TVERDOKHLEBOV, TVERDOKHLEBOVA, MINIKH, SURKOV & BENTON, 2005
Locality: Yaman-Yushatyr, Right bank of the Yaman Yushatyr River, opposite Davletkulovo village, Ural Drainage basin, South Urals, Orenburg Region, European Russia.
Horizon: Belebey Svita.
Biostratigraphy:
Age: Top of the Kazanian substage, Capitanian Stage, Upper Guadelupian Stage, Middle Zechstein Epoch, Late Middle Permian.
Material:
Number: Not given: Isolated postcranial bones.