.

Biostratigraphy:

Age: Lancian age, upper Maastrichtian Stage, Upper Senonian subepoch, Upper Upper Gulf epoch, Late Cretaceous.

Material:

SDSM 12697: 1 vertebra.

 

Locality: SDSM Locality V851B, Zerbst Hadrosaur Level, Niobrara County, Wyoming.

Horizon: Lance Formation.

Biostratigraphy:

Age: Lancian age, upper Maastrichtian Stage, Upper Senonian subepoch, Upper Upper Gulf epoch, Late Cretaceous.

Material:

SDSM 12387: 1 vertebra.

 

WROBLEWSKI,1998

Locality: Western Hanna Basin, Carbon County, Southcental Wyoming.

Horizon: Ferris Formation.

Biostratigraphy:

Age: Lancian age, uppermost Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.

Material:

Number: Not given:

 

= Adelphesiren olivae GOIN & AUFFENBERG 1958
Etymology:

Holotype: CM 6467

Locality: Niobrara County, Wyoming.

Horizon: Lance Formation.

Biostratigraphy:

Age: Lancian age, upper Maastrichtian Stage, Upper Senonian subepoch, Upper Upper Gulf epoch, Late Cretaceous.

Material: Dorsal vertebra.

Referred material:

CM 6448: Dorsal vertebra.

_____________________________________________________________________________________

Species: prodilatus GARDNER, 2003
Etymology: Latin, pro, "before" and Latin, dilatus, "spread or expanded"; specific epithet of the type species in reference to the earlier age and possible ancestral status of the new congener.

Holotype: UALVP 43906

Locality: Irvine microvertebrate locality, Cypress County, Alberta Province, Canada.

Horizon: Dinosaur Park Formation, upper Judith River Group.

Biostratigraphy:

Age: Judithian age, Middle Campanian Stage, middle Senonian subepoch, upper Gulf Epoch, Late Cretaceous.

Material: Right premaxilla missing distal end of vomerine process and most of pars dorsalis and preserving an intact tooth row with three complete teeth, 1 nearly complete tooth and bases of 3 others.

Referred material:

UALVP 43902-43905: Premaxillae.

UALVP 43907-43909: Dentaries.

UALVP 40051: Atlantal centrum.