Genus: Tanystropheus MEYER, 1855
Etymology: Greek, tany, "long, stretched" and Greek, strophos, "strap, rope, ribbon"; long ribbion in reference to the slender, elongate shape of the bones.
= Macroscelosaurus MUENSTER, 1830
Etymology:
= Pectenosaurus HUENE, 1902
Etymology:
= Procerosaurus HUENE, 1902
Etymology:
= Tanystrophaeus COPE, 1886 (sic)
= Tribelesodon BASSANI, 1886
Etymology:

Species: conspicuus MEYER, 1847-1855
Etymology: Latin, conspicuus, "visible, striking."
= Macroscelosaurus conspicuus (MEYER, 1847-1855) KUHN, 1937

Holotype: UM-O no. BT 732

Locality: Bindlacher Berg near Bayreuth, Bayern (Bavaria) State, Germany.

Horizon: Upper Muschalkalk.

Biostratigraphy:

Age: Ladinian Stage, Upper Mid Triassic Epoch, Upper Middle Triassic.

Material: Cervical, dorsal, caudal vertebrae, and left humerus.

Referred material:

DIEDRICH, 2012

SMF no. R280: Middle Thoracic vertebra.

SMF no. R285: Cervical 11.

SMF no. R4034: Left humerus.

SMF no. R4092: Anterior caudal vertebra.

SMF no. 54690: Middle thoracic vertebra.

SNSD no. R287: Distal caudal vertebra.

UM-O no. Bayr-65: Tooth.

UM-O No. Bayr-66: Left huemrus of a medium-sized indivdual.

UM-O no. BT 736: Cervical 10.

UM-O no. BT 741: Anterior thoracic vertebra.

UM-O unnumbered: Cervical 5.

UM-O unnumbered: Cervical 9.

 

Locality: Kulmbach, Bayern (Bavaria) State, Germany.

Horizon: Upper Muschalkalk.

Biostratigraphy:

Age: Ladinian Stage, Upper Mid Triassic Epoch, Upper Middle Triassic.

Material:

Number: Not given: Cervical vertebra.

 

Locality: Dorfels near Kronach, Bayern (Bavaria) State, Germany.

Horizon: Upper Muschalkalk.

Biostratigraphy:

Age: Ladinian Stage, Upper Mid Triassic Epoch, Upper Middle Triassic.

Material:

Number: Not given: Dorsal vertebra.

 

Locality: Hegnabrunn near Kulmbach, Bayern (Bavaria) State, Germany.

Horizon: Upper Muschalkalk.

Biostratigraphy:

Age: Ladinian Stage, Upper Mid Triassic Epoch, Upper Middle Triassic.

Material:

Number: Not given: Left femur and a sacral vertebra.

 


Locality: Olmuhle near Crailsheim, Baden-Wurttemberg State, Germany.

Horizon: Upper Muschalkalk.

Biostratigraphy:

Age: Ladinian Stage, Upper Mid Triassic Epoch, Upper Middle Triassic.

Material:

Number: Not given: Broken neural arch of a caudal vertebra.

 


Locality: Bindlach near Bayreuth, Bayern (Bavaria) State, Germany.

Horizon: Upper Muschalkalk.

Biostratigraphy:

Age: Ladinian Stage, Upper Mid Triassic Epoch, Upper Middle Triassic.

Material:

Number: Not given: Sacral vertebra.

 

Locality: Laryszow (Miedary) near Tarnowskie Gory, Upper Silesia, southern Poland.

Horizon: Miedary beds. (Equivalent of the Lowermost Keuper).

Biostratigraphy:

Age: Ladinian Stage, Upper Mid Triassic Epoch, Upper Middle Triassic.

Material:

M.B.S. 11 (number 538, 'Tarnowitz'): Cervical vertebra.

 

ADAMS

Locality: Frankische Grenzxchichten.

Horizon:

Biostratigraphy:

Age: Late Triassic.

Material:

Number: Not given: A cervical vertebrae missing the anterior end.

 

BROILI

Locality: Bayreuth, Bayern (Bavaria) State, Germany.

Horizon: Muschalkalk.

Biostratigraphy:

Age: Mid Triassic Epoch, Upper Middle Triassic.

Material:

Number: Not given: Cervical vertebra.

 

DIEDRICH, 2015

Locality: Enodis/posseckeri bonebed, Lamerden, Hessen State, Germany.

Horizon: Upper Muscheklalk.

Biostratigraphy:

Age: Ladinian Stage, Late Mid Triassic Epoch, Late Middle Triassic.

Material:

LAM-1508a:Lumbar vertebra processus lateralis.

LAM-1508b: Left lumbar vertebra processus lateralis.

 

= Tanystropheus cf. conspicuus SCHOCH, 1999

Locality: N 49°12’52”, F9°40’52”, Autobahn near Kupferzell-Bauersbach, Hohenlohekreis, Baden-Wurttemberg State, Germany.

Horizon: Untere Grau Mergel, Upper Lettenkeuper.

Biostratigraphy:

Age: Longobardian, Upper Ladinian Stage, Upper Mid Triassic Epoch, Late Middle Triassic.

Material:

Number: Not given:

 

= Chelyzoon blezinger HUENE, 1902
Etymology:

Holotype:

Locality: Crailsheim, Baden-Wurttemberg State, Germany.

Horizon: Grenzbonebed.

Biostratigraphy:

Age: Mid Triassic Epoch, Middle Triassic.

Material: Sacral vertebra.

 

= Chelyzoon latum HUENE, 1902
Etymology: Latin, latus, "side."

Holotype:

Locality: Bindlacher berg near Bayreuth, Bayern (Bavaria) State, Germany.

Horizon: Upper Muschelkalk.

Biostratigraphy:

Age: Ladinian Stage, Upper Mid Triassic Epoch, Upper Middle Triassic.

Material: Sacral vertebra.

 

= Procerosaurus cruralis HUENE, 1902
Etymology: Latin, cruralis, "leg."

Holotype:

Locality: Olmuhle near Crailsheim, Baden-Wurttemberg State, Germany.

Horizon: Grenzbonebed.

Biostratigraphy:

Age: Mid Triassic Epoch, Middle Triassic.

Material: Femur.

 

= Pectenosaurus strunzi HUENE, 1902
Etymology:

Holotype:

Locality:

Horizon:

Biostratigraphy:

Age:

Material:

 

= Thecodontosaurus latespinatus HUENE, 1905
Etymology:

Holotype: SMF no. R282

Locality: Bayreuth, Crailsheim, Gottingen and Thuringen, Baden-Wurttemberg State, Germany.

Horizon: Upper Muschelkalk and Grenzbonebed.

Biostratigraphy:

Age: Ladinian Stage, Upper Mid Triassic Epoch, Middle Triassic.

Material: Caudal vertebrae, cervical and dorsal vertebrae.

 

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Species: longobardicus (BASSANI, 1886) PEYER, 1931
Etymology:
= Tribelesodon longobardicus BASSANI, 1886
= Macroscelosaurus longobardicus (BASSANI, 1886) KUHN, 1937

Neotype: PIMUZ T 2791 (examplar a of WILD, 1973)

Locality: Besano, Varesse Province, Italy.

Horizon: Besano Formation.

Biostratigraphy: Grenzbitumen zone.

Age: Anisian /Ladinian Stage, Mid Triassic Epoch, Middle Triassic.

Material: Skull and skeleton.

Referred material:

Locality: Monte San Giorgio, Tessin Canton, Switzerland.

Horizon:

Biostratigraphy:

Age: Anisian /Ladinian Stage, Mid Triassic Epoch, Middle Triassic.

Material: Skull and skeleton.

Skull after Wild, 1973

New interperatation of skull.

Exampler c.

Manus and foot.

Pectoral and pelvic girdle.

New interpertation of a juvenile skull.

 

Locality:

Horizon: Grenzbitumenzone.

Biostratigraphy:

Age:

Material:

T2483 (examplar g of WILD, 1973), T2792 (examplar f of WILD, 1973), T2794 (examplar s of WILD, 1973), T2795 (examplar e of WILD, 1973): Skeletons.

T1270 (examplar x of WILD, 1973), T2782 (examplar n of WILD, 1973), T2789 (examplar y of WILD, 1973): Skulls and skeletons.

T1277 (examplar w of WILD, 1973): Partial skeleton.

T1307 (examplar z of WILD, 1973): Skull, cervical, dorsal and caudal vertebrae.

T2480 (examplar l of WILD, 1973): Skull.

T2481 (examplar t of WILD, 1973): Fragmentary skeleton.

T2482 (examplar d of WILD, 1973): Skull, anterior cervicals.

T2484 (examplar r of WILD, 1973): Caudal, dorsal vertebrae and gastralia.

T2485 (examplar h of WILD, 1973): Posterior cervicals.

T2779 (examplar b of WILD, 1973): Fragmentary remains.

T2780 (examplar o of WILD, 1973): Skull, and cervical vertebrae.

T2781 (examplar aa of WILD, 1973): 2 isolated coracoids, limb bones and dorsal vertebrae.

T2783 (examplar u of WILD, 1973): Deformed skull, and dorsal vertebrae.

T2788 (examplar ab of WILD, 1973): Fragmentary skeleton.

T2817 (examplar k of WILD, 1973): Skull and cervicals.

 

= Tanystropheus cf. T. longobardicus DALLA VECCHIA, 2006

Locality: Site 1, Aupa Valley, Moggio Udinese, Udine Province, Friuli Venezia Giulia Region, Northeastern Italy.

Horizon: "Torbidti d'Aupa"

Biostratigraphy:

Age: Late Anisian Stage, Lower Mid Triassic, Lower Middle Triassic.

Material:

MFSN 31545: Posterior cervical vertebra.

 

Locality: Site 2, Aupa Valley, Moggio Udinese, Udine Province, Friuli Venezia Giulia Region, Northeastern Italy.

Horizon: "Torbidti d'Aupa"

Biostratigraphy:

Age: Late Anisian Stage, Lower Mid Triassic, Lower Middle Triassic.

Material:

MFSN 27269, 31544: Cervical vertebrae.

MFSN 27274: Dorasal vertebra.

MFSN 27276: Dorsal neural arch.

MFSN 27277: Neural arch of a dorsal or caudal.

MFSN 27281: Proximal cervical verebra.

MFSN 28521: Fragmentary anterior cervical vertebra.

 

Locality: Site 4, Aupa Valley, Moggio Udinese, Udine Province, Friuli Venezia Giulia Region, Northeastern Italy.

Horizon: "Torbidti d'Aupa"

Biostratigraphy:

Age: Late Anisian Stage, Lower Mid Triassic, Lower Middle Triassic.

Material:

MFSN 31554: Tooth.

 

Locality: Site 5, Aupa Valley, Moggio Udinese, Udine Province, Friuli Venezia Giulia Region, Northeastern Italy.

Horizon: "Torbidti d'Aupa"

Biostratigraphy:

Age: Late Anisian Stage, Lower Mid Triassic, Lower Middle Triassic.

Material:

MFSN 31547: Fragmentary anterior cervical vertebrae.

 

Locality: Site 6, Aupa Valley, Moggio Udinese, Udine Province, Friuli Venezia Giulia Region, Northeastern Italy.

Horizon: "Torbidti d'Aupa"

Biostratigraphy:

Age: Late Anisian Stage, Lower Mid Triassic, Lower Middle Triassic.

Material:

MFSN 31545: Partial posterior cervical vertebra.

MFSN 31548: Fragmentary cervical vertebra.

MFSN 31549:Fragmentary caudal vertebra.

MFSN 31550: Proximal caudal vertebra.

MFSN 31551: Fragmentary ?dorsal neural arch.

MFSN 31552: Sacaral vertebra.

MFSN 31553: Clavical.

MFSN 31556, 31557: 2 teeth.

MFSN 31560, 31561: 2 teeth.

MFSN 31562: Fragmentary centa.

MFSN 31573: Ilium.

MFSN 31574: Distal caudal vertebra.

MFSN 31575: Distal caudal vertebra.

MFSN 31576: Cervical centra.

MFSN 31579: Complete and isolated cervical vertebra.

MFSN 31580: Distal caudal vertebra.

MFSN 31581: Mid-posterior dorsal vertebrae.

MFSN 31596: Mid-posterior dorsal vertebra.

MFSN 27271: Dorsal vertebra.

MFSN 27272: Fragmenty of vertebra.

MFSN 27273: Fragmmentary dorsal vertebra.

MFSN 27275: Fragmentary caudal vertebra.

 

DIEDRICH, 2009

Locality: Bissendorf, Osnabrucker Bergland, Lower Saschen (Saxony) State, northwestern Germany.

Horizon: Meißner Formation, Upper Muschelkalk.

Biostratigraphy:

Age: Anisian/Landinan Boundary, Mid Triasic Epoch, Middle Triassic.

Material:

Coll.-No. Pal 327-534: Tooth.

NOSOTTI, 1999

Locality: Besano, Lombardy, Varesse Province, Northern Italy.

Horizon: Besano Formation.

Biostratigraphy:

Age: Anisian-Lower Ladinian Stage, Lower Mid Triassic Epoch, Middle Triassic.

Material:

BES SC 265, 1018, 3730: Nearly complete specimens.

Dorsal and lateral view of skull (after NOSOTTI, 2007)

 

= Tanystropheus cf. T. longobardicus RENESTO, 2005

Locality: Val Serrata, Monte San Giorgio, Canton Tessen, Switzerland.

Horizon: Lower Cava inferiore beds, Meride Limestone.

Biostratigraphy:

Age: Ladinian Stage, Lower Mid Triassic Epoch, Middle Triassic.

Material:

MCSN 4451: Fragmentary skeleton lacking the head and first 8 cervical vertebrae, and most bones fo both hind limbs.

Note: Has skin impressions.

 

= Tanystropheus cf. T. longobardicus RIEPPEL, JIANG, FRASER, HAO, MOTANI, SUN & SUN, 2010

Locality: Heshangsi of Nimaigu Village, Wusha District, Xingyi City, Guizhou Province, China.

Horizon: Zhuganpo Member, Falang Formation.

Biostratigraphy:

Age: Latest Ladinian Stage, Lower Mid Triassic Epoch, Middle Triassic.

Material:

GMPKU-P-1537: Nearly complete skeleton lacking the skull and first few cervical vertebrae.


= Tanystropheus meridensis WILD, 1980
Etymology:

Holotype: T/3901 PIUZ

Locality: Monte San Giorgio, Tessin Canton, Switzerland.

Horizon: Lower Meridekalke.

Biostratigraphy:

Age: Ladinian Stage, Upper Mid Triassic Epoch, Lower Late Triassic.

Material: Skull, lower jaws, and 5 cervical vertebrae with ribs.

New interpertation of the skull.

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Species: biharicus JURSAK, 1975
Etymology: In reference to Bihor County, Romania.

Holotype: Nr. X, Muzeul Tairl Crrisurilor, Oradea, inv. nr. 8988.

Locality: Lugasu de Sus, judetul Bihor County, Romania.

Horizon:

Biostratigraphy:

Age: Lower Anisian Stage, Lower Mid Triassic Epoch, Middle Triassic.

Material: Cervical vertebra.

_____________________________________________________________________________________

Species: fossai WILD, 1983
Etymology:

Holotype: Nr 4035 MCSNB

Locality: North Hang Canto Alto, Val Brembana, Italy.

Horizon:

Biostratigraphy:

Age: Upper Norian Stage, Late Triassic Epoch, Late Triassic.

Material: 4 cervical vertebrae and ribs.
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Species: haasi RIEPPEL, 2001
Etymology: In honor of the late Prof. G. Haas, who initiated the study of fossil reptile fauna from Makhtesh Ramon.
= Tanystropheus sp PEYER, 1953

Holotype: HUJ-Pal. TR. 1

Locality: Makhtesh Ramon, Negev, Isreal.

Horizon: Muschelkalk.

Biostratigraphy:

Age: Upper Anisian- lower Ladinian Stage, Middle Mid Triassic Epoch, Middle Triassic.

Material: Posterior end of 8th cervical vertebra.

Referred material:

HUJ-Pal. TR 1447: Posterior end of a cervical vertebra with the middle cervical region (cf 6 through 8, probably cv 8.

HUJ-PAL. TR 9, 11, -13, 17, 20, 26-28, 39, 44, 50, 51, 1451, 1452, 1456, 1464, 1465, 2126, 2127, 2130, 2258, 2264, 2736: Incomplete fragments of cervical vertebrae.

2 uncatalogued specimens.

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Species: hydroides SPIEKMAN, NEENAN, FRASER, FERNANDEZ, RIEPPEL, NOSOTTI & SCHEYER, 2020
Etymology: Hydra refering to the long-necked mythical sea monster of Greek mythology, and Greek, -oides, "related to or resembling."
Note: Large individuals of Tanystropheus longobardicus.

Holotype: PIMUZ T2790

Locality: Monte San Giorgio on the border of Switzerland (Canton Ticino) and Italy (Lombardy). ,

Horizon: Besano Formation.

Biostratigraphy: Grenzbitumen zone.

Age: Anisian /Ladinian Stage, Mid Triassic Epoch, Middle Triassic.

Material: Skull and caudal vertebrae.

Referred material:

MSNM V 3663:

PIMUZ T183 (examplar v of WILD, 1973): Skull and skeleton.

PIMUZ T2787 (examplar i of WILD, 1973): Skull and skeleton.

PIMUZ T2793 (examplar c of WILD, 1973): Skull and skeleton.

PIMUZ T2818 (examplar m of WILD, 1973): Skull and skeleton.

PIMUZ T2819 (examplar q of WILD, 1973): Skeleton.

SNSB-BSPG 1953 XV 2: