Species: rhaeticus HUENE, 1935
Etymology: In reference to the Rhaetian, Late Triassic.

Holotype: NHMUK PV (old BMNH) R747

Locality: Blue Anchor Pont, (National Grid Reference ST 034 435), Somerset County, England.

Horizon: Westbury Formation, Penarth Group.

Biostratigraphy:

Age: Rhaetian Stage, Uppermost Late Triassic Epoch, Late Triassic.

Material: Dorsal vertebrae ribs, fibula and claws.

Referred material:

BARDET, CUNY, LACHKAR, SUBERBIOLA, MAZIN, THOMEL, BARBIERI, VASSE, PAROT & EMILY, 1998

Locality: Pic des Courmettes, near the city of Grasse (Alpes-Maritimes), Eastern Province, France.

Horizon:

Biostratigraphy:

Age: Rhaetian Stage, Uppermost Late Triassic Epoch, Late Triassic.

Material:

Number: Not given:

 

QUINN, MATHEAU-RAVEN, WHITESIDE, MARSHALL, HUTCHINSON & BENTON, 2024

Locality: Cliff fall, Somerset County, England.

Horizon: Westbury Formation, Penarth Group.

Biostratigraphy:

Age: Rhaetian Stage, Uppermost Late Triassic Epoch, Late Triassic.

Material:

BRSMG Cg3102: Block with 62 vertebrae (mostly dorsal centra, including elements from throughout the vertebral column), representing two or more scattered individuslas, probodials (humeri and femora), epipodials (radius and probable tibia), ribs, phalanges, and hundreds fo unidentifiable fragments.

Note: Also found with it are a Birgeria sp dentary, dinosaur phalanges, probable coelacanth basisphenoid, adn numerous chondrichthyan teeth.

 

SANDER, WINTRICH, KINDLIMANN & SCHWERMANN, 2016

Locality: Pit III, a claypit operated by the August Lücking GmbH & Co. KG Brick Company in Bonenburg, Warburg City, Kreis Höxter, North Rhine-Westphalia State, Germany.

Horizon: Contorta Beds.

Biostratigraphy:

Age: Rhaetian Stage, Latest Late Triassic Epoch, Late Triassic.

Material:

LWL MfN P64364: Caudal centra.

LWL MfN P64365: Humerus.

LWL MfN P64367: Interclavicle.

LWL MfN P64368: ?Scapula.

LWL MfN P64369: Ectopterygoid.

 

Locality: Bonebed 2, a claypit operated by the August Lücking GmbH & Co. KG Brick Company in Bonenburg, Warburg City, Kreis Höxter, North Rhine-Westphalia State, Germany.

Horizon: Contorta Beds.

Biostratigraphy:

Age: Rhaetian Stage, Latest Late Triassic Epoch, Late Triassic.

Material:

Number: Not given: Centra.

 

STORRS & GOWER, 1993

Localities: Aust and Garden Cliffs, And Hampstead Farm Quarry, Avon and Gloucestershire County, England.

Horizon: Westbury Formation, Penarth Group.

Biostratigraphy:

Age: Rhaetian Stage, Uppermost Late Triassic Epoch, Late Triassic.

Material:

NHMUK PV (old BMNH) R.3711: Dorsal Rib.

BRSMG acc. no. 45/1991: Gasralium.

BRSMG Ce 17165: Dorsal vertebra.

BRSMG Cb4887: Vertebral centrum.

BRSUG 25302: Caudal vertebra.

BRSUG Cb4907: Left humerus.

BRSUG 7010: Right femur.

NHMUK PV (old BMNH) R.2728: Proximal portion of right ilium.

NHMUK PV (old BMNH) R.4837: Distal portion of left ilium.

BRSMG Ce 17200: Interclavicale.

BRSMG Ce 17081: Interclavicle.

STORRS, GOWER & LARGE, 1996

BATGM C 18, C25, C25A: Anterior caudal? Vertebra.

BATGM C28: Anterior dorsal vertebra.

BATGM M205: Large matrix-free, dorsal vertebra.

BGS GSM 610, GSM 53508: Large femur.

BGS GSM 5394-53598, GSM 112033, GSM 112036, GSM 112039, zr 8601-8604: Isolated, matrix-free ectopterygoid.

NHMUK PV (old BMNH) R395, R2728: Right iliac actabulum.

NHMUK PV (old BMNH) R4837: Large left illiac blade.

NHMUK PV (old BMNH) R6242: Dorsal rib.

NHMUK PV (old BMNH) R6243: Dorsal centrum.

NHMUK PV (old BMNH) R6245: Caudal rib.

NHMUK PV (old BMNH) R6259, R6851: Mid-series dorsal vertebra.

NHMUK PV (old BMNH) R12466-12478: Mid-series caudal vertebra.

NHMUK PV (old BMNH) R12479-12488: Slab with large femur.

NHMUK PV (old BMNH) R12535: Includes a large partial pubis and ilium.

NHMUK PV (old BMNH) R12528: Slab baring an ectopterygoid.

NHMUK PV (old BMNH) R12537: Indeterminate epipodial.

NHMUK PV (old BMNH) R 12554: Miscellaneous postcranial remains on Garden Cliff slabs.

NHMUK PV (old BMNH) R12489-12494, R12496-12498, R12500, R12505, R12507-12508, R12510-12512, R12514, R12516-12517, R12520-12521, R12523-12525, R12527, R12529, R12531-12354, R12536, R12539-12543, R12546-12548, R12550-12443:

NHMUK PV (old BMNH) 44835: Block containing anterior dorsal vertebra and rib.

BRSMG Cb9065: Slab of associated remains.

BRSMG Cd2422: Proximal end of large right humerus.

BRSMG Cd2678: Large right humerus.

BRSMG Ce1684: Immature right femur.

BRSMG Ce17081: Interclavicle.

BRSMG Ce17146: Dorsal neural arch.

BRSMG Ce17165: Series of dorsal vertebra.

BRSMG Ce17166: Anterior cervical vertebra.

BRSMG Ce17200: Interclavicle.

BRSMG Ce17330: Small femur.

BRSMG Cd3174, Cd3365, Ce17082, Ce17093-17094, Ce17102, Ce17112, Ce17151, Ce17173, Ce17191-17192, Ce17322-17323:

BRSMG Ce17656: Distal caudal vertebra.

BRSMG Ce17770: Slab with associated postcrania.

BRSMG Ce 17775: Small femur.

BRSMG Ce17777-17778: Juvenile humerus.

BRSMG Ce17801: And numerous specimens in lot acc. No. 45/1991: Including a large, matrix-free, right clavicle, a juvinle right ilium, a ?tibia and phalanges, and also a matrix-free gastralium.

BRSUG 7010.4: Right femur.

BRSUG 17037: Mid-series cervical neural arch.

BRSUG 19415.3, 19969: Interclavicle.

BRSUG 25301: Dorsal centrum.

BRSUG 25302.1: Mid-series caudal vertebra.

BRSUG 2503.1: Anterior caudal? Vertebra.

BRSUG 25314: Dorsal rib.

BRSUG 25320: Dorsal vertebra.

BRSUG 25323: Dorsal rib.

BRSUG 23525: Anterior dorsal vertebra.

BRSUG 25329: Left ilium.

BRSUG 25333: Ectopterygoid.

BRSUG 25330: Isolated left pubis.

BRSUG 25331: Sacral vertebra and rib.

BRSUG 25302.2, 25304, 25306, 25308-25313, 25315-25319, 25321-25322, 25324, 25326-25328, 25332, 26161:

CMNHS VP4112: Humerus.

CMNHS VP4114: Ectopterygoid.

CMNHS VP4117: Humerus.

CMNHS 4120: Right humerus.

CMNHS VP4215: Partial pubis.

CMNHS VP4128: Small ectopterygoid.

CMNHS VP4132: Propodials.

CMNHS VP4135: Dorsal vertebra.

CMNHS VP4133: Vertebra.

CMNHS VP4139: Vertebrae and partial femur.

CMNHS VP4642: Femur.

CMNHS VP4113, VP4115-4116, VP4118, VP4119, VP4121-4124, VP42126-VP4127, VP4129-4131, VP4134, VP4136-4138, VP4140, VP4644:

GPIT 19552: One half of a sacral centrum, phalanx, and isolated neural arches.

LEIUG 88988: Proximal end of a humerus.

LEIUG 88989: Juvenile humerus.

LEIUG 88990: ?Sacral vertebral centrum.

MM LL.8018: and others: Associated postcrania from “Slime Road” Cliff.

MNHN SNP101: Abraded dorsal centra.

RSM 1888.87.1, 1911.5.5887: Matrix-free interclavicle.

RSM 1911.5.5918: Proximal end of matrix-free, left humerus.

SMNS 58791: 6 isolated dorsal centra.

STGCM 60.62/1: Slab of associated remains, including vertebrae and limbs, potentially a single individual.

 

= Rysosteus (no species given) OWEN, 1841
= Rysosteus oweni WOODWARD & SHERBORN, 1890
Etymology: In reference to Richard Owen.

Holotype:

Locality: Aust Cliff, near Bristol, Avon County, England.

Horizon: ‘Rhaetic Bone Bed’.

Biostratigraphy:

Age: Rhaetian Stage, Uppermost Late Triassic Epoch, Late Triassic.

Material: Single vertebra.

Referred material:

Locality: Aust and Garden Cliff, Westbury-on-Severn, Gloucestershire County, England.

Horizon:

Biostratigraphy:

Age: Rhaetian Stage, Uppermost Late Triassic Epoch, Late Triassic.

Material:

SO 717 130:

 

Locality: Westbury-on-Severn, Gloucestershire County, England.

Horizon: Rhatic.

Biostratigraphy:

Age: Rhaetian Stage, Uppermost Late Triassic Epoch, Late Triassic.

Material:

Number: Not given:

 

BROWNE, 1894

Locality: ‘Bone-bed’, Aust Cliff, Gloucestershire County, England.

Horizon: Rhatic Strata.

Biostratigraphy:

Age: Rhaetian Stage, Uppermost Late Triassic Epoch, Late Triassic.

Material:

Number: Not given: Dorsal vertebrae, vertebrae and other remains.

 

BROWNE, 1895

Locality: Near East Leake, Nottinghamshire County, England.

Horizon: Rhatic Strata.

Biostratigraphy:

Age: Rhaetian Stage, Uppermost Late Triassic Epoch, Late Triassic.

Material:

Number: Not given:

 

DUFIN, 1980

Locality: Basal bone-bed, Wb 1, Chilcolmpton, Somerset County, England.

Horizon:

Biostratigraphy:

Age: Rhaetian Stage, Uppermost Late Triassic Epoch, Late Triassic.

Material:

Dufin collection: Centrum.

 

MARTILL & DAWN, 1986

Locality: Staple Pit, Newark, Nottinghamshire County, England.

Horizon: Westbury Formation, Penarth Group.

Biostratigraphy:

Age: Rhaetian Stage, Uppermost Late Triassic Epoch, Late Triassic.

Material:

LEIUG 88988: Proximal end of a humerus.

LEIUG 88989: Ilium.

LEIUG 88990: ?Sacral vertebrae.

Also neural arches.