Genus: Endothiodon OWEN, 1876
Etymology: Greek, endothi, “within” and Greek, odon, “tooth”: internal tooth.
= Emydochampsa BROOM, 1912
Etymology: Greek, emydos, (emys), “turtle” and Greek, khampsa, “crocodile”: Turtle crocodile.
= Endogomphodon BROOM, 1932
Etymology: Greek, endon, “within”, Greek, gomphios, “molar” and Greek, odon, “tooth”: Internal molar-tooth.
= Esoterodon SEELEY, 1895
Etymology: Greek, esoteros, “interior, inner” and Greek, odon, “tooth”: Interior tooth.
= Pachytegos HAUGHTON, 1932
Etymology: Greek, pakhys, “thick” and Greek, tegos, “roof”: Thick roof.= Prodicynodon BROOM, 1904
Etymology: Greek, pro-, “before”, and Dicynodon: Before Dicynodon.

Species: bathystoma OWEN, 1876
Etymology: Latin, bathys, "deep" and Greek, stoma, "mouth."

Holotype: BMNH R1646

Locality: Sneewberg range, Graaff Reinet district, Eastern Cape Province, South Africa.

Horizon: Hoedemaker member, Middle Teekloof Formation.

Biostratigraphy: Tropidostoma zone (Tropidostoma-Endothidon Assemblage zone KEYSER, 1979, SACS, 1980, Tropidostoma microtrema zone KEYSER & SMITH, 1977-78, Lower Cistecephalus zone KITCHING, 1970, 1977, Endothiodon zone, WATSON, 1914).

Age: Tatarian Age, Lopingian Subepoch, Zechstein Epoch, Late Permian.

Material: Anterior half of skull and mandible.

Referred material:

ANTUNES, 1975

Locality: Near Maniamba, Mozambique, South Africa.

Horizon: Lunho Series.

Biostratigraphy:

Age: Late Permian.

Material:

Numbers: Not given: Fragments of skulls and skeletons.

 

= Endothiodon cf. bathystoma ARAUJO, FERNANDEZ, RABBITT, EKDALE, ANTUNES, CASTANHINHA, FRÖBISCH

& MARTINS, 2018

Locality: Metangula Graben, Niassa Province, Mozambique, South Africa.

Horizon: K5 Formation.

Biostratigraphy:

Age: Late Permian.

Material:

= Endothiodon cf. E. bathystoma BARBERENA & ARAUJO, 1975, BOOS, SCHULTZ, VEGA & AUMOND, 2013

Locality: A rail way cut near Serra do Cadeado area, Parana State, Brazil.

Horizon: Rio do Rasto Formation.

Biostratigraphy:

Age: Ufimian to Early Tatarian age, Guadelupian Stage, Middle Zeichstein Epoch, Middle Permian.

Material:

FURB PV0226P: Fragmentary skull and fragmentary lower jaw.

Referred material:

Number: Not given: Fragmentary skeleton.

 

BROOM, 1915

Locality: Beaufort West, Western Cape Province, South Africa.

Horizon: Hoedemaker member, Middle Teekloof Formation.

Biostratigraphy: Tropidostoma zone (Tropidostoma-Endothidon Assemblage zone KEYSER, 1979, SACS, 1980, Tropidostoma microtrema zone KEYSER & SMITH, 1977-78, Lower Cistecephalus zone KITCHING, 1970, 1977, Endothiodon zone, WATSON, 1914).

Age: Tatarian Age, Lopingian Subepoch, Zechstein Epoch, Late Permian.

Material:

AMNH 5614: Skull, mandible, and cervical vertebrae.

 

COX, 1964

Locality: South Africa.

Horizon: Hoedemaker member, Middle Teekloof Formation.

Biostratigraphy: Tropidostoma zone (Tropidostoma-Endothidon Assemblage zone KEYSER, 1979, SACS, 1980, Tropidostoma microtrema zone KEYSER & SMITH, 1977-78, Lower Cistecephalus zone KITCHING, 1970, 1977, Endothiodon zone, WATSON, 1914).

Age: Tatarian Age, Lopingian Subepoch, Zechstein Epoch, Late Permian.

Material:

AMNH 5614, BMNH R4142: Fragmentary skulls.

BMNH R1646: Skull.

 

HAUGHTON, 1932

Locality: B32, below Usili Mt, near Ngaka-Ruhuhu confluence, Tanganyika, East Africa.

Horizon: Kawinga Formation.

Biostratigraphy: Cistecephalus zone (formerly Auacephalodon-Cistecephalus zone KEYSER, 1979, SACS, 1980, Aulacephalodon baini zone KEYSER & SMITH, 1977-78, Upper Cistecephalus Zone, KITCHING, 1970, 77, Lower Cistecephalus zone WATSON, 1914).

Age: Tatarian Age, Lopingian Subepoch, Zechstein Epoch, Late Permian.

Material:

Number: Not given: Fragmentary right ramus.

 

LYDEKKER, 1890

Locality: Gouph district near Beaufort West, Western Cape Province, South Africa.

Horizon:

Biostratigraphy:

Age: Late Permian.

Material:

BMNH 49415: Anterior portion of a mandible.

 

SEELEY, 1892

Locality: Oude Kloof, Nieuwveldt Mountains, Cape Colony, South Africa.

Horizon: Hoedemaker member, Middle Teekloof Formation.

Biostratigraphy: Tropidostoma zone (Tropidostoma-Endothidon Assemblage zone KEYSER, 1979, SACS, 1980, Tropidostoma microtrema zone KEYSER & SMITH, 1977-78, Lower Cistecephalus zone KITCHING, 1970, 1977, Endothiodon zone, WATSON, 1914).

Age: Tatarian Age, Lopingian Subepoch, Zechstein Epoch, Late Permian.

Material:

Number: Not given: Nearly complete left dentary and a left squamosal.

 

= Emydochampsa oweni BROILI & SCHRODER, 1936
Etymology: In honor of Richard Owen (1804-1892), British anatomist and Paleontologist
= Endothiodon oweni (BROILI & SCHRODER, 1936)
= Endogomphodon oweni (BROILI & SCHRODER, 1936)

Holotype: Munich 1934 viii 31

Locality: Farm Lombaardskraal, 12 miles south of Beaufort West, Western Cape Province, South Africa.

Horizon: Hoedemaker member, Middle Teekloof Formation.

Biostratigraphy: Tropidostoma zone (Tropidostoma-Endothidon Assemblage zone KEYSER, 1979, SACS, 1980, Tropidostoma microtrema zone KEYSER & SMITH, 1977-78, Lower Cistecephalus zone KITCHING, 1970, 1977, Endothiodon zone, WATSON, 1914).

Age: Tatarian Age, Lopingian Subepoch, Zechstein Epoch, Late Permian.

Material: Skull and lower jaw.

= Endothiodon uniseries OWEN, 1879
Etymology:
= Esoterodon uniseries (OWEN, 1879) SEELEY, 1895

Holotype: BMNH 49414

Locality: On the flanks of the Nieuwveldt range, Beaufort West, Western Cape Province, South Africa.

Horizon: Hoedemaker member, Middle Teekloof Formation.

Biostratigraphy: Tropidostoma zone (Tropidostoma-Endothidon Assemblage zone KEYSER, 1979, SACS, 1980, Tropidostoma microtrema zone KEYSER & SMITH, 1977-78, Lower Cistecephalus zone KITCHING, 1970, 1977, Endothiodon zone, WATSON, 1914).

Age: Tatarian Age, Lopingian Subepoch, Zechstein Epoch, Late Permian.

Material: Anterior half of skull.

Referred material:

BROOM, 1915

Locality: Beaufort West, Western Cape Province, South Africa.

Horizon: Hoedemaker member, Middle Teekloof Formation.

Biostratigraphy: Tropidostoma zone (Tropidostoma-Endothidon Assemblage zone KEYSER, 1979, SACS, 1980, Tropidostoma microtrema zone KEYSER & SMITH, 1977-78, Lower Cistecephalus zone KITCHING, 1970, 1977, Endothiodon zone, WATSON, 1914).

Age: Tatarian Age, Lopingian Subepoch, Zechstein Epoch, Late Permian.

Material:

AMNH 5612: Skull.

AMNH 5613: Fragmentary skull and skeleton.

AMNH 5334: Skull and skeleton.

 

WATSON, 1949

Locality: Beaufort West, Western Cape Province, South Africa.

Horizon: Hoedemaker member, Middle Teekloof Formation.

Biostratigraphy: Tropidostoma zone (Tropidostoma-Endothidon Assemblage zone KEYSER, 1979, SACS, 1980, Tropidostoma microtrema zone KEYSER & SMITH, 1977-78, Lower Cistecephalus zone KITCHING, 1970, 1977, Endothiodon zone, WATSON, 1914).

Age: Tatarian Age, Lopingian Subepoch, Zechstein Epoch, Late Permian.

Material:

R 79? Fragmentary skull.

BMNH R 4044: Anterior end of right lower jaw.

AMNH 5615: Articular.

 

= Endothiodon platyceps BROOM, 1912
Etymology: Greek, platy, "broad, flat, wide" and Latin, -ceps, "head."
= Emydochampsa platyceps (BROOM, 1912)

Holotype: AMNH 5570

Locality: Beaufort West, Western Cape Province, South Africa.

Horizon: Hoedemaker member, Middle Teekloof Formation.

Biostratigraphy: Tropidostoma zone (Tropidostoma-Endothidon Assemblage zone KEYSER, 1979, SACS, 1980, Tropidostoma microtrema zone KEYSER & SMITH, 1977-78, Lower Cistecephalus zone KITCHING, 1970, 1977, Endothiodon zone, WATSON, 1914).

Age: Tatarian Age, Lopingian Subepoch, Zechstein Epoch, Late Permian.

Material: Skull.

Referred material:

AMNH 5571: Skull.

 

RAY & BANDYOPADHYAY, 2003

Locality: Near Golet, Adilabad District, Andhra Pradesh State, India.

Horizon: Lower Gondwana Kundaram Formation.

Biostratigraphy: Equivalent to the Tropidostoma zone (Tropidostoma-Endothidon Assemblage zone KEYSER, 1979, SACS, 1980, Tropidostoma microtrema zone KEYSER & SMITH, 1977-78, Lower Cistecephalus zone KITCHING, 1970, 1977, Endothiodon zone, WATSON, 1914).

Age: Tatarian Age, Lopingian Subepoch, Zechstein Epoch, Late Permian.

Material:

ISIR 208: Partial skull and lower jaw.

 

= Endothiodon seeleyi BROOM, 1915
Etymology: In honor of H. G. Seeley.

Holotype: AMNH 5615 (BMNH 1969?)

Locality: Oude Kloof, Beaufort West, Western Cape Province, South Africa.

Horizon: Hoedemaker member, Middle Teekloof Formation.

Biostratigraphy: Tropidostoma zone (Tropidostoma-Endothidon Assemblage zone KEYSER, 1979, SACS, 1980, Tropidostoma microtrema zone KEYSER & SMITH, 1977-78, Lower Cistecephalus zone KITCHING, 1970, 1977, Endothiodon zone, WATSON, 1914).

Age: Tatarian Age, Lopingian Subepoch, Zechstein Epoch, Late Permian.

Material: Mandibles.

 

= Endothiodon paucidens BROOM, 1915
Etymology:
= Esoterodon paucidens (BROOM, 1915)

Holotype: AMNH 5572

Locality: Beaufort West, Western Cape Province, South Africa.

Horizon: Hoedemaker member, Middle Teekloof Formation.

Biostratigraphy: Tropidostoma zone (Tropidostoma-Endothidon Assemblage zone KEYSER, 1979, SACS, 1980, Tropidostoma microtrema zone KEYSER & SMITH, 1977-78, Lower Cistecephalus zone KITCHING, 1970, 1977, Endothiodon zone, WATSON, 1914).

Age: Tatarian Age, Lopingian Subepoch, Zechstein Epoch, Late Permian.

Material: Skull.

Referred material:

AMNH 5573: Skull.

AMNH 5574: Fragmentary skeleton.

 

= Endothiodon angusticeps BROOM, 1915
Etymology:
= Esoterodon angusticeps (BROOM, 1915)

Holotype: AMNH 5612

Locality: Beaufort West, Western Cape Province, South Africa.

Horizon: Hoedemaker member, Middle Teekloof Formation.

Biostratigraphy: Tropidostoma zone (Tropidostoma-Endothidon Assemblage zone KEYSER, 1979, SACS, 1980, Tropidostoma microtrema zone KEYSER & SMITH, 1977-78, Lower Cistecephalus zone KITCHING, 1970, 1977, Endothiodon zone, WATSON, 1914).

Age: Tatarian Age, Lopingian Subepoch, Zechstein Epoch, Late Permian.

Material: Skull.

 

= Endogomphodon minor BROOM, 1932
Etymology: Latin, minor, "minor, small."
= Endothiodon minor (BROOM, 1932)

Holotype: SAM 2676

Locality: Beaufort West Flats, Western Cape Province, South Africa.

Horizon: Hoedemaker member, Middle Teekloof Formation.

Biostratigraphy: Tropidostoma zone (Tropidostoma-Endothidon Assemblage zone KEYSER, 1979, SACS, 1980, Tropidostoma microtrema zone KEYSER & SMITH, 1977-78, Lower Cistecephalus zone KITCHING, 1970, 1977, Endothiodon zone, WATSON, 1914).

Age: Tatarian Age, Lopingian Subepoch, Zechstein Epoch, Late Permian.

Material: Skull.

= Endothiodon whaitsi BROOM 1912
Etymology: In honor of Rev. John H. Whaits (1870-1947).
= Esoterodon whaitsi (BROOM, 1912)

Holotype: AMNH 5565

Locality: Beaufort West, Western Cape Province, South Africa.

Horizon: Hoedemaker member, Middle Teekloof Formation.

Biostratigraphy: Tropidostoma zone (Tropidostoma-Endothidon Assemblage zone KEYSER, 1979, SACS, 1980, Tropidostoma microtrema zone KEYSER & SMITH, 1977-78, Lower Cistecephalus zone KITCHING, 1970, 1977, Endothiodon zone, WATSON, 1914).

Age: Tatarian Age, Lopingian Subepoch, Zechstein Epoch, Late Permian.

Material: Skull, mandible, a few vertebrae, ribs and limb bones.

Referred material:

= Endothiodon crassus BROOM, 1932
Etymology: Latin, crassus, "dense, thick."
= Endogomphodon crassus (BROOM, 1932)

Holotype: Kimberly Museum, McBMK 4236

Locality: Dunedin, Beaufort West District, South Africa.

Horizon: Hoedemaker member, Middle Teekloof Formation.

Biostratigraphy: Tropidostoma zone (Tropidostoma-Endothidon Assemblage zone KEYSER, 1979, SACS, 1980, Tropidostoma microtrema zone KEYSER & SMITH, 1977-78, Lower Cistecephalus zone KITCHING, 1970, 1977, Endothiodon zone, WATSON, 1914).

Age: Tatarian Age, Lopingian Subepoch, Zechstein Epoch, Late Permian.

Material: Skull.

= Pachytegos stockleyi HAUGHTON, 1932
Etymology:

Holotype: SAM 10639 (Stockley number S559)

Locality: B32, below Usili Mt, near Ngaka-Ruhuhu Valley, Tanzania, East Africa.

Horizon: Kawinga Formation.

Biostratigraphy: Cistecephalus zone (formerly Auacephalodon-Cistecephalus zone KEYSER, 1979, SACS, 1980, Aulacephalodon baini zone KEYSER & SMITH, 1977-78, Upper Cistecephalus Zone, KITCHING, 1970, 77, Lower Cistecephalus zone WATSON, 1914).

Age: Tatarian Age, Lopingian Subepoch, Zechstein Epoch, Late Permian.

Referred material:

GAY & CRUICKSHANK, 1999

SAM 10642 (Stockley number S559): Pineal region.

_____________________________________________________________________________________

Species: mahalanobisi RAY, 2000
Etymology: In honor of Professor Prasanta Chandra Mahalanobis, the founder of the Indian Statistical Institute who initiated systematic vertebrate Paleontological research in the Geological Studies Unit of ISI.

Holotype: ISI R201

Locality: Localities I and II, near Golet, Adilabad District, Andhra Pradesh State, India.

Horizon: Lower Gondwana, Kundaram Formation.

Biostratigraphy: Equivalent to the Tropidostoma zone (Tropidostoma-Endothidon Assemblage zone KEYSER, 1979, SACS, 1980, Tropidostoma microtrema zone KEYSER & SMITH, 1977-78, Lower Cistecephalus zone KITCHING, 1970, 1977, Endothiodon zone, WATSON, 1914).

Age: Tatarian Age, Lopingian Subepoch, Zechstein Epoch, Late Permian.

Material: A nearly complete skull with an associated jaw.

Paratypes:

ISI R214: A nearly complete skull except the squamosals.

ISI R202, R211-215, R218, R220, R340, R342-343, R345-347, R359: Several snouts and anterior parts of skull.

ISI R341, R344: Anterior part of skulls and assocated lower jaws.

ISI R219, R356-357: Posterior parts of skull.

ISI R206-207, R216, R348-350, R352, R354-355, R358, R360: Several fragments of lower jaw.

_____________________________________________________________________________________

Species: tolani COX & ANGIELCZYK, 2015
Etymology: In honor of Steve Tolan of the Chipembele Wildlife Education Trust (Mfuwe, Luangwa Valley, Zambia), as a token of thanks for putting the authors in touch with one another, and in recognition of his activites in advancing research on the paleontological heritage of Zambia and combating the illegal fossil tade in the country.

Holotype: NHMUK PV R12443

Locality: Locality L4, close to the heads of the Matamondo and Rupaha streams, about 5.5 km north of the village of Litumba Ndyosi, Ruhuhu Basin, Tanzania.

Horizon: Ruhuhu Formation.

Biostatigraphy:

Age: Late Permian.

Material: A partial skull and complete mandible.

Paratype: NHMUK PV R12444: The anterior portion of the skull wiht pterygoid region and occipital portions of the right and left squamosals.

Referred material:

NHMUK PV R12446, PV R12447, PV R12449, PV R12452:

 

Locality: Locality L2, close to the heads of the Matamondo and Rupaha streams, about 5.5 km north of the village of Litumba Ndyosi, Ruhuhu Basin, Tanzania.

Horizon: Ruhuhu Formation.

Biostatigraphy:

Age: Late Permian.

Material:

NHMUK PV R12248, PV R12350:

 

Locality: Locality L3, close to the heads of the Matamondo and Rupaha streams, about 5.5 km north of the village of Litumba Ndyosi, Ruhuhu Basin, Tanzania.

Horizon: Ruhuhu Formation.

Biostatigraphy:

Age: Late Permian.

NHMUK PV R12445: A smaller fragmentary skull consisting of right and left parts of snout, frontal region back to the anterior margin of the pineal foramen, median part of pterygoids and brancase back to occipital condyles, occipital portion of left squamosal, occipital portion and the base of the zygomatic process of the right squamosal, and the bases of right and left postorbital bars with suborbital bosses.

NHMUK PV R122443, PV R12451:

_____________________________________________________________________________________

Endothidon sp (Note: The following species are considered to be Endothidon sp, though they already have a species name and should keep the species name)

Species: sp (peartonensis (BROOM, 1904) (nomen dubium))
= Prodicynodon peartonensis BROOM, 1904
Etymology: In reference to Pearston, Eastern Cape Province, South Africa.

Holotype: Albany Museum 2551

Locality: Pearston, Eastern Cape Province, South Africa.

Horizon: Hoedemaker member, Middle Teekloof Formation.

Biostratigraphy: Tropidostoma zone (Tropidostoma-Endothidon Assemblage zone KEYSER, 1979, SACS, 1980, Tropidostoma microtrema zone KEYSER & SMITH, 1977-78, Lower Cistecephalus zone KITCHING, 1970, 1977, Endothiodon zone, WATSON, 1914).

Age: Tatarian Age, Lopingian Subepoch, Zechstein Epoch, Late Permian.

Material: Imperfect skull.
_____________________________________________________________________________________

Species: sp (beaufortensis (BROOM, 1912) (nomen dubium))
= Prodicynodon beaufortensis BROOM, 1912
Etymology: In reference to Beaufort West, Western Cape Province, South Africa.

Holotype: AMNH 5509

Locality: Kuilspoort, Beaufort West, Western Cape Province, South Africa.

Horizon: Hoedemaker member, Middle Teekloof Formation.

Biostratigraphy: Tropidostoma zone (Tropidostoma-Endothidon Assemblage zone KEYSER, 1979, SACS, 1980, Tropidostoma microtrema zone KEYSER & SMITH, 1977-78, Lower Cistecephalus zone KITCHING, 1970, 1977, Endothiodon zone, WATSON, 1914).

Age: Tatarian Age, Lopingian Subepoch, Zechstein Epoch, Late Permian.

Material: Fragmentary skull.