Genus: Trilophosaurus CASE, 1928
Etymology: Greek, tri, "three," Greek, lophus, "crest" and Greek, sauros, "lizard."
= Chinleogomphius SUES & OLSEN, 1993
Etymology: In reference to the Chinle Formation, Arizona, Greek, gomphios, “cheek, tooth”: Chinle cheek-tooth.

Species: buettneri CASE, 1928
Etymology: In honor of William Buettner.

Holotype: UMMP 2338

Locality: Walker's Tank, Crosby County, Western Texas.

Horizon: Tecovas Member, Dockum Formation.

Biostratigraphy:

Age: Carnian Stage, Late Triassic Epoch, Late Triassic.

Material: An incomplete right dentary fragment bearing parts of 5 teeth..

Referred material:

 

GREGORY, 1945, DEMAR & BOLT, 1981, SPIELMANN, LUCAS, RINEHART & HECKERT, 2008

Locality: Trilophosaurus Quarry 1 (NMMNH locality 860, TMM locality 31025, YPM 5830) , 800 feet south and 1100 feet west of the northeast corner of a 4-section tract belonging to Mrs. Sarah Mina Hyman, Mrs. Otis Chalk, and Mrs. Doris Roberts, it lies in the extreme southeast corner of sec. 58, blk, 29, Waco and Northwestern Railroad Company Survey, about 3 miles north of Otis Chalk, Howard County, Texas.

Horizon: Colorado City Formation (“Pre-Tecovas Horizon”), Dockum Group.

Biostratigraphy:

Age: Carnian Stage, Lower Late Triassic Epoch, Late Triassic.

Material:

TMM 31025-1, 31025-2, 31025-5, 31025-93, 31025-96, 31025-103-104, 31025-108-111, 31025-115-116, 31025-119, 31025-125, 31025-175, 31025-194, 31025-196, 31025-205-206, 31025-209, 31025-214, 31025-217-221, 31025-233-224, 31025-226, 31025-233, 31025-236-238, 31025-248-250, 31025-254, 31025-630, 31025-633-635, 31025-679, 31025-754-758, 31025-763, 31025-871: Mandibulae.

TMM 31025-4: Frontal and parietal.

TMM 31025-10, 31025-67, 31025-67TT, 31025-693-695, 31025-826-828: Femora.

TMM 31025-13-18, 31025-20-33, 31025-35-40, 31025-44-45, 31025-47-48, 31025-50-55, 31025-57-62, 31025-84-85, 31025-123-124, 31025-161-162, 31025-173, 31025-212, 31025-275, 31025-578, 31025-627, 31025-824: Vertebrae.

TMM 31025-41-42, 31025-141, 31025-195: Skeletons.

TMM 31025-46: Middle cervical vertebra.

TMM 31025-66, 31025-184, 31025-406, 31025-689-692, 31025-705-708, 31025-743, 31025-792, 31025-830-831, 31025-849-851, 31025-892-893, 31025-897: Humerus.

TMM 31025-68, 31025-411, 31025-487, 31025-506, 31025-523, 31025-535, 31025-687-688, 31025-714-717, 31025-801-802, 31025-802, 31025-821, 31025-836, 31025-852, 31025-869: Scapula.

TMM 31025-68R: Right scapula.

TMM 31025-69, 31025-803, 31025-825, 31025-871-872: Coracoid.

TMM 31025-69H: Right coracoid.

TMM 31025-70-75, 3107-79, 31025-122, 31025-167, 31025-186-188, 31025-200, 31025-480, 31025-484-486, 31025-488, 31025-515, 31025-589, 31025-680-681, 31025-686, 31025-699-702, 31025-720-722, 31025-724-725, 31025-748-753, 31025-762, 31025-796-800, 31025-822-823, 31025-835, 31025-864-867, 31025-898-900: Pelvis.

TMM 31025-76, 31025-185: Scapulocoracoid.

TMM 31025-77: Right ilium.

TMM 31025-77D: Left ilium.

TMM 31025-78, 31025-78I: Left ischia.

TMM 31025-79B: Pubis.

TMM 31025-80, 31025-201, 31025-516, 31025-696-698, 31025-713, 31025-739-741, 31025-747, 31025-759, 31025-788-789, 31025-829, 31025-847-848, 31025-887-889: Tibia.

TMM 31025-80F: Tibia.

TMM 31025-80P: Left tibia.

TMM 31025-81, 31025-407-408, 31025-492, 31025-623, 31025-682-685, 31025-704, 31025-709, 31025-744-745, 31025-793, 31025-795, 31025-832, 31025-853, 31025-863, 31025-895-896: Ulna.

TMM 31025-81H: Right ulna.

TMM 31025-82: Right radius.

TMM 31025-82P: Left radius.

TMM 31025-83, 31025-189, 31025-481-482, 31025-703, 31025-742, 31025-790-791, 31025-890-891: Fibula.

TMM 31025-83A, 31025-83C: Fibulae.

TMM 31025-83M: Left radius.

TMM 31025-86: Fibulae.

TMM 31025-87, 31025-277: Rib.

TMM 31025-88, 31025-113, 31025-129, 31025-131, 31025-134-136, 31025-170-171, 31025-202, 31025-410, 31025-412-413, 31025-447, 31025-449-451, 31025-456-459, 31025-461, 31025-464-467, 31025-471-472, 31025-475-478, 31025-525-529, 31025-531, 31025-533-534. 31025-536, 31025-540, 31025-543, 31025-550, 31025-554-555, 31025-557, 31025-567, 31025-572-577, 31025-584, 31025-598, 31025-604, 31025-606-608, 31025-610, 31025-612-613, 31025-616-617, 31025-620-621, 31025-624, 31025-626, 31025-638, 31025-640-651, 31025-658-661, 31025-726-729, 31025-735, 31025-764, 31025-766-769, 31025-772-773, 31025-775-777, 31025-806-808, 31025-937, 31025-840-841, 31025-843, 31025-854-857, 31025-874-876: Phalanges.

TMM 31025-89, 31025-203: Tarsal.

TMM 31025-90: Caudal chevrons.

TMM 31025-91-92, 31025-94-95, 31025-97-102, 31025-106-107, 31025-112, 31025-120-121, 31025-127, 31025-138, 31024-142, 31025-143, 31025-160, 31025-208, 31025-210-211, 31025-213, 31025-215-216, 31025-222, 31025-225, 31025-228-231, 31025-234-235, 31025-239-244, 31025-251-253, 31025-255-256, 31025-274, 31025-499, 31025-518, 31025-629. 31025-631-632, 31025-678: Skulls.

TMM 31025-105, 31025-127: Skulls missing posterior end.

TMM 31025-128, 31025-132-133, 31025-137, 31025-159, 31025-199, 31025-446, 31025-452, 31025-460, 31025-468-469, 31025-474-475, 31025-537-539, 31025-541-542, 31025-547-549, 31025-551-552, 31025-556, 558-560, 31025-565, 31025-571, 31025-579-581, 31025-590, 31025-593, 31025-595, 31025-597, 31025-599, 31025-603, 31025-609, 31025-611, 31025-625, 31025-639, 31025-652-653, 31025-655, 31025-663-665, 31025-668-672, 31025-730-734, 31025-765, 31025-770-771, 31025-778-779, 31025-810-819, 31025-838-839, 31025-842, 31025-858, 31025-879-881: Metatarsal.

TMM 31025-129, 31025-409, 31025-453, 31025-462-463, 31025-483, 31025-546, 31025-596, 31025-600-601, 31025-615, 31025-618-619, 31025-673-675, 31025-859-860, 31025-878: Metacarpal.

TMM 31025-140: Skull missing anterior end, dentary missing anterior end, 8 cervical, 2 dorsal, 2 sacral, and 1 caudal vertebrae, ilium, ribs, pectoral girdle, left humerus, right ulna, femur, right astragalus, left astragalus, right calcaneus, right navicular, left navicular, right pes, and left manus.

TMM 31025-143: Skull missing posterior end.

TMM 31025-144, 31025-174: Interclavicle.

TMM 31025-145-146, 31025-166, 31025-448, 31025-491: Metatacarpal/metatarsal?

TMM 31025-168-169: Sacrum.

TMM 31025-183, 31025-191, 31025-198, 31025-245, 31025-247: Specimen.

TMM 31025-190, 31025-414, 31025-710-712, 31025-738, 31025-774, 31025-786-787, 31025-846, 31025-862, 31025-884-886: Femur.

TMM 31025-193: Fibulare.

TMM 31025-197, 31025-507: Carpal.

TMM 31025-204, 31025-493, 496, 31025-500-504, 31025-508-511-514, 31025-564, 31025-628: Chevron.

TMM 31025-232: Skull? mandible.

TMM 31025-246: Articular.

TMM 31025-276: Limb bone.

TMM 31025-405, 31025-455, 31025-479, 31025-489, 31025-520-522, 31025-524, 31025-532, 31025-545, 31025-677, 31025-718, 31025-760-761, 31025-809, 31025-883: Calcanium.

TMM 31025-417, 31025-746, 31025-794, 31025-833-834, 31025-861, 31025-868: Radius.

TMM 31025-454, 31025-470, 31025-505, 31025-544, 31025-561, 31025-568, 31025-654, 31025-656, 31025-844: Tarsal element.

TMM 31025-519, 31025-530, 31025-582-583, 31025-585, 31025-587, 31025-667, 31025-676, 31025-882: Astragalus.

TMM 31025-661: Right humerus.

TMM 31025-870: Scapula?

NMMNH P-18418: Right distal femur.

NMMNH P-18420: Right proximal humerus.

NMMNH P-18421: 3 proximal phalanges and 2 terminal phalanges.

NMMNH P-18425: 4 caudal centra.

NMMNH P-29396: Proximal left femur.

NMMNH P-29397: 2 unguals.

NMMNH P-29398: 1 vertebra.

NMMNH P-34010: 1 incomplete tooth.

NMMNH P-34051: 2 distal left humeri.

NMMNH P-34059: 1 caudal centrum.

NMMNH P-34070: 1 proximal femur.

NMMNH P-37367: 1 phalanx.

NMMNH P-37373: 1 chevron.

NMMNH P-37378: 5 assorted postcranial fragments.

NMMNH P-37383: 1 partial vertebra.

NMMNH P-37396: 1 elongate caudal centrum.

NMMNH P-40927: 1 ?atlas.

NMMNH P-40928: 1 ?atlas.

NMMNH P-41349: 1 badly worn, transversely broad tooth fragment.

NMMNH P-42251: 1 left maxilla with parts of 5 teeth.

NMMNH P-42252: 1 left? lower jaw fragment with 4 teeth.

NMMNH P-42253: 1 left? lower jaw fragment with 3 teeth.

NMMNH P-42254: 1 right lower jaw fragment with 3 concreted teeth.

NMMNH P-42255: 1 concreted jaw with 6 teeth.

NMMNH P-42256: 1 jaw fragment with 2 teeth.

NMMNH P-43357: 1 cervical centrum.

NMMNH P-42258: 1 cervical? vertebra.

NMMNH P-42259: 1 dorsal vertebra.

NMMNH P-42260: 1 concreted dorsal vertebra.

NMMNH P-42261: 3 caudal vertebrae.

NMMNH P-42262: 1 tiny distal left humerus of juvenile/hatchling.

NMMNH P-42263, 42264: 1 right calcaneum.

NMMNH P-42265, 42266: 1 right calcaneum fragment.

NMMNH P-42267: 1 left astragalus.

NMMNH P-42268: 1 left calcaneum.

NMMNH P-42269: 18 ungual phalanx fragments.

NMMNH P-42270: 1 posterior cervical centrum.

NMMNH P-42271: 1 tiny caudal vertebra.

NMMNH P-42272: 1 concreted dorsal vertebra in matrix.

NMMNH P-42273: 7 caudal vertebrae.

NMMNH P-42274: 4 fragmentary caudal vertebrae.

NMMNH P-42275: 1 proximal pubis.

NMMNH P-42276: 1 proximal femur.

NMMNH P-42277: 34 proximal metapodials and/or phalanges.

NMMNH P-42278: 5 distal phalanges.

NMMNH P-42279: 9 distal phalanges.

NMMNH P-44763: 1 juvenile anterior right? maxillary fragment.

NMMNH P-44764: 1 incomplete tooth.

NMMNH P-44765: 1 small neural spine.

NMMNH P-44987: 2 tiny distal phalanges.

YPM 4186: Incomplete lower jaws.

YPM 41878, 4820: Braincases.

LONG & MURRY, 1995

TMM 31025-207, TMM 31025-211, TMM 31025-239, TMM 31025-241: Partial skulls.

TMM 31025-142, TMM 31025-240, TMM 31025-244: Braincases.

TMM 31025-265: Left femur of a robust trilophosaurid.


DEMAR & BOLT, 1981

Locality:

Horizon:

Biostratigraphy:

Age:

Material:

TMM 31099: 1, 13C, 13D, 13F, 14, 15, 15A, 16-22.

 

Locality: Petrified Forest National Park, Apache County, Arizona.

Horizon: Lower Petrified Forest Member, Chinle Formation.

Biostratigraphy:

Age: Late Carnian Stage, Lower Late Triassic Epoch, Late Triassic.

Material:

PEFO 3893, 3894, 3895: 3 teeth.

 

HECKERT, 2001

Locality: Lower Kalgary site, NMMNH Locality 1312, Crosby County, Texas.

Horizon: Colorado City Member, Dockum Formation, Chinle Group (Lower Kalgary).

Biostratigraphy:

Age: Upper Otischalkian, Late Carnian Stage, Lower Late Triassic Epoch, Late Triassic.

Material:

NMMNH P-34291: Tooth.

NMMNH P-34292: Fragmentary polycuspate teeth.

= Trilophosaurus? sp

NMMNH P-34293: Numerous fragmentary teeth.

NMMNH P-34294: Enamel-less polycuspate teeth.

SMU 67496-67511, 67513-67562: isolated teeth.

 

Locality: NMMNH 1430, at the head of the south fork of Home Creek north of Kalgary Locality, UTM 3701500N, 398850E, zone 14, NAD 27, Crosby County, Texas.

Horizon: Colorado City Member, Tecovas Formation, Chinle Group.

Biostratigraphy:

Age: Adamanian, Latest Carnian Stage, Late Triassic Epoch, Late Triassic.

Material:

NMMNH P-26422: Numerous teeth with low cusps lacking cingula.

NMMNH P-34074: Incomplete tooth.

NMMNH P-34075: High-cupsed tooth.

NMMNH P-34373, 34374, 34375: Enamel-less teeth.

NMMNH P-34376: Nearly complete teeth.


KAYE & PADIAN, 1992/KAYE & PADIAN, 1994

Locality: UCMP loc. A269, the Placerias quarry, in a small valley 6 1/2 miles S 65"W of St. Johns, Located 0.3 miles northwest of Romero Springs at approximately Long. 109'28'W and lat. 34"27'N on the USGS St. Johns quadrangle of 1921, Apache County, Arizona.

Horizon: Bluewater Creek Formation, Lower Chinle Group.

Biostratigraphy:

Age: Adamanian, Latest Carnian Stage, Late Triassic Epoch, Late Triassic.

Material:

UCMP V3730: Jaw fragment.

 

MUELLER, HUTTENLOCKER, SMALL, PINTO, DEAN-WALLACE & CHATTERJEE, 2022

Locality: Boren Quarry (MOTT VPL 3869), Garza County, Texas.

Horizon: Tecovas Formation, Dockum Group.

Biostratigraphy: Otischalkian LVF.

Age: Early Norian Stage, Middle Late Triassic Epoch, Middle Late Triassic.

Material:

Number: Not given:

 

MURRY, 1986

Locality: Saurian Valley, PEFO 085002 (UCMP V82251, PF97) (part of UCMP V7034), NW 1/4 of SE 1/4, Sec. 15, and NE 1/4 of SW 1/4, Sec. 15. T 19N, R 23E, Lithodendron Wash Area, 6 to 8 m below? Sonsela Sandstone, Devil’s Playground Area, Petrified Forest, Apache County, Arizona.

Locality: Potter County, Texas.

Horizon: Dockum Formation.

Biostratigraphy:

Age: Carnian Stage, Lower Late Triassic Epoch, Late Triassic.

Material:

Number: Not given:

 


MURRY, 1989

Locality: Sunday Canyon Locality, SMU 123, 34'57"18N, 101'42"15W, badland along Sundy Creek, Fortress Cliff 7.5' Quadrangle, Randall County, Texas.

Horizon: Tecovas Member, Dockum Formation.

Biostratigraphy:

Age: Carnian Stage, Late Triassic Epoch, Late Triassic.

Material:

Number: Not given:


Locality: Rotten Hill/Sierrita de la Cruz Locality, SMU 121, 35'23"06N, 102'09"40W, badlands along Sierrita de la Cruz Creek, Ady 7.5 Quadrangle, Potter County, Texas.

Horizon: Tecovas Member, Dockum Formation.

Biostratigraphy:

Age: Carnian Stage, Late Triassic Epoch, Late Triassic.

Material:

Number: Not given:

LONG & MURRY, 1995

Locality: Placerias quarry (UCMP A269, MNA 207C), Big Hollow Wash near Romero Springs, a small valley 6 1/2 miles (approximately 10.4 km) S 65"W of St. Johns, Located 0.3 miles northwest of Romero Springs at approximately Long. 109'28'W and lat. 34"27'N on the USGS St. Johns quadrangle of 1921, Apache County, Arizona.

Horizon: Bluewater Creek Formation, Lower Chinle Group.

Biostratigraphy:

Age: Adamanian, Latest Carnian Stage, Late Triassic Epoch, Late Triassic.

Material:

UCMP 32390: Proximal two-thirds of left femur.

UCMP 25877: Distal extremity of left humerus.

UCMP 25825, 25839: Proximal and distal extremities of left tibia.

UCMP 25826, 15798: Proximal and distal extremities of right sacral vertebrae.

UCMP 25790: Proximal extremity of right tibia.

UCMP 25424: Second sacral vertebra.

MNA Pl. 2924: Sacrals 1 and 2.

MAN Pl. 2956, 2928: Left and right ilia

Note: All MNA specimens may represent a single individual.

 

Locality: Agate Bridge N (UCMP V82339, PF 161), NE 1/4 of SE 1/4 and SE 1/4 of NE 1/4, SEC. 4, T 17N, R 24E, near base of black/grey siltstone, approximately 6 m below Sonsela Sandstone, resting on a loose sandstone, Lot’s Wife Area, Petrified Forest, Apache County, Arizona.

Horizon: Lower Petrified Forest Member, Chinle Formation.

Biostratigraphy:

Age: Late Carnian Stage, Lower Late Triassic Epoch, Late Triassic.

Material:

UCMP: Caudal vertebrae.

NMMNH: 2 caudal vertebrae.

Note: HUNT, LUCAS, HECKERT, SULLIVAN & LOCKLEY, 1998 state that pers. comm. from LONG, that Coelophysis cervical (elongate centra) vertebrae from this locality belong to Trilophosaurus.

 

Locality: Saurian Valley, PEFO 085002 (UCMP V82251, PF97) (part of UCMP V7034), NW 1/4 of SE 1/4, Sec. 15, and NE 1/4 of SW 1/4, Sec. 15. T 19N, R 23E, Lithodendron Wash Area, 6 to 8 m below? Sonsela Sandstone, Devil’s Playground Area, Petrified Forest, Apache County, Arizona.

Horizon: Lower Petrified Forest Member, Chinle Formation.

Biostratigraphy:

Age: Late Carnian Stage, Lower Late Triassic Epoch, Late Triassic.

Material:

UCMP 126749: Caudal vertebra.

UCMP: Ilium.


Locality: Hole at Canada Colorado, 4 km east of Lamy, Santa Fe County, New Mexico.

Horizon: Los Esteros Member, Santa Rosa Formation.

Biostratigraphy:

Age: Lower Carnian Stage, Lower Late Triassic Epoch, Late Triassic.

Material:

NMMNH: Proximal extremity of right tibia of a very large individual.

 

Locality: Trilophosaurus Site 1 (TMM Locality 31213), Borden County, Texas.

Horizon: Colorado City Formation (“Pre-Tecovas Horizon”), Dockum Group.

Biostratigraphy:

Age: Carnian Stage, Lower Late Triassic Epoch, Late Triassic.

Material:

TMM 31213-8: Large humerus.

TMM 31213-12: Nearly complete proximal and distal portions of humeri.

TMM 31213-11: Proximal ends of three femora.

TMM 31213-13: Vertebrae (mixed with dorsal vertebrae and other taxa).

 

Locality: Quarry 3, (TMM Locality TMM 31100-443, Near Otis Chalk, Howard County, Texas.

Horizon: Los Esteros Member, Santa Rosa Formation.

Biostratigraphy:

Age: Lower Carnian Stage, Lower Late Triassic Epoch, Late Triassic.

Material:

TMM 31100-443: Braincase.

 


MURRY & LONG, 1989

Locality: Dying Ground (PF 121, SMU #228), Sec. 23, T 18N, R 24E in variegated siltstone bed, 34'56"36N, 109'45"55W, Blue Mesa W Area, Petrified Forest, Apache County, Arizona.

Horizon: Chinle Formation.

Biostratigraphy:

Age: Late Triassic.

Material:

Number: Not given:


SPIELMANN, LUCAS, RINEHART & HECKERT, 2008

Locality: Trilophosaurus quarry 2 (NMMNH locality 4208, TMM Locality 31099), Howard County, Texas.

Horizon: Colorado City Formation (“Pre-Tecovas Horizon”), Dockum Group.

Biostratigraphy:

Age: Carnian Stage, Lower Late Triassic Epoch, Late Triassic.

Material:

NMMNH P-36232: Postcrania.
Note: Associated with a small skull of Buettneria.

NMMNH P-36233: 1 distal femur.

NMMNH P-36235: Postcrania associated with clavicle.

NMMNH P-36326: 1 proximal metapodal.

NMMNH P-39327: 1 centrum.

NMMNH P-39328: 1 centrum fragment.

NMMNH P-39793: 1 left tibia fragment.

NMMNH P-39794, P-39795: 1 distal left tibia.

NMMNH P-39796: 1 left tibia.

NMMNH P-39797, P-39798: 1 proximal right tibia.

NMMNH P-39799: 1 distal right tibia.

NMMNH P-39800: 1 left tibia.

NMMNH P-39801-P39803: Distal right tibia.

NMMNH P-39804: Right tibia.

NMMNH P-39805: 1 distal right tibia.

NMMNH P-39806: 1 left humerus.

NMMNH P-39807: 1 right tibia.

NMMNH P-39808-P39809: 1 proximal left humerus.

NMMNH P-39810: Distal left humerus.

NMMNH P-39811: 1 left humerus.

NMMNH P-39812: 1 incomplete right humerus.

NMMNH P-39813: 1 distal left tibia.

NMMNH P-39814, P-39815: 1 right humerus.

NMMNH P-39816-P39817: 1 proximal right humerus.

NMMNH P-39818: 1 left femur.

NMMNH P-39819: 1 distal left femur.

NMMNH P-39820-P39821: 1 proximal left femur.

NMMNH P-P39022: 1 right humerus.

NMMNH P-39823: 1 left femur.

NMMNH P-39824-P-39825: 1 distal left femur.

NMMNH P-39826-P39827: 1 proximal left humerus.

NMMNH P-39828-P39830: Distal left femur.

NMMNH P-39831: 1 distal right? femur.

NMMNH P-39832: 1 right femur.

NMMNH P-39833: 1 proximal right femur.

NMMNH P-39834-P-39835: 1 right distal femur.

NMMNH P-39836-P-39837: 1 proximal right femur.

NMMNH P-39838: 1 right femur.

NMMNH P-39849, P-39842: 1 distal right femur.

NMMNH P-39843, P-39844: Proximal right femur.

NMMNH P-39845-P-39847: 1 proximal left ulna.

NMMNH P-39848-P-39851: 1 distal left ulna.

NMMNH P-29853: 1 distal right ulna.

NMMNH P-29854, P-39855: 1 proximal right ulna.

NMMNH P-39856-P-39862: 1 distal right ulna.

NMMNH P-39863: 1 right scapula fragment.

NMMNH P-39864: 1 left coracoid fragment.

NMMNH P-39865: 1 left? coracoid fragment.

NMMNH P-39866: 1 left scapula fragment.

NMMNH P-39867, P-39868: 1 interclavicle? fragment.

NMMNH P-39869: 36 ribs and rib fragments.

NMMNH P-39870: numerous metapodial fragments.

NMMNH P-39871: 3 cervical vertebrae.

NMMNH P-39872: 1 dorsal vertebra.

NMMNH P-39873: 2 anterior caudal vertebrae.

NMMNH P-39874: 4 caudal? vertebrae.

NMMNH P-39875: 20 dorsal vertebrae.

NMMNH P-39876: 20 caudal vertebrae.

NMMNH P-39939: 78 caudal vertebrae.

NMMNH P-39940: 12 sacral vertebrae.

NMMNH P-39941: 15 dorsal vertebrae.

NMMNH P-39942: 50 miscellaneous centra.

NMMNH P-39949: 24 miscellaneous centra.

NMMNH P-39953, P-39954: 1 cervical vertebra.

NMMNH P-39960: 1 distal right ulna.

NMMNH P-39961: 18 caudal vertebrae.

NMMNH P-39962, P-39963: 1 dorsal vertebra.

NMMNH P-33964: 1 anterior caudal vertebra.

NMMNH P-39965: 1 anterior right fragmentary fragment.

NMMNH P-39966: 1 right dentary.

NMMNH P-39967: 1 anterior half of left dentary.

NMMNH P-39968: 2 miscellaneous cranial fragment.

NMMNH P-40152: 1 right coracoid.

NMMNH P-40153: 1 right ilium.

NMMNH P-41743: 1 anterior cervical centrum.

NMMNH P-41744: 1 terminal phalanx (claw).

NMMNH P-41745: 1 proximal rig vragment.

NMMNH P-41746, P-41747: 1 dorsal vertebra.

NMMNH P-41748: Numerous caudal vertebrae.

NMMNH P-41749: 1 tiny cervical centrum.

NMMNH P-45029: 1 incomplete left scapulocoracoid.

NMMNH P-45030: 1 incomplete right scapulocoracoid.

NMMNH P-45031: 1 nearly complete right scapulocoracoid.

NMMNH P-45032: 1 right tibia.

NMMNH P-45033: 1 right femur.

NMMNH P-45034: 1 right humerus.

NMMNH P-45035: 1 left? tibia.

NMMNH P-45036: 1 left? fibula?

NMMNH P-45038: 1 metatarsal 1.

NMMNH P-45039: 1 small right ischium.

NMMNH P-45040: 1 dorsal vertebra.

NMMNH P-45041: 1 proximal caudal vertebra.

NMMNH P-45042: 1 small anterior dorsal vertebra.

NMMNH P-45568: 1 complete right dentary with teeth.

NMMNH P-45569: 4 associated bone fragments.

NMMNH P-50175: 1 concretaed porixmla humerous.

NMMNH P-50176: 1 phalanx.

NMMNH P-50177: 2 associated dorsal vertebrae.

NMMNH P-50178: 8 bone fragment.s

TMM 31099-37, 31099-38: Femora.

TMM 31099-39, 31099-40: Tibae.

TMM 31099-118: Mandible.

 


= Trilophosaurus jacobsi (sic?) POLCYN, WINKLER, JACOS & NEWMAN, 2002

Locality: SMU locality 252, vicinity of North Stinking Springs Mountian, what use to be Seven Springs Ranch, northwest of St. Johns, Apache County, Arizona.

Horizon: Lowe Petrified Forest Formation, Chinle Group.

Biostratigraphy:

Age: Late Carnian Stage, Late Triassic Epoch, Late Triassic.

Material:

Number: Not given:


_____________________________________________________________________________________

Species: jacobsi MURRY, 1987 emend HECKERT, LUCAS, RINEHART, SPIELMANN, & KAHLE, 2003
= Trilophosaurus? jacobsi MURRY, 1987
= Chinleogomphius jacobsi (MURRY, 1987) SUES & OLSEN, 1993
Etymology: In honor of Professor Louis L. Jacobs.

Holotype: MNA V3192

Locality: Placerias quarry (NMMNH locality 858, UCMP A269, MNA 207C), Big Hollow Wash near Romero Springs, a small valley 6 1/2 miles (approximately 10.4 km) S 65"W of St. Johns, Apache County, Arizona. (Located 0.3 miles northwest of Romero Springs at approximately Long. 109'28'W and lat. 34"27'N on the USGS St. Johns quadrangle of 1921.)

Horizon: Lower Bluewater Creek Formation (Lower Petrified Forest Formation).

Biostratigraphy:

Age: Otischalkian, Late Triassic (LUCAS, HECKET & HUNT, 1995) (Old Lower Petrified Forest Member, Chinle Formation, Adamanian, Late Triassic.)

Material: Left maxilla fragment.

Topotype:

MNA V3193: Left dentary fragment.

MNA V3194: Right maxilla fragment.

MNA V3197: Right maxilla fragment.

MNA V3198: Posterior right maxilla.

MNA V3199: Left dentary fragment.

Referred material:

SPIELMANN, LUCAS, RINEHART & HECKERT, 2008

MNA Pl.2924: Sacrals 1 and 2.

MNA Pl. 2956, Pl. 2928: Left and right ilia.
Note: All MNA specimens may represent a single individual).

UCMP 32390: Proximal two-thirds of left femur.

UCMP 35877: Distal extremity of left humerus.

UCMP 25825, 25839: Proximal and distal right tibiae.

UCMP 25826, 25798: Proximal and distal right tibiae.

UCMP 25790: Proximal right tibia.

UCMP 25424: Second sacral vertebra.


HECKERT, 2001

Locality: NMMNH 1430, at the head of the south fork of Home Creek north of Kalgary Locality, UTM 3701500N, 398850E, zone 14, NAD 27, Crosby County, Texas.

Horizon: Colorado City Member, Tecovas Formation, Chinle Group.

Biostratigraphy:

Age: Adamanian, Latest Carnian Stage, Late Triassic Epoch, Late Triassic.

Material:

NMMNH P-26421: Tooth.

NMMNH P-26442: Posterior

NMMNH P-34073: Tooth.

NMMNH P-34372: Tooth fragment.

 

Locality: NMMNH Locality 2739, Sixmile Springs, New Mexico.

Horizon: Bluewater Creek Formation, Lower Chinle Group.

Biostratigraphy:

Age: Adamanian, Latest Carnian Stage, Late Triassic Epoch, Late Triassic.

Material:

NMMNH P-34447: Tooth fragment.

 

MUELLER, HUTTENLOCKER, SMALL, PINTO, DEAN-WALLACE & CHATTERJEE, 2022

Locality: Boren Quarry (MOTT VPL 3869), Garza County, Texas.

Horizon: Tecovas Formation, Dockum Group.

Biostratigraphy: Otischalkian LVF.

Age: Early Norian Stage, Middle Late Triassic Epoch, Middle Late Triassic.

Material:

Number: Not given:

 

SPIELMANN, HECKERT & LUCAS, 2005

Locality: NMMNH Locality 3775 (Kahle Trilophosaurus quarry), West Texas.

Horizon: Trujillo Formation.

Biostratigraphy:

Age: Late Triassic.

Material:

NMMNH P-39917: Right femur.

NMMNH P-39936: Left humerus.

NMMNH P-39937: Left humeral head.

HECKERT, LUCAS, RINEHART, SPIELMANN, HUNT & KAHLES, 2006

NMMNH P-41400: Incomplete skull and mandible of a juvenile.

SPIELMANN, LUCAS, RINEHART & HECKERT, 2008

NMMNH P-29959: 1 maxillary? fragment.

NMMNH P-29960: 1 tooth fragment.

NMMNH P-29961: 2 complete caudal vertebrae.

NMMNH P-29962: 1 complete vertebra.

NMMNH P-29963: 1 proximal humerus? fragment.

NMMNH P-29964: 1 distal humerus.

NMMNH P-29965: 2 distal ulnae.

NMMNH P-29966: 4 proximal radii.

NMMNH P-29967: 1 carpal (I or V).

NMMNH P-29968: 1 poximal femur.

NMMNH P-29969: 1 distal femur.

NMMNH P-29970: 1 astragalus.

NMMNH P-29971: 2 terminal phalanges.

NMMNH P-29972: 1 carpal.

NMMNH P-29973: 1 phalanx.

NMMNH P-29974: 3 podials.

NMMNH P-29975: 1 end of metopodial.

NMMNH P-34101: 1 left humerus.

NMMNH P-34102: 1 tibia.

NMMNH P-34103: 1 caudal vertebra.

NMMNH P-34104: 2 femur.

NMMNH P-34105: 1 caudal vertebra.

NMMNH P-34113: 2 terminal phalanges.

NMMNH P-34122: 2 caudal centra.

NMMNH P-36699: 1 left calcaneum.

NMMNH P-36700, 36701: 1 right calcaneum.

NMMNH P-36702: 1 right pubis.

NMMNH P-36703-P36706: 1 left pubis.

NMMNH P-36706: 1 right ulnae.

NMMNH P-36708-P36719: 1 right astragalus.

NMMNH P-36720-P-36728: 1 left astrgalus.

NMMNH P-36729: 1 astragalus fragment.

NMMNH P-36730-P-36737: 1 right radaliae (?).

NMMNH P-36738-P-36744: 1 left radaliae?

NMMNH P-36745-P-36751: 1 right first distal tarsal?

NMMNH P-36752-P-36760: 1 left first distal tarsal?

NMMNH P-36761-P-36764: 1 right ulnae?

NMMNH P-36765-36771: 1 left ulnae?

NMMNH P-36772: 1 right calcaneum.

NMMNH P-36773, P-36774: 1 left third carpal.

NMMNH P-36775-P-36802: Mesopodial.

NMMNH P-36803: 1 proximal phalanx.

NMMNH P-36804: 3 intercentra.

NMMNH P-36807: Numerous carpals/tarsals.

NMMNH P-36808: 1 right maxillary fragment with 6 teeth.

NMMNH P-36809: 1 dentary fragment with 4 teeth.

NMMNH P-36810: 1 dentary fragment with 8 teeth.

NMMNH P-36811: 1 right maxillary fragment with 11 teeth.

NMMNH P-36812: 1 right maxillary? fragment with 7 teeth.

NMMNH P-39880: 1 left distal humerus.

NMMNH P-39881, P-39882: Left proximal humerus.

NMMNH P-39883, P-39884: Left distal humerus.

NMMNH P-39887: 1 left distal humerus.

NMMNH P-39888: 1 left proximal humerus.

NMMNH P-39889-P-28981: Right proximal hemerus.

NMMNH P-39892-38999: 1 right proximal tibia.

NMMNH P-39000: 1 right tibia.

NMMNH P-39901: 1 right femur.

NMMNH P-39904: 1 left proximal femur.

NMMNH P-39905: 1 left distal femur.

NMMNH P-39906-P-39908: 1 left proximal femur.

NMMNH P-39909-P-39910: 1 left distal femur.

NMMNH P-39911: 1 left proximal femur.

NMMNH P-39912: 1 right distal femur.

NMMNH P-39913: 1 left distal femur.

NMMNH P-39914-P-39916: 1 right distal femur.

NMMNH P-39918, P-39919: 1 right distal femur.

NMMNH P-39920-P-29922: 1 right distal femur.

NMMNH P-39924: 1 right proximal fibula.

NMMNH P-39925-P-39927: 1 left proximal fibula.

NMMNH P-39928, P-39929: 1 distal fibula.

NMMNH P-39930: 1 right proximal fibula.

NMMNH P-39931: 1 left proximal fibula.

NMMNH P-39932, P-39933: 1 distal fibula.

NMMNH P-39934: 1 left humerus.

NMMNH P-39935: 1 right proximal humerus.

NMMNH P-39938: 1 right humerus.

NMMNH P-39969: 1 left tibia.

NMMNH P-39970: 2 left proximal tibiae.

NMMNH P-39971: 1 right proximal tibia.

NMMNH P-39972-P-39980: 1 left proximal tibia.

NMMNH P-39981: 1 right tibia.

NMMNH P-39982, P-39983: 1 left tibia.

NMMNH P-39984: 1 left proximal tibia.

NMMNH P-39985: 1 left tibia.

NMMNH P-39986: 1 proximal tibia.

NMMNH P-39987, P-39988: 1 right distal humerus.

NMMNH P-39989-P-39992: 1 right proximal humerus.

NMMNH P-39993: 1 distal humerus.

NMMNH P-39994: 1 humeral shaft.

NMMNH P-39995: I right ilium.

NMMNH P-39996: 1 left ilium fragment.

NMMNH P-39997: 1 distal humerus.

NMMNH P-39998, P-39999: 1 left proximal humerus.

NMMNH P-40001: 3 humeral fragments.

NMMNH P-40002: 1 bone fragment.

NMMNH P-40003-P-40005: 1 left proximal fibula.

NMMNH P-40006: 4 fibula fragments.

NMMNH P-40007: 1 left proximal fibula.

NMMNH P-40008: 1 fibular shaft.

NMMNH P-40009: 1 proximal fibula.

NMMNH P-40010: 1 distal fibula.

NMMNH P-40011: 1 left distal humerus.

NMMNH P-40084, P-40085: 1 left femur.

NMMNH P-40086: 1 left proximal femur.

NMMNH P-40087: 1 left distal femur.

NMMNH P-40088: 1 left proximal femur.

NMMNH P-40089-P-40091: 1 distal femur.

NMMNH P-40092: 1 right distal tibia.

NMMNH P-40093: 2 right proximal tibia fragments.

NMMNH P-40094-P-40097: 1 right distal tibia.

NMMNH P-40098-P-40102: 1 left distal tibia.

NMMNH P-40103, P-40104: 1 left proximal tibia.

NMMNH P-40105: 8 distal tibia fragments.

NMMNH P-40106-P-40109: 1 distal tibia.

NMMNH P-40110: 1 proximal? tibia.

NMMNH P-40111: 1 distal tibia.

NMMNH P-40112: 1 distal? tibia.

NMMNH P-40113: 3 tibia fragments.

NMMNH P-40114: 1 distal femur.

NMMNH P-40115: 1 left proximal femur.

NMMNH P-40116: 1 proximal femur.

NMMNH P-40117-P-40141: 2 terminal phalanx (claw).

NMMNH P-43258: 1 distal left ulna fragment.

NMMNH P-43259, P-43260: 1 proximal left ulna fragment.

NMMNH P-43261: 1 complete left ulna.

NMMNH P-43262: 2 distal right? ulna fragment.

NMMNH P-43263: 1 proximal right? ulna fragment.

NMMNH P-43264: 1 distal left ulna fragment.

NMMNH P-43265: 2 distal right ulna fragment.

NMMNH P-43266: 1 distal ulna fragment.

NMMNH P-43267: 1 left radilae.

NMMNH P-43267: 1 proximal left metatarsal IV fragment.

NMMNH P-43269, P-43270: 1 distal fragment pedal phalanx.

NMMNH P-43271: 1 proximal fragment of metatarsal IV.

NMMNH P-43272: 1 left pubis.

NMMNH P-43273: 1 left radiale.

NMMNH P-43274: 1 proximal fragment of pedal phalanx.

NMMNH P-43275: 1 distal fragment of manual phalanx.

NMMNH P-43276: 1 proximal metatarsal fragment.

NMMNH P-43277: 1 right metatarsal 1.

NMMNH P-43278: 1 right astragalus.

NMMNH P-43279: 1 left first distal tarsal.

NMMNH P-43280: 1 proximal fragment of manual phalanx.

NMMNH P-43281: 1 distal fragment of right metatarsal IV.

NMMNH P-43282: 1 right manual phalanx of digit 2.

NMMNH P-43282: 1 tooth.

NMMNH P-43283: 1 tooth.

NMMNH P-43284: 1 left astragalus.

NMMNH P-43285: 1 distal left ulna.

NMMNH P-43286: 1 left proximal ulna.

NMMNH P-43287: 1 proximal left? ulna shaft.

NMMNH P-43288: 1 distal left ulna.

NMMNH P-43289: 1 proximal right ulna.

NMMNH P-43290: 1 distal left ulna.

NMMNH P-43291: 2 proximal right ulna.

NMMNH P-43293-P-43295: 1 proximal right ulna.

NMMNH P-43296: 1 distal right ulna.

NMMNH P-43297: 1 proximal right ulna.

NMMNH P-43298: 1 distal left ulna.

NMMNH P-43299: 1 right? ulna shaft.

NMMNH P-43300: 1 proximal right ulna.

NMMNH P-43301: 1 proximal left ulna.

NMMNH P-43302: 1 distal left ulna.

NMMNH P-43559: 5 caudal vertebrae.

NMMNH P-43560: 1 left scapula fragment.

NMMNH P-43561: 1 left ulna missing distal end.

NMMNH P-43562: 2 miscellaneous bone fragments.

NMMNH P-43563: 1 dorsal vertebra.

NMMNH P-43564: 1 cervical vertebra.

NMMNH P-43565: 1 large caudal vertebra.

NMMNH P-43566: 1 caudal centrum fragment.

NMMNH P-43567: 1 caudal vertebra.

NMMNH P-43569: 1 centrum fragment.

NMMNH P-44274: 1 posterior cervical vertebra.

NMMNH P-44275: 1 anterior cervical vertebra.

NMMNH P-44276: 1 posterior cervical vertebra.

NMMNH P-44277: 1 anterior caudal vertebra.

NMMNH P-44278: 1 right pubis.

NMMNH P-44279: 1 right coracoid.

NMMNH P-44280: 1 phalanx 2 of digit 2 of left manus.

NMMNH P-44281: 1 phalanx 1 of digit 3 of left manus.

NMMNH P-44282: 1 left carpal.

NMMNH P-44283: 1 left carpal II.

NMMNH P-44284: 1 left radiale.

NMMNH P-44285: 1 right ilium.

NMMNH P-44286: 1 left metacarpal.

NMMNH P-44287: 1 right tarsal IV.

NMMNH P-44288: 1 right metatarsal IV.

NMMNH P-44289: 1 large sacral vertebra.

NMMNH P-45021: 17 caudal vertebrae.

NMMNH P-45022: 1 sacral vertebra.

NMMNH P-45023: 1 caucal vertebra.

NMMNH P-45024: 1 caudal vertebra found with ischium.

NMMNH P-45025: 1 right ischium.

NMMNH P-45026- P-45028: 1 proximal caudal vertebra.

NMMNH P-45566: 1 right maxillary? fragment with 8 teeth.

NMMNH P-45567: 1 left maxillary? fragment with 8 teeth.

NMMNH P-46131: 1 sacral vertebrae in two pieces.

NMMNH P-50164, P-50165: 1 right pubis.

NMMNH P-50166: 1 partial right ilium.

NMMNH P-50167: 1 left coracoid.

NMMNH P-50168: 1 left pubis.

NMMNH P-50169: 1 proximal humerus.

NMMNH P-50170: 2 tibial shafts.

NMMNH P-50171: 1 anterior right maxilla and associated fragments.

NMMNH P-50172: 1 partial phalanx.

NMMNH P-54141: 4 jaw fragments.

 

= Trilophosaurus cf. T. jacobsi SPIELMANN, LUCAS, HECKERT & KRZYZANOWSKI, 2013

Locality: NMMNH Locality 3764, Krzyzanowski Site, Arizona.

Horizon: Lower Blue Mesa Member, Petrified Forest Formation, Chinle Group.

Biostratigraphy:

Age: Adamanian, Latest Carnian Stage, Late Triassic Epoch, Late Triassic.

Material:

NMMNH P-40937: Dorsal vertebra.

NMMNH P-40939: Partial dorsal vertebra, conglomerated mass of vertebrae including a cervical vertebra and small left ilium.

NMMNH P-42189: Right astragalus.

NMMNH P-44164: Left tibia.

NMMNH P-44174: Left pelvic girdle with a nearly complete ilium adn incomplete ischium.

NMMNH P-60208: Partial cervical vertebra, distal right humerus, right radius, and left and right femora.

NMMNH P-63571: Incomplete left femur.

NMMNH P-63572: Right ulna.

_____________________________________________________________________________________

Species: dornorum MUELLER & PARKER, 2006
Etymology: In honor of PEFO Chief of Resource Management Karen Dorn, under who the paleontology program of the park has been revitalized; and Chuck Dorn, a PEFO Law Enforcement ranger, for his ongoing protection of fossil resources.
= Trilophosaurus sp IRMIS, 2005
= Trilophosaurus sp PARKER, 2005
= Trilophosaurus sp PARKER & IRMIS, 2005

Holotype: PEFO 31165

Locality: PFV 191, Flag Canyon Locality, Petrified Forest National Park, Apache County, Arizona.

Horizon: Sonsel Member, Petrified Forest Formation, Chinle Group.

Biostratigraphy:

Age: Late Carnian-Early Norian Stage, Late Triassic Epoch, Late Triassic.

Material: Partial left maxillary with teeth and partial left dentary with teeth.

Referred material:

Locality: MOTT VPL 3624, Post Quarry, lat 33°31’17”N, long. 101°18’54”W, 14.5 km southeast of Post, R. C. Miller Ranch, Garza County, Texas.

Horizon: Cooper Canyon Member, Dockum Formation.

Biostratigraphy:

Age: Carnian Epoch, Middle Late Triassic Epoch, Late Triassic.

Material:

TTU-P09497: Isolated tooth.

 

Locality: MOTT VPL 3869, Garza County, Texas.

Horizon: Cooper Canyon Member, Dockum Formation.

Biostratigraphy:

Age: Carnian Epoch, Middle Late Triassic Epoch, Late Triassic.

Material:

TTU-P10413: Partial right maxilla.

TTU-P10582: Partial right maxilla.

TTU-P10583: Partial left maxilla.

TTU-P10586: Right and left dentary fragments.

 

Locality: MOTT VPL 3878, Garza County, Texas.

Horizon: Cooper Canyon Member, Dockum Formation.

Biostratigraphy:

Age: Carnian Epoch, Middle Late Triassic Epoch, Late Triassic.

Material:

TTU-P10447: Partial maxilla.

 

MUELLER, HUTTENLOCKER, SMALL, PINTO, DEAN-WALLACE & CHATTERJEE, 2022

Locality: Boren Quarry (MOTT VPL 3869), Garza County, Texas.

Horizon: Tecovas Formation, Dockum Group.

Biostratigraphy: Otischalkian LVF.

Age: Early Norian Stage, Middle Late Triassic Epoch, Middle Late Triassic.

Material:

Number: Not given:


_____________________________________________________________________________________

Species: phasmalophos KLIGMAN, MARSH, NESBITT, PARKER & STOCKER, 2020
Etymology: Greek, phasma, "phantom or ghost" and Greek, lopho, "crest," referring to the unique two cusped condition of this trilophosaurid taxon.

Holotype: PEFO 42082

Locality: PFV 089 (the Bowman Site), Petrified Forest National Park, Apache County, Arizona.

Horizon: Upper Sonsela Member.

Biostratigraphy: Lower part of the Jim Camp Wash Beds, Revueltian land vertebrtae holochronozone.

Age: Norian Stage, Late Triassic Epoch, Late Triassic.

Material: Isoalted tooth from the maxilla or dentary.

Paratypes:

PEFO 42117, 42167, 42209, 42231, 42261, 42271, 42278, 42292: Isolated teeth.

Referred material:

DMNH PAL 2018-05-0012, -0013: Isolated teeth.