Genus: Howesia BROOM, 1905
Etymology: In honor of Prof. G. B. Howes, whose work on the allied Sphenodon will probably ever remain the standard work on the development of the reptilian skeleton.

Species: browni BROOM, 1905
Etymology: In honor of Alfred Brown.

Holotype: SAM-PK-5884 (specimen A)

Locality: Precise location is unknown, but near the town of Aliwal North, Eastern Cape Province, South Africa.

Horizon: Burgersdorp Formation, Tarkastad Subgroup, Beaufort Group, Karro Supergroup.

Biostratigraphy: Cynognathus Assemblage zone.

Age: Early Anisian Stage, lowermost Mid Triassic Epoch, Early to Middle Triassic.

Material: A partial skull with palate and incomplete lower jaws.

Hypodigm:

 SAM-PK-5885 (Specimen B): A dorsoventrally crushed skull with partial palate, braincase, and atlas-axis complex.

 SAM-PK-5886 (Specimen C): A partial, articulated postcranium consisting of the posterior four dorsal vertebrae, sacrals, first dozen caudals, incomplete pelvic girdles, a partial left hind limb, and a complex right tarsus.

 

SPIEKMAN, BROWNING, TSHIBALANGANDA, MUKHERJEE, SMITH, du PLESSIS AND WOLVAARDT, 2026

Locality: Farm Lemoenfontein (no. 44), c 14 km from the type locality, near the town of Aliwal North, Eastern Cape Province, South Africa.

Horizon: Lower Middle Burgersdorp Formation, Tarkastad Subgroup, Beaufort Group, Karro Supergroup.

Biostratigraphy: Cynognathus Assemblage zone.

Age: Early Anisian Stage, lowermost Mid Triassic Epoch, Early to Middle Triassic.

Material:

SAM-PK-K10180: A virtually complete skull with mandible, right ceratobranchial, distal ends of a tibia and fibula, and a single metataral, and a largely complete right manus.

SAM-PK-K1181: A partial skull with mandible including an articulated front part of the snout, and 2 associated metatarsals.

SAM-PK-K10159: A partial skull with mandible missing the anterior end, right ceratobranchial, and limited remains of the front limb.