Species: kayi AUFFENBERG, 1964
Etymology:

Holotype: CM 6468

Locality: Niobrara County, Wyoming.

Horizon: Lance Formation.

Biostratigraphy:

Age: Lancian age, upper Maastrichtian Stage, Upper Senonian subepoch, Upper Upper Gulf epoch, Late Cretaceous.

Material: Middle Trunk vertebra.

Referred material:

ARMSTRONG-ZIEGLER, 1978

Locality: UA-P 75137, SE 1/4, SE 1/4, sec. 34, T24N, R13W, Alamo Mesa West, 7 1/2 quadrangle, San Juan Basin, New Mexico.

Horizon: Fruitland Formation.

Biostratigraphy:

Age: Judithian age, Late Campanian Stage, Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.

Material:

Number: Not given:

 

EBERTH, 1987

Locality: Dinosaur Provincial Park, Newell County, Alberta Province, Canada.

Horizon: Dinosaur Park Formation, upper Judith River Group.

Biostratigraphy:

Age: Judithian age, Middle Campanian Stage, middle Senonian subepoch, upper Gulf Epoch, Late Cretaceous.

Material:

Number: Not given:

BRAYNT, 1990

Locality: Northeastern Montana.

Horizon: Hell Creek Formation.

Biostratigraphy:

Age: Lancian age, upper Maastrichtian Stage, Upper Senonian subepoch, Upper Upper Gulf epoch, Late Cretaceous.

Material:

UCMP 130684: Vertebrae.

UCMP 123488: Vertebrae.

 

BREITHUPT, 1982

Locality: East flank of Rock Springs Uplift, Sweetwater County, Wyoming.

Horizon: Lance Formation.

Biostratigraphy:

Age: Lancian age, upper Maastrichtian Stage, Upper Senonian subepoch, Upper Upper Gulf epoch, Late Cretaceous.

Material:

UW 14573: Right dentary fragment.

UW 14574: Right dentary fragment.

UW 14575-14577: left dentary fragments.

UW 15028: left dentary fragment.

UW 15029: left dentary fragment.

UW 14578: left vomer fragment.

UW 14580: Pterygopalatine fragment.

UW 15026: Pterygoplatine fragment.

UW 14581: Vertebral fragment.

UW 15027: Miscellaneous vomer.

 

BREITHAUPT, 1985

Locality: Wyoming.

Horizon: "Mesaverde" Formation.

Biostratigraphy:

Age: Judithian age, Campanian, Senonian subepoch, Upper Gulf epoch, Late Cretaceous.

Material:

Number: Not given:

BRINKMAN, 1990

Locality: Dinosaur Provincial Park, Newell County, Alberta Province, Canada.

Horizon: Dinosaur Park Formation, upper Judith River Group.

Biostratigraphy:

Age: Judithian age, Middle Campanian Stage, middle Senonian subepoch, upper Gulf Epoch, Late Cretaceous.

Material:

Number: Not given:

 

CARPENTER, 1979

Locality: UCM loc. 77062, Weld County, Colorado.

Horizon: Laramie Formation.

Biostratigraphy:

Age: Lancian age, upper Maastrichtian Stage, Upper Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.

Material:

UCM 38765: Dentary.

UCM 38766: Dentary.

UCM 38714: Atlas.

UCM 38718: Atlas.

UCM 38719: Atlas.

 

ESTES, 1964

Locality: V-5003, S 1/2, Sec. 20, T37N, R64W, Lance Creek, Powder River Basin, Niobara County, Wyoming.

Horizon: Lance Formation.

Biostratigraphy:

Age: Lancian age, upper Maastrichtian Stage, Upper Senonian subepoch, Upper Upper Gulf epoch, Late Cretaceous.

Material:

UC 55821: A-most complete second cervical vertebra.

Other unnumbered vertebrae also present.

 

Localities: UCMP V-5616, Altman Blowout, this large blowout, site of the UCMP field-party camps form 1956 to 1960, is just east of the crest of the divide separating Lance Creek and Hancock Creek and 100 yards east of a dirt road. Fossils were found in anthills, talus, and sandstone exposed in the blowout, which is in the NE 1/4, Sec. 9, T37N, R64W, Niobrara County (Formerly Converse County), Wyoming.

Horizon: Lance Formation.

Biostratigraphy:

Age: Lancian age, upper Maastrichtian Stage, Upper Senonian subepoch, Upper Upper Gulf epoch, Late Cretaceous.

Material:

Unnumbered: Vertebrae.

 

Locality: UC V-5618, Niobara County (Formerly Converse County), Eastern Wyoming.

Horizon: Lance Formation.

Biostratigraphy:

Age: Lancian age, upper Maastrichtian Stage, Upper Senonian subepoch, Upper Upper Gulf epoch, Late Cretaceous.

Material: Vertebrae.

 

Locality: UCMP V-5620, Lull Quarry, near the head of a tributary of Buck Creek, Powder River Basin, Niobrara County (Formerly Converse County), Wyoming.

Horizon: Lance Formation.

Biostratigraphy:

Age: Lancian age, upper Maastrichtian Stage, Upper Senonian subepoch, Upper Upper Gulf epoch, Late Cretaceous.

Material:

UC 49501-49504, 46078, 49526: Dentaries.

UC 49528-49532: Vomers.

UC 47817, 46063, 50021: Maxillae.

UC 49520-49521: Atlases.

UC 49512-49514: Vertebrae.

Many other unnumbered specimens of above bones.

 

Locality: UC V-5624, Niobara County (Formerly Converse County), Eastern Wyoming.

Horizon: Lance Formation.

Biostratigraphy:

Age: Lancian age, upper Maastrichtian Stage, Upper Senonian subepoch, Upper Upper Gulf epoch, Late Cretaceous.

Material:

Number: Not given: Atlas.

 

Locality: UCMP V-5711, Bushy Tailed Blowout, Quarry area includes the floor of a small blowout approximately 10 yards northwest of the highest point on the ridge forming the northeastern side of the blowouts of loc. V-5003, S 1/2, Sec. 20, T37N, R64W, Lance Creek, Powder River Basin, Niobrara County, Wyoming.

Horizon: Lance Formation.

Biostratigraphy:

Age: Lancian age, upper Maastrichtian Stage, Upper Senonian subepoch, Upper Upper Gulf epoch, Late Cretaceous.

Material:

UC 49544: Premaxilla.

Many unnumbered vertebrae, atlases, dentaries, vomers and premaxillae.

 

Locality: UCMP V-5815, Hatcher Number 2, western side of a cross-stratified sandstone erosional remnant near the center of a large blowout on the crest of a ridge, the distance form the nearest point on the road, approximately 150 yards south of the quarry, to the Johnson ranch is 0.4 miles, Niobrara County (Formerly Converse County), Eastern Wyoming.

Horizon: Lance Formation.

Biostratigraphy:

Age: Lancian age, upper Maastrichtian Stage, Upper Senonian subepoch, Upper Upper Gulf epoch, Late Cretaceous.

Material:

Number: Not given: Vertebrae.

 

Locality: UCMP V-5817, False Disciple Point, cross-stratified sandstone and sandy shales cropping out on the top of the northern end of a long, northwestward-trending ridge that extends from the crest of the divide between the Hancock and Lance Creek drainages, Niobrara County (Formerly Converse County), Wyoming.

Horizon: Lance Formation.

Biostratigraphy:

Age: Lancian age, upper Maastrichtian Stage, Upper Senonian subepoch, Upper Upper Gulf epoch, Late Cretaceous.

Material:

Number: Not given: A few vertebrae and dentaries.

 

ESTES, BERBERIAN & MESZOELY, 1969

Locality: Bug Creek Anthills, McCone County, Montana.

Horizon: Hell Creek Formation.

Biostratigraphy:

Age: Lancian age, upper Maastrichtian Stage, Upper Senonian subepoch, Upper Upper Gulf epoch, Late Cretaceous.

Material:

MCZ 3676: Vertebrae, Atlantes, dentaries, exocciptials, parietals, premaxillae and maxillae.

 

HOGANSON, ERICKSON & HOLLAND, 2007

Locality: NDSFC Locality LO64, Logan County, North Dakota.

Horizon: Fox Hills Formation.
Note: Interfingering of Western Interior Seaway with the Hell Creek Formation.

Biostratigraphy:

Age: Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.

Material:

ND 99-6.2: Dorsal vertebra.

 

LANGSTON, STANDHARDT, & STEVENS, 1989

Locality: Big Bend, Brewster County, Texas.

Horizon: Lower Tornillo.

Biostratigraphy:

Age: Lancian Age, Maastrichtian, Upper Senonian subepoch, Upper Upper Gulf Epoch, Late Cretaceous.

Material:

Number: Not given:

NAYLOR, 1979

Locality: Alberta Province, Canada.

Horizon: Judith River Formation, Judith River Group.

Biostratigraphy:

Age: Judithian age, Campanian Stage, Senonian subepoch, Upper Gulf epoch, Late Cretaceous.

Material:

UA 12096: Atlas.

 

SAHNI, 1972

Locality: Montana.

Horizon: Judith River Formation.

Biostratigraphy:

Age: Judithian age, Campanian Stage, Senonian subepoch, Upper Gulf epoch, Late Cretaceous.

Material:

AMNH 8481: The cotyles of a partial atlas.

AMNH 8112: Dorsal vertebrae.

AMNH 8485: Fragmentary dentary.

 

STENERSON & O’CONNER, 1994

Locality: Northwestern South Dakota.

Horizon: Hell Creek Formation.

Biostratigraphy:

Age: Lancian age, upper Maastrichtian Stage, Upper Senonian subepoch, Upper Upper Gulf epoch, Late Cretaceous.

Material:

Number: Not given:

 

WHITMORE, 1988

Locality: SDSM Locality V851, Zerbst Locality, east of the Cheyenne River, Niobrara County, Wyoming.

Horizon: Lance Formation.

Biostratigraphy:

Age: Lancian age, upper Maastrichtian Stage, Upper Senonian subepoch, Upper Upper Gulf epoch, Late Cretaceous.

Material:

SDSM 12380: 5 atlantes.

SDSM 12386: 41 vertebrae.

 

Locality: SDSM Locality V852, Hanson Locality, Niobrara County, Wyoming.

Horizon: Lance Formation.

Biostratigraphy:

Age: Lancian age, upper Maastrichtian Stage, Upper Senonian subepoch, Upper Upper Gulf epoch, Late Cretaceous.

Material:

SDSM 12385: 8 vertebrae.

 

Locality: SDSM Locality V685, Greasewood Creek Locality, northern side of the south fork of the Greasewood Creek, Niobrara County, Wyoming.

Horizon: Lance Formation.

Biostratigraphy:

Age: Lancian age, upper Maastrichtian Stage, Upper Senonian subepoch, Upper Upper Gulf epoch, Late Cretaceous.

Material:

SDSM 12743: 1 vertebrae.

 

Locality: SDSM Locality V851B, Zerbst Hadrosaur Level, Niobrara County, Wyoming.

Horizon: Lance Formation.

Biostratigraphy:

Age: Lancian age, upper Maastrichtian Stage, Upper Senonian subepoch, Upper Upper Gulf epoch, Late Cretaceous.

Material:

SDSM 15112: 1 vertebral fragment.