Genus: Scutosaurus HARTMANN-WEINBERG, 1930
Etymology: Latin, Scutum, “leather shield”, Greek, sauros, “lizard”: Shield lizard.
= Proelginia HARTMANN-WEINBERG, 1937
Etymology: Before Elginia.

Species: karpinskii (AMALITSKI, 1922) HARTMANN-WEINBERG, 1930
Etymology:
= Pareiasaurus karpinskii AMALITSKI, 1922
= Pareiosaurus karpinskii (AMALITSKI, 1922) (sic)
= Pareiosaurus karpinskyi WATSON, 1917) (sic)

Holotype: PIN N 2005/1532

Locality: Sokolki, North Divina River, Kotlasskii (Kotlas) District, Arkhangel’sk Province, North European Russia.

Horizon: Tatarian Series, Upper Vyatkian Substage, Vyatkian Horizon (Northern Dvina Horizon, bottom of the Vyatkian Horizon).

Biostratigraphy: Sokolki Faunal Assemblage, Sokolki Faunal Subassemblage, Scutosaurus karpinskii zone (Dicynodon Assemblage zone, LUCAS, 2005), Zone IV.

Age: Upper Tatarian Age, Lopingian Subepoch, Upper Zechstein Epoch, Late Permian.

Material:

Referred material:

N 200/1538: Skull.

LEE, 1997

PIN 2005/1533, 2005/1535, 2005/1536, 2005/1537, 2005/1538: 6 skeletons including skull, lower jaw, complete vertebral column, ribs, most of shoulder pelvic girdles, and limbs. Osteoderms are only associated with the first specimen.

PIN 2005/1534: Skull and mandible, 17 presacrals, 5 sacrals, 14 caudals, pelvis and parts of both hindlimbs.

PIN 2005/2471: Partial skull including prepared braincase.

PIN 11/215: Skull and lower jaws.

PIN 3919: Braincase, and fragments of skull and lower jaws, osteoderms and numerous unidentifiable fragments. This number also includes a left dentary, left premaxilla, and partial fragments of a second pareiasaur.

PIN 2895: Skull and lower jaw fragments.

PIN 156/2, 156/3: 2 juvenile skulls, missing lower jaws.

PIN 156/305, 156/306: 2 juvenile lower jaws.

PIN 2005/1885: Weathered braincase.

PIN 2005/2473: Atlas-axis complex, sectioned sagitally.

PIN 2005/1877: Partial pes.

PIN 2005/2489: Phalangeal elements with associated sesamoids.

PIN 2005/123: Astragalocalcaneum, sectioned.

PIN 2005/2070: Right fibula.

PIN 2005/1543: Interclavicle.

PIN 2005/455: Partial forelimb.

PIN 11/28: Intermedium and ulnare.

PIN 2005/698: Osteoderm and rib.

 

IVAKHNENKO, 2008

Locality: Zavarzhye (Zavrazh’e) locality, Malaya Severnaya Dvina River, Kotlasskii (Kotlas) District, Arkhangel’sk Province, North European Russia.

Horizon: Tatarian Series, Vyatkian Substage, Vyatkian Horizon, Vyatka Regional Stage.

Biostratigraphy: Sokolki Faunal Assemblage, Sokolki Faunal Substage (Dicynodon Assemblage zone, LUCAS, 2005).

Age: Upper Tatarian Age, Lopingian Subepoch, upper Zechstein Epoch, Late Permian.

Material:

Numbers: Not given:

 

= Pareiasuchus elegans AMALITSKI, 1922
Etymology:
= Pareiosaurus elegans (AMALITSKI, 1922) (sic)

Holotype:

Locality: European Russia.

Horizon:

Biostratigraphy:

Age: Late Permian.

Material:

 

= Pareiasuchus horridus AMALITSKI, 1922
Etymology:
= Pareiasaurus horridus (AMALITSKI, 1922)
= Pareiosaurus horridus (AMALITSKI, 1922) (sic)

Holotype:

Locality: European Russia.

Horizon:

Biostratigraphy:

Age: Late Permian.

Material: Skull and dentary.

 

= Pareiasuchus tuberculatus AMALITSKI, 1922
Etymology:
= Scutosaurus tuberculatus (AMALITSKI, 1922)
= Pareiosaurus tuberculatus (AMALITSKI 1922) (sic)

Holotype: N 2005/1533

Locality: Sokolki, North Divina River, Kotlasskii (Kotlas) District, Arkhangel’sk Province, North European Russia.

Horizon: Tatarian Series, Upper Vyatkian Substage, Vyatkian Horizon (Northern Dvina Horizon, bottom of the Vyatkian Horizon).

Biostratigraphy: Sokolki Faunal Assemblage, Sokolki Faunal Subassemblage, Scutosaurus karpinskii zone (Dicynodon Assemblage zone, LUCAS, 2005), Zone IV.

Age: Upper Tatarian Age, Lopingian Subepoch, Upper Zechstein Epoch, Late Permian.

Material: Skull.

 

= Proelginia permiana HARTMANN-WEINBERG, 1937
Etymology:
= Scutosaurus permianus (HARTMANN-WEINBERG, 1937)

Holotype: N 156/1

Locality: Semin Ravine, near the Ilyinskoe Village, Tetyushky region, European Russia.

Horizon:

Biostratigraphy:

Age: Tatarian Age, Lopingian Subepoch, Zechstein Epoch, Late Permian.

Material: Skull and dentary of a juvenile.

 

IVAKHNENKO, 2008b

Locality: Semin Ovrag Locality, Semin (Syomin) Ovrag Ravine, Cemin Ravin, village of Il’inskoe, right bank of the Volga River, Tetyushi District, Tatarstan, European Russia.

Horizon: Serverodvinian (Dvina) horizon, Semen Formation.

Biostratigraphy: Sokolki Faunal Assemblage, Ilinskoe Faunal Subassemblage.

Age: Upper Tatarian Age, Lopingian Subepoch, upper Zechstein Epoch, Late Permian.

Material: Not given.

 

TVERDOKHLEBOV, TVERDOKHLEBOVA, MINIKH, SURKOV & BENTON, 2005

= Proelginia cf. perminana

Locality: Babentzevo, 3.8 km NE of Babentzevo village, right slope of nameless gully, right tributary of Konduzla River, Samara drainage basin, Orenburg Region, European Russia.

Horizon: Malokinelskaya Svita, Severodovinian Gorizont.

Biostratigraphy:

Age: Upper Tatarian Age, Lopingian Subepoch, upper Zechstein Epoch, Late Permian.

Material:

Number: Not given: Fragments of vertebral column.

Locality: Preobrazhenka, 2.8 km NE of Preobrazhenka village, Vyazovii stream, a right tributary of Konduzla River, Samara drainage basin, Orenburg Region, European Russia.

Horizon: Malokinelskaya Svita, Severodovinian Gorizont.

Biostratigraphy:

Age: Upper Tatarian Age, Lopingian Subepoch, upper Zechstein Epoch, Late Permian.

Material:

Number: Not given:

 

 

= Scutosaurus itilensis IVACHNEKO & LEBEDEV, 1987
Etymology:

Holotype: N 3919/1

Locality: European Russia.

Horizon:

Biostratigraphy:

Age: Late Permian.

Material: Fragmentary skull.