Genus: Pachystropheus HUENE, 1935
Etymology: Greek, pachys, "thick" and Greek, stropheus, "vertebra."
= Pachastropheaus ORLOV, 1964 (sic)
= Rysosteus OWEN, 1841
Etymology:
= Ryzosteus HALSTEAD & NICOLL, 1971 (sic)

Species: rhaeticus HUENE, 1935
Etymology: In reference to the Rhaetian, Late Triassic.

Holotype: BMNH R747

Locality: Blue Anchor Pont, (National Grid Reference ST 034 435), Somerset County, England.

Horizon: Westbury Formation, Penarth Group.

Biostratigraphy:

Age: Rhaetian Stage, Uppermost Late Triassic Epoch, Late Triassic.

Material: Dorsal vertebrae ribs, fibula and claws.

Referred material:

BARDET, CUNY, LACHKAR, SUBERBIOLA, MAZIN, THOMEL, BARBIERI, VASSE, PAROT & EMILY, 1998

Locality: Pic des Courmettes, near the city of Grasse (Alpes-Maritimes), Eastern Province, France.

Horizon:

Biostratigraphy:

Age: Rhaetian Stage, Uppermost Late Triassic Epoch, Late Triassic.

Material:

Number: Not given:

 

SANDER, WINTRICH, KINDLIMANN & SCHWERMANN, 2016

Locality: Pit III, a claypit operated by the August Lücking GmbH & Co. KG Brick Company in Bonenburg, Warburg City, Kreis Höxter, North Rhine-Westphalia State, Germany.

Horizon: Contorta Beds.

Biostratigraphy:

Age: Rhaetian Stage, Latest Late Triassic Epoch, Late Triassic.

Material:

LWL MfN P64364: Caudal centra.

LWL MfN P64365: Humerus.

LWL MfN P64367: Interclavicle.

LWL MfN P64368: ?Scapula.

LWL MfN P64369: Ectopterygoid.

 

Locality: Bonebed 2, a claypit operated by the August Lücking GmbH & Co. KG Brick Company in Bonenburg, Warburg City, Kreis Höxter, North Rhine-Westphalia State, Germany.

Horizon: Contorta Beds.

Biostratigraphy:

Age: Rhaetian Stage, Latest Late Triassic Epoch, Late Triassic.

Material:

Number: Not given: Centra.

 


STORRS & GOWER, 1993

Localities: Aust and Garden Cliffs, And Hampstead Farm Quarry, Avon and Gloucestershire County, England.

Horizon: Westbury Formation, Penarth Group.

Biostratigraphy:

Age: Rhaetian Stage, Uppermost Late Triassic Epoch, Late Triassic.

Material:

BMNH R.3711: Dorsal Rib.

BRSMG acc. no. 45/1991: Gasralium.

BRSMG Ce 17165: Dorsal vertebra.

BRSMG Cb4887: Vertebral centrum.

BRSUG 25302: Caudal vertebra.

BRSUG Cb4907: Left humerus.

BRSUG 7010: Right femur.

BMNH R.2728: Proximal portion of right ilium.

BMNH R.4837: Distal portion of left ilium.

BRSMG Ce 17200: Interclavicale.

BRSMG Ce 17081: Interclavicle.

STORRS, GOWER & LARGE, 1996

BATGM C 18, C25, C25A: Anterior caudal? Vertebra.

BATGM C28: Anterior dorsal vertebra.

BATGM M205: Large matrix-free, dorsal vertebra.

BGS GSM 610, GSM 53508: Large femur.

BGS GSM 5394-53598, GSM 112033, GSM 112036, GSM 112039, zr 8601-8604: Isolated, matrix-free ectopterygoid.

BMNH R395, R2728: Right iliac actabulum.

BMNH R4837: Large left illiac blade.

BMNH R6242: Dorsal rib.

BMNH R6243: Dorsal centrum.

BMNH R6245: Caudal rib.

BMNH R6259, R6851: Mid-series dorsal vertebra.

BMNH R12466-12478: Mid-series caudal vertebra.

BMNH R12479-12488: Slab with large femur.

BMNH R12535: Includes a large partial pubis and ilium.

BMNH R12528: Slab baring an ectopterygoid.

BMNH R12537: Indeterminate epipodial.

BMNH R 12554: Miscellaneous postcranial remains on Garden Cliff slabs.

BMNH R12489-12494, R12496-12498, R12500, R12505, R12507-12508, R12510-12512, R12514, R12516-12517, R12520-12521, R12523-12525, R12527, R12529, R12531-12354, R12536, R12539-12543, R12546-12548, R12550-12443:

BMNH 44835: Block containing anterior dorsal vertebra and rib.

BRSMG Cb9065: Slab of associated remains.

BRSMG Cd2422: Proximal end of large right humerus.

BRSMG Cd2678: Large right humerus.

BRSMG Ce1684: Immature right femur.

BRSMG Ce17081: Interclavicle.

BRSMG Ce17146: Dorsal neural arch.

BRSMG Ce17165: Series of dorsal vertebra.

BRSMG Ce17166: Anterior cervical vertebra.

BRSMG Ce17200: Interclavicle.

BRSMG Ce17330: Small femur.

BRSMG Cd3174, Cd3365, Ce17082, Ce17093-17094, Ce17102, Ce17112, Ce17151, Ce17173, Ce17191-17192, Ce17322-17323:

BRSMG Ce17656: Distal caudal vertebra.

BRSMG Ce17770: Slab with associated postcrania.

BRSMG Ce 17775: Small femur.

BRSMG Ce17777-17778: Juvenile humerus.

BRSMG Ce17801: And numerous specimens in lot acc. No. 45/1991: Including a large, matrix-free, right clavicle, a juvinle right ilium, a ?tibia and phalanges, and also a matrix-free gastralium.

BRSUG 7010.4: Right femur.

BRSUG 17037: Mid-series cervical neural arch.

BRSUG 19415.3, 19969: Interclavicle.

BRSUG 25301: Dorsal centrum.

BRSUG 25302.1: Mid-series caudal vertebra.

BRSUG 2503.1: Anterior caudal? Vertebra.

BRSUG 25314: Dorsal rib.

BRSUG 25320: Dorsal vertebra.

BRSUG 25323: Dorsal rib.

BRSUG 23525: Anterior dorsal vertebra.

BRSUG 25329: Left ilium.

BRSUG 25333: Ectopterygoid.

BRSUG 25330: Isolated left pubis.

BRSUG 25331: Sacral vertebra and rib.

BRSUG 25302.2, 25304, 25306, 25308-25313, 25315-25319, 25321-25322, 25324, 25326-25328, 25332, 26161:

CMNHS VP4112: Humerus.

CMNHS VP4114: Ectopterygoid.

CMNHS VP4117: Humerus.

CMNHS 4120: Right humerus.

CMNHS VP4215: Partial pubis.

CMNHS VP4128: Small ectopterygoid.

CMNHS VP4132: Propodials.

CMNHS VP4135: Dorsal vertebra.

CMNHS VP4133: Vertebra.

CMNHS VP4139: Vertebrae and partial femur.

CMNHS VP4642: Femur.

CMNHS VP4113, VP4115-4116, VP4118, VP4119, VP4121-4124, VP42126-VP4127, VP4129-4131, VP4134, VP4136-4138, VP4140, VP4644:

GPIT 19552: One half of a sacral centrum, phalanx, and isolated neural arches.

LEIUG 88988: Proximal end of a humerus.

LEIUG 88989: Juvenile humerus.

LEIUG 88990: ?Sacral vertebral centrum.

MM LL.8018: and others: Associated postcrania from “Slime Road” Cliff.

MNHN SNP101: Abraded dorsal centra.

RSM 1888.87.1, 1911.5.5887: Matrix-free interclavicle.

RSM 1911.5.5918: Proximal end of matrix-free, left humerus.

SMNS 58791: 6 isolated dorsal centra.

STGCM 60.62/1: Slab of associated remains, including vertebrae and limbs, potentially a single individual.

 

= Rysosteus (no species given) OWEN, 1841
= Rysosteus oweni WOODWARD & SHERBORN, 1890
Etymology: In reference to Richard Owen.

Holotype:

Locality: Aust Cliff, near Bristol, Avon County, England.

Horizon: ‘Rhaetic Bone Bed’.

Biostratigraphy:

Age: Rhaetian Stage, Uppermost Late Triassic Epoch, Late Triassic.

Material: Single vertebra.

Referred material:

Locality: Aust and Garden Cliff, Westbury-on-Severn, Gloucestershire County, England.

Horizon:

Biostratigraphy:

Age: Rhaetian Stage, Uppermost Late Triassic Epoch, Late Triassic.

Material:

SO 717 130:

 


Locality: Westbury-on-Severn, Gloucestershire County, England.

Horizon: Rhatic.

Biostratigraphy:

Age: Rhaetian Stage, Uppermost Late Triassic Epoch, Late Triassic.

Material:

Number: Not given:

 

BROWNE, 1894

Locality: ‘Bone-bed’, Aust Cliff, Gloucestershire County, England.

Horizon: Rhatic Strata.

Biostratigraphy:

Age: Rhaetian Stage, Uppermost Late Triassic Epoch, Late Triassic.

Material:

Number: Not given: Dorsal vertebrae, vertebrae and other remains.

 

BROWNE, 1895

Locality: Near East Leake, Nottinghamshire County, England.

Horizon: Rhatic Strata.

Biostratigraphy:

Age: Rhaetian Stage, Uppermost Late Triassic Epoch, Late Triassic.

Material:

Number: Not given:

 

DUFIN, 1980

Locality: Basal bone-bed, Wb 1, Chilcolmpton, Somerset County, England.

Horizon:

Biostratigraphy:

Age: Rhaetian Stage, Uppermost Late Triassic Epoch, Late Triassic.

Material:

Dufin collection: Centrum.

 

MARTILL & DAWN, 1986

Locality: Staple Pit, Newark, Nottinghamshire County, England.

Horizon: Westbury Formation, Penarth Group.

Biostratigraphy:

Age: Rhaetian Stage, Uppermost Late Triassic Epoch, Late Triassic.

Material:

LEIUG 88988: Proximal end of a humerus.

LEIUG 88989: Ilium.

LEIUG 88990: ?Sacral vertebrae.

Also neural arches.