Species: declivis (OWEN, 1860)
Etymology:
= Dicynodon declivis OWEN, 1860
= Ptychognathus declivis (OWEN, 1860) OWEN, 1876
= Ptychosiagum declive (OWEN, 1860) LYDEKKER in NICHOLSON & LYDEKKER, 1889

Holotype: BMNH 36221

Locality: Rhenosterberg, Graaff Reinet District, Eastern Cape Province, South Africa.

Horizon: Katberg Formation, Beaufort Group.

Biostratigraphy: Lystrosaurus Assemblage zone (Lystrosaurus-Thrinaxodon zone, KEYSER, 1979, SACS, 1980, Procolophon zone, Lystrosaurus zone, WATSON, 1914, BROOM, 1906).

Age: Early Scythian Epoch, Early Triassic.

Material: Skull.

Modified from Colbert, 1974.

Referred material:

BOTHA-BRINK, HUTTENLOCKER, & MODESTO, 2014

Locality: Nooitgedacht 68, Bethulie District,  Eastern Cape Province, South Africa.

Horizon: Palingkloof Member, Balfour Formation.

Biostratigraphy: Lystrosaurus Assemblage Zone.

Age: Early Scythian Epoch, Early Triassic.

Material:

BP/1/3990: Skull and complete skeleton.

NMQR 1655, 3209, 3688, 3925, 3926, 3929, 3933: Skulls (Uppermost slopes of Loskop and Spitskop).

 

CLUVER, 1971

Locality: South Africa.

Horizon: Katberg Formation, Beaufort Group.

Biostratigraphy: Lystrosaurus Assemblage zone (Lystrosaurus-Thrinaxodon zone, KEYSER, 1979, SACS, 1980, Procolophon zone, Lystrosaurus zone, WATSON, 1914, BROOM, 1906).

Age: Early Scythian Epoch, Early Triassic.

Material:

SAM 171, K1392, 3455: Skulls.

JASINOSKI, RAYFIELD & CHINSAMY, 2010

SAM-PK-K 4800: Skull.

TTHACKERAY, DURAND, & MEYER, 1998

TM 27, 30, 38, 113, 117, 139, 165, 215, 1004, 3580, 4049, 4387, 4429, ghg 186, 240, 244, 255, 283, 355, 386, 387, 402, 404, 425, wb 503: Skulls.

 

= Dicynodon latirostris OWEN, 1860
Etymology: Laitn, lati, "broad, wide" and Latin, rostris, "nose, snout."
= Ptychognathus latirostris (OWEN, 1860) OWEN, 1876
= Lystrosaurus latirostris (OWEN, 1860)
= Ptychosiagum latirostris (OWEN, 1860) LYDEKKER in NICHOLSON & LYDEKKER, 1889

Holotype: BMNH 36222

Locality: Rhenosterberg branch of the Sneewberg range, Graaff Reinet District, Eastern Cape Province, South Africa.

Horizon: Katberg Formation, Beaufort Group.

Biostratigraphy: Lystrosaurus Assemblage zone (Lystrosaurus-Thrinaxodon zone, KEYSER, 1979, SACS, 1980, Procolophon zone, Lystrosaurus zone, WATSON, 1914, BROOM, 1906).

Age: Early Scythian Epoch, Early Triassic.

Material: Fragmentary skull.

Referred material:

BROOM 1915

Locality: Smithfield, Free (Orange Free) State, South Africa.

Horizon: Katberg Formation, Beaufort Group.

Biostratigraphy: Lystrosaurus Assemblage zone (Lystrosaurus-Thrinaxodon zone, KEYSER, 1979, SACS, 1980, Procolophon zone, Lystrosaurus zone, WATSON, 1914, BROOM, 1906).

Age: Early Scythian Epoch, Early Triassic.

Material:

AMNH 5600: Skull.

 

Locality: Harrismith, Free (Orange Free) State, South Africa.

Horizon: Katberg Formation, Beaufort Group.

Biostratigraphy: Lystrosaurus Assemblage zone (Lystrosaurus-Thrinaxodon zone, KEYSER, 1979, SACS, 1980, Procolophon zone, Lystrosaurus zone, WATSON, 1914, BROOM, 1906).

Age: Early Scythian Epoch, Early Triassic.

Material:

AMNH 5629: Skull of a juvenile.

 

CLUVER, 1971

Locality: South Africa.

Horizon: Katberg Formation, Beaufort Group.

Biostratigraphy: Lystrosaurus Assemblage zone (Lystrosaurus-Thrinaxodon zone, KEYSER, 1979, SACS, 1980, Procolophon zone, Lystrosaurus zone, WATSON, 1914, BROOM, 1906).

Age: Early Scythian Epoch, Early Triassic.

Material:

Nat. Mus. No. C150: Skull, mandible and postcranial scraps.

Nat. Mus. No. C403: Skull and mandible.

Nat. Mus. No. C391, Nat. Mus. No. C171: Skulls.

SAM K 1284: Fragmentary skull and mandible.

 

van HOEPEN, 1913

Locality:

Horizon:

Biostratigraphy:

Age:

Material:

112, 21, 4000-: Skulls.

 

LYDEKKER, 1890

Locality: Styl-Krantz, Graaff Reinet District, Eastern Cape Province, South Africa.

Horizon: Katberg Formation, Beaufort Group.

Biostratigraphy: Lystrosaurus Assemblage zone (Lystrosaurus-Thrinaxodon zone, KEYSER, 1979, SACS, 1980, Procolophon zone, Lystrosaurus zone, WATSON, 1914, BROOM, 1906).

Age: Early Scythian Epoch, Early Triassic.

Material:

BMNH 47064: Fragmentary skull.

 

 

Locality: Rhenosterberg, Graaff Reinet District, Eastern Cape Province, South Africa.

Horizon: Katberg Formation, Beaufort Group.

Biostratigraphy: Lystrosaurus Assemblage zone (Lystrosaurus-Thrinaxodon zone, KEYSER, 1979, SACS, 1980, Procolophon zone, Lystrosaurus zone, WATSON, 1914, BROOM, 1906).

Age: Early Scythian Epoch, Early Triassic.

Material:

BMNH 47342: Fragmentary skull.

 

= Dicynodon alfredi OWEN, 1861
Etymology:
= Ptychognathus alfredi (OWEN, 1861) OWEN, 1876
= Lystrosaurus alfredi (OWEN, 1861)

Holotype: BMNH 47342

Locality: Rhenostereberg, Graaff Reinet District, Eastern Cape Province, South Africa.

Horizon: Katberg Formation, Beaufort Group.

Biostratigraphy: Lystrosaurus Assemblage zone (Lystrosaurus-Thrinaxodon zone, KEYSER, 1979, SACS, 1980, Procolophon zone, Lystrosaurus zone, WATSON, 1914, BROOM, 1906).

Age: Early Scythian Epoch, Early Triassic.

Material: Fragmentary skull.

 

= Dicynodon depressus OWEN, 1876
Etymology:
= Ptychognathus depressus (OWEN, 1876) OWEN, 1876
= Lystrosaurus depressus (OWEN, 1876)

Holotype: BMNH 47064

Locality: Styl-Krantz, Graaff Reinet District, Eastern Cape Province, South Africa.

Horizon: Katberg Formation, Beaufort Group.

Biostratigraphy: Lystrosaurus Assemblage zone (Lystrosaurus-Thrinaxodon zone, KEYSER, 1979, SACS, 1980, Procolophon zone, Lystrosaurus zone, WATSON, 1914, BROOM, 1906).

Age: Early Scythian Epoch, Early Triassic.

Material: Flattened skull.

 

= Lystrosaurus wagneri van HOEPEN, 1916
Etymology:

Holotype: Transvall Museum 28

Locality: Harrismith, Free (Orange Free) State, South Africa.

Horizon: Katberg Formation, Beaufort Group.

Biostratigraphy: Lystrosaurus Assemblage zone (Lystrosaurus-Thrinaxodon zone, KEYSER, 1979, SACS, 1980, Procolophon zone, Lystrosaurus zone, WATSON, 1914, BROOM, 1906).

Age: Early Scythian Epoch, Early Triassic.

Material: Skull.

 

= Lystrosaurus primitivus TOERIEN, 1954
Etymology: Latin, primitivus, "primitive."

Holotype: BP 330

Locality: Farm Ripplemead, New Bethesda District, north of Graaff Reinet District, Eastern Cap Province, South Africa.

Horizon: Katberg Formation, Beaufort Group.

Biostratigraphy: Lystrosaurus Assemblage zone (Lystrosaurus-Thrinaxodon zone, KEYSER, 1979, SACS, 1980, Procolophon zone, Lystrosaurus zone, WATSON, 1914, BROOM, 1906).

Age: Early Scythian Epoch, Early Triassic.

Material: Skull.

Lystrosaurus primitivus modified from TOERIEN, 1954.

_____________________________________________________________________________________

Species: murrayi (HUXLEY, 1859) COPE, 1870
Etymology:
= Dicynodon murrayi HUXLEY, 1859
= Ptychosiagum murrayi (HUXLEY, 1859) LYDEKKER in NICHOLSON & LYDEKKER, 1889

Holotype: BMNH R1291

Locality: Middle Beaufort Beds, Colesburg District, Cape Province, South Africa.

Horizon: Katberg Formation, Beaufort Group.

Biostratigraphy: Lystrosaurus Assemblage zone (Lystrosaurus-Thrinaxodon zone, KEYSER, 1979, SACS, 1980, Procolophon zone, Lystrosaurus zone, WATSON, 1914, BROOM, 1906).

Age: Early Scythian Epoch, Early Triassic.

Material: Imperfect skull.

Modified from Colbert, 1974.

Modified from Colbert, 1974.

Referred material:

BMNH R1292, R1294: Fragmentary skulls.

RAY, CHINSAMY & BANDYOPADHYAY, 2005

SAM-PK-11184: Humerus, radius, ulna, femur and rib.

JASINOSKI, RAYFIELD & CHINSAMY, 2010

SAM-PK-K 1342: Fragmentary skull.

TTHACKERAY, DURAND, & MEYER, 1998

TM 15, 17, 18, 20, 21, 67, 69, 70, 71, 112, 154, 213, 288.1, 3534, 4051, 4058, 4113, O 12: Skulls.

 

BOTHA-BRINK, HUTTENLOCKER, & MODESTO, 2014

Locality: Nooitgedacht 68, Bethulie District, Eastern Cape Province, South Africa.

Horizon: Palingkloof Member, Balfour Formation.

Biostratigraphy: Lystrosaurus Assemblage Zone.

Age: Early Scythian Epoch, Early Triassic.

Material:

BP/1/3974, 3975, 3977, 3978, 3979, 4030, 4039, 4040: Skulls and skeletons.

NMQR 3212, 3649, 3932, 3927: Skulls and associated postcrania (Uppermost slopes of Koskop and Spitskop).

CAMP, 1956

Locality: 2 1/2 miles South East of Thaba n'Chu, Free (Orange Free) State, South Africa.

Horizon: Katberg Formation, Beaufort Group.

Biostratigraphy: Lystrosaurus Assemblage zone (Lystrosaurus-Thrinaxodon zone, KEYSER, 1979, SACS, 1980, Procolophon zone, Lystrosaurus zone, WATSON, 1914, BROOM, 1906).

Age: Early Scythian Epoch, Early Triassic.

Material:

UCMP 42007: Skull.

 

BRINK, 1982

Locality: Valkfontein, Middelburg, Albany District, Eastern Cape Province, South Africa.

Horizon: Katberg Formation, Beaufort Group.

Biostratigraphy: Lystrosaurus Assemblage zone (Lystrosaurus-Thrinaxodon zone, KEYSER, 1979, SACS, 1980, Procolophon zone, Lystrosaurus zone, WATSON, 1914, BROOM, 1906).

Age: Early Scythian Epoch, Early Triassic.

Material:

BP M112: Skull and lower jaw.

 

 

Locality: Elim, Burgersdorp, Albany District, Eastern Cape Province, South Africa..

Horizon: Katberg Formation, Beaufort Group.

Biostratigraphy: Lystrosaurus Assemblage zone (Lystrosaurus-Thrinaxodon zone, KEYSER, 1979, SACS, 1980, Procolophon zone, Lystrosaurus zone, WATSON, 1914, BROOM, 1906).

Age: Early Scythian Epoch, Early Triassic.

Material:

BP M116: Skull and lower jaw.

 

BROOM, 1915

Locality: Unknown, South Africa.

Horizon: Katberg Formation, Beaufort Group.

Biostratigraphy: Lystrosaurus Assemblage zone (Lystrosaurus-Thrinaxodon zone, KEYSER, 1979, SACS, 1980, Procolophon zone, Lystrosaurus zone, WATSON, 1914, BROOM, 1906).

Age: Early Scythian Epoch, Early Triassic.

Material:

AMNH 5629: Snout.

 

CLUVER, 1971

Locality: South Africa.

Horizon: Katberg Formation, Beaufort Group.

Biostratigraphy: Lystrosaurus Assemblage zone (Lystrosaurus-Thrinaxodon zone, KEYSER, 1979, SACS, 1980, Procolophon zone, Lystrosaurus zone, WATSON, 1914, BROOM, 1906).

Age: Early Scythian Epoch, Early Triassic.

Material:

Nat. Mus. No. C 282, Nat. Mus. No. C211, Nat. Mus. No. C228: Skulls and mandibles.

SAM-PK-K1495: Complete skeleton.

Nat. Mus. No. C6457: Skull, mandible and cervical vertebrae.

SAM 1336: Fragmentary skull and mandible.

 

COLBERT, 1974

Locality: Northwest slope of Coalsack Bluff, Transantarctic Mountains, East Antarctica, Antarctica.

Horizon: Fremouv Formation.

Biostratigraphy:

Age: Anisian Stage, Lower Mid Triassic Epoch, Early Middle Triassic.

Material:

AMNH 9302: Right maxilla with tooth.

AMNH 9308: Left maxilla, lacking tooth.

AMNH 9302, modified from Colbert, 1974.

KITCHING

Locality: Nooitgedacht, Bethulie, Free (Orange Free) State, South Africa.

Horizon: Katberg Formation, Beaufort Group.

Biostratigraphy: Lystrosaurus Assemblage zone (Lystrosaurus-Thrinaxodon zone, KEYSER, 1979, SACS, 1980, Procolophon zone, Lystrosaurus zone, WATSON, 1914, BROOM, 1906).

Age: Early Scythian Epoch, Early Triassic.

Material:

BP 409, 411: Skulls, lower jaws and fragmentary skeletons.

BP 410: Skull lower jaw and almost complete skeleton.

BP 413: Skull and dentary.

 

 

Locality: Orange, Bethulie, Free (Orange Free) State, South Africa.

Horizon: Katberg Formation, Beaufort Group.

Biostratigraphy: Lystrosaurus Assemblage zone (Lystrosaurus-Thrinaxodon zone, KEYSER, 1979, SACS, 1980, Procolophon zone, Lystrosaurus zone, WATSON, 1914, BROOM, 1906).

Age: Early Scythian Epoch, Early Triassic.

Material:

BP 407: Skull and lower jaw of a juvenile.

 

Locality: Old Brickfield Donga, Harrismith, Free (Orange Free) State, South Africa.

Horizon: Katberg Formation, Beaufort Group.

Biostratigraphy: Lystrosaurus Assemblage zone (Lystrosaurus-Thrinaxodon zone, KEYSER, 1979, SACS, 1980, Procolophon zone, Lystrosaurus zone, WATSON, 1914, BROOM, 1906).

Age: Early Scythian Epoch, Early Triassic.

Material:

BP 408: Skull, lower jaw, cervical vertebrae and shoulder girdle of a juvenile.

 

 

Locality: Donovan, Bethulie, Free (Orange Free) State, South Africa.

Horizon: Katberg Formation, Beaufort Group.

Biostratigraphy: Lystrosaurus Assemblage zone (Lystrosaurus-Thrinaxodon zone, KEYSER, 1979, SACS, 1980, Procolophon zone, Lystrosaurus zone, WATSON, 1914, BROOM, 1906).

Age: Early Scythian Epoch, Early Triassic.

Material:

BP 412: Skull and lower jaw of a juvenile.

 

Locality: Edenvale, Naauwpoort, South Africa.

Horizon: Katberg Formation, Beaufort Group.

Biostratigraphy: Lystrosaurus Assemblage zone (Lystrosaurus-Thrinaxodon zone, KEYSER, 1979, SACS, 1980, Procolophon zone, Lystrosaurus zone, WATSON, 1914, BROOM, 1906).

Age: Early Scythian Epoch, Early Triassic.

Material:

BP 106: Skull and lower jaw.

BRINK, 1982

BP M111: Skull and lower jaw.

BP M117: Skull and lower jaw.

 

KITCHING, 1978

Locality: Harrismith, Free (Orange Free) State, South Africa.

Horizon: Katberg Formation, Beaufort Group.

Biostratigraphy: Lystrosaurus Assemblage zone (Lystrosaurus-Thrinaxodon zone, KEYSER, 1979, SACS, 1980, Procolophon zone, Lystrosaurus zone, WATSON, 1914, BROOM, 1906).

Age: Early Scythian Epoch, Early Triassic.

Material:

BP M134: Fragmentary skeleton mixed with 7 Lydekkerinids remains.

RAY, 2005

SAM-PK-3596: Skull and lower jaw.

RAY, CHINSAMY & BANDYOPADHYAY, 2005

SAM-PK-K1415: Dorsal rib fragments.

LYDEKKER, 1890

Locality: Rhenosterberg branch of the Sneewberg range, Graaff Reinet district, Eastern Cape Province, South Africa.

Horizon: Katberg Formation, Beaufort Group.

Biostratigraphy: Lystrosaurus Assemblage zone (Lystrosaurus-Thrinaxodon zone, KEYSER, 1979, SACS, 1980, Procolophon zone, Lystrosaurus zone, WATSON, 1914, BROOM, 1906).

Age: Early Scythian Epoch, Early Triassic.

Material:

BMNH 36224, 36253, 36223: Fragmentary skulls.

 

RAY, 2005

Locality: Free (Orange Free) State, South Africa.

Horizon: Katberg Formation, Beaufort Group.

Biostratigraphy: Lystrosaurus Assemblage zone (Lystrosaurus-Thrinaxodon zone, KEYSER, 1979, SACS, 1980, Procolophon zone, Lystrosaurus zone, WATSON, 1914, BROOM, 1906).

Age: Early Scythian Epoch, Early Triassic.

Material:

SAM-PK-3531: Skull, lower jaw and postcranials.

STATE (OFS)

SAM-PK-3593: Skull, and partial lower jaw.

SAM-PK-K1396: Skull and lower jaw.

SAM-PK-1397: Skull, lower jaw and postcranials.

SAM-PK-3594: Partial skull, and lower jaw.

SAM-PK-K1398: Skull.

 

Locality: Bethuile, Free (Orange Free) State, South Africa.

Horizon: Katberg Formation, Beaufort Group.

Biostratigraphy: Lystrosaurus Assemblage zone (Lystrosaurus-Thrinaxodon zone, KEYSER, 1979, SACS, 1980, Procolophon zone, Lystrosaurus zone, WATSON, 1914, BROOM, 1906).

Age: Early Scythian Epoch, Early Triassic.

Material:

SAM-PK-K8013: Skull, lower jaw and partial skeleton.

SAM-PK-K8038: Complete skeleton.

RAY, CHINSAMY & BANDYOPADHYAY, 2005

SAM-PK-K8012: Scapula, vertebra and rib.

 

Locality: Middleburg District, Eastern Cape Province, South Africa.

Horizon: Katberg Formation, Beaufort Group.

Biostratigraphy: Lystrosaurus Assemblage zone (Lystrosaurus-Thrinaxodon zone, KEYSER, 1979, SACS, 1980, Procolophon zone, Lystrosaurus zone, WATSON, 1914, BROOM, 1906).

Age: Early Scythian Epoch, Early Triassic.

Material:

SAM-PK-K7: SKull, lower jaw and postcranials.

RAY, CHINSAMY & BANDYOPADHYAY, 2005

SAM-PK-K8: Left humerus and rib fragments.

 

 

Locality: Cape Province, South Africa.

Horizon: Katberg Formation, Beaufort Group.

Biostratigraphy: Lystrosaurus Assemblage zone (Lystrosaurus-Thrinaxodon zone, KEYSER, 1979, SACS, 1980, Procolophon zone, Lystrosaurus zone, WATSON, 1914, BROOM, 1906).

Age: Early Scythian Epoch, Early Triassic.

Material:

SAM-PK-K21: Skull, and lower jaw.

SAM-PK-K20: Partial skull.

 

Locality: KwaZulu Natal Province, South Africa.

Horizon: Katberg Formation, Beaufort Group.

Biostratigraphy: Lystrosaurus Assemblage zone (Lystrosaurus-Thrinaxodon zone, KEYSER, 1979, SACS, 1980, Procolophon zone, Lystrosaurus zone, WATSON, 1914, BROOM, 1906).

Age: Early Scythian Epoch, Early Triassic.

Material:

SAM-PK-K8311: Skull and lower jaw.

 

Locality: Unknown, South Africa.

Horizon: Katberg Formation, Beaufort Group.

Biostratigraphy: Lystrosaurus Assemblage zone (Lystrosaurus-Thrinaxodon zone, KEYSER, 1979, SACS, 1980, Procolophon zone, Lystrosaurus zone, WATSON, 1914, BROOM, 1906).

Age: Early Scythian Epoch, Early Triassic.

Material:

SAM-PK-1165: Skull and postcranial.

SAM-PK-1268: Skull, lower jaw and postcranials.

Nat. Mus. C. 282, SAM-PK-NII: Skulls.

SAM-PK-K8524: Skull, and partial postcranials.

 

 

RAY, CHINSAMY & BANDYOPADHYAY, 2005

Locality: Meadows, South Africa.

Horizon: Katberg Formation, Beaufort Group.

Biostratigraphy: Lystrosaurus Assemblage zone.

Age: Early Scythian Epoch, Early Triassic.

Material:

SAM-PK-8991a: Humerus, radius, ulna, scapula and tibia.

SAM-PK-8991bD: Humerus and femur.

SAM-PK-8796a: Scapula, femur, tibia, and phalanx.

SAM-PK-8796b: Femur.

 

Locality: Banspatail Locality, Damodar Basin, West Bengal State, India.

Horizon: Panchet Formation.

Biostratigraphy:

Age: Early Triassic.

Material:

ISIR758, 756, 765, 757, 761, 764: Humeri.

ISIR 762, 763, 751, 750: Humeri.

ISIR 759: Tibia.

ISIR 760: Fibula.

ISIR 754: Clavicle.

ISIR 752: Dorsal rib.

 

TRIPATHI & SATSANGI, 1963

Locality: Talkunri, Raniganji Coalfield, West Bengal State, India.

Horizon: Panchet beds.

Biostratigraphy:

Age: Early Triassic.

Material:

GSI Type 17851,17856, 17846, 17853, 17859: Skulls and lower jaws.

GSI Type 17854, 17847, 17864: Partial skulls and lower jaw.

 

Locality: Batiata, Raniganji Coalfield, West Bengal State, India.

Horizon: Panchet beds.

Biostratigraphy:

Age: Early Triassic.

Material:

GSI Type 17852, 17858: Skulls and lower jaw.

GSI 17844: Skull, lower jaw and a few postcranials.

 

Locality: Iswarda, Raniganji Coalfield, West Bengal State, India.

Horizon: Panchet beds.

Biostratigraphy:

Age: Early Triassic.

Material:

GSI Type 17855, 17857: Partial skulls and lower jaw.

 

Locality: Kukrakhuri, Raniganji Coalfield, West Bengal State, India.

Horizon: Panchet beds.

Biostratigraphy:

Age: Early Triassic.

Material:

GSI Type 17848: Partial skull and lower jaw.

 

Locality: Banspetall, Raniganji Coalfield, West Bengal State, India.

Horizon: Panchet beds.

Biostratigraphy:

Age: Early Triassic.

Material:

GSI Type 17849: Skull and lower jaw.

RAY, 2005

ISIR 766: Nearly complete skull and lower jaw.

ISIR 767: Partial skull and lower jaw.

ISIR 768: Occiput, and partila lower jaw.

 

Locality: Unknown, Raniganji Coalfield, West Bengal State, India.

Horizon: Panchet beds.

Biostratigraphy:

Age: Early Triassic.

Material:

GSI Type 17843, 17863: Skulls.

GSI Type 17860: Skull and lower jaw.

 

= Lystrosaurus platyceps TRIPATHI & SATSANGI, 1963

Locality: Talkunri, Raniganji Coalfield, West Bengal State, India.

Horizon: Panchet beds.

Biostratigraphy:

Age: Early Triassic.

Material:

GSI Type 17865: Lower jaw and a few postcranial elements.

Modified from Colbert, 1974.

= Dicynodon orientalis HUXLEY, 1865
Etymology: Latin, orientalis, "eastern, from the east."
= Ptychognathus orientalis (HUXLEY, 1865) OWEN, 1876
= Ptychosiagum orientalis (HUXLEY, 1865) LYDEKKER in NICHOLSON & LYDEKKER, 1889
= Lystrosaurus orientalis (HUXLEY, 1865) DAS GUPTA, 1922

Holotype: BMNH R1026

Locality: Near Raniganji Coalfield, West Bengal State, India.

Horizon: Panchet beds.

Biostratigraphy:

Age: Early Triassic.

Material: Left occiputal region.

Referred material:

BMNH R1026a, R1026b: Fragmentary scapula.

BMNH R1026c: Right coracoid.

BMNH R1026d: Right humerus.

BMNH R1026e: 3 fragmentary dorsal vertebrae.

BMNH R1026f: Anterior portion of the centrum and right rib of a sacral vertebra.

Fragmentary pelvis.

 

TRIPATHI & PURI, 1962

Locality: Locality K47/579, 3 furlongs to the W.N.W. of Deoli, 23°39’, 86°53’, in the Damodar river, Raniganji Coalfield, West Bengal State, India.

Horizon: Panchet beds.

Biostratigraphy:

Age: Early Triassic.

Material:

GSI 17815: Left maxilla fragment with tusk.

GSI 17817: Premaxilla.

GSI 17840: Right humeri.

GSI 17821: Left ulna.

GSI 17822: Right ulna.

GSI 17842: Right ulna of a juvenile.

GSI 17825: Left femur.

GSI 17826: Fragmentary left femur.

GSI 17829: Fragmentary left fibula.

GSI 17830: Fragmentary right fibula.

GSI 17833: Left scapula.

GSI 17836: Fragmentary left ilium.

 

Locality: Locality K47/578, ½ miles to the south of Dhenua, 23°38’00”, 86°58’00”, in the Damodar river, Raniganji Coalfield, West Bengal State, India.

Horizon: Panchet beds.

Biostratigraphy:

Age: Early Triassic.

Material:

GSI 17816: Right maxilla with tusk.

GSI 17820: Right humerus.

 

Locality: Locality K47/586, northern side of the railway line, 5 furlongs N.W. of Dihika, Raniganji Coalfield, West Bengal State, India.

Horizon: Panchet beds, Early Triassic.

Material:

GSI 17827, 17828: Fragmentary tibiae.

 

Locality: Locality K47/580, left bank of the Damodar river, ½ mile to the W.N.W. of Dihika, 23°38’30”, 86°54’30”, Raniganji Coalfield, West Bengal State, India.

Horizon: Panchet beds.

Biostratigraphy:

Age: Early Triassic.

Material:

GSI 17818: Left humerus.

GSI 17831: Fragmentary fibula.

GSI 17837: Right ischium.

 

Locality: Locality K47/588, near the right bank of the Damodar river, 1 mile south east of Bagulia, 23°33’30”, 86°53’30”, Raniganji Coalfield, West Bengal State, India.

Horizon: Panchet beds.

Biostratigraphy:

Age: Early Triassic.

Material:

GSI 17834, A, B, C, (K47/581, K47/582, K47/584): Fragmentary scapula.

GSI 17835: Fragmentary coracoid.

 

 

Locality: Locality K47/589, 1 ¼ miles south east of Bagulia, Raniganji Coalfield, West Bengal State, India.

Horizon: Panchet beds.

Biostratigraphy:

Age: Early Triassic.

Material:

GSI 17819: Left humerus.

 

Locality: Locality K47/591, right bank of the Khanchi Jhor, ½ mile east of Kesharkundi, Raniganji Coalfield, West Bengal State, India.

Horizon: Panchet beds.

Biostratigraphy:

Age: Early Triassic.

Material:

GSI 17822: Right radius.

 

 

Locality: Locality K47/592, right bank of the Khanchi Jhor, 3 ½ furlongs east of Rajpura, 23°37’, 86°56’, Raniganji Coalfield, West Bengal State, India.

Horizon: Panchet beds.

Biostratigraphy:

Age: Early Triassic.

Material:

GSI 17832: Left scapula.

 

= Lystrosaurus frontosus COPE, 1870
Etymology:
= Ptychognathus frontosus (COPE, 1870)

Holotype: YPM 2225

Locality: 200 miles from Cape of Good Hope, Orange River, South Africa.

Horizon: Katberg Formation, Beaufort Group.

Biostratigraphy: Lystrosaurus Assemblage zone (Lystrosaurus-Thrinaxodon zone, KEYSER, 1979, SACS, 1980, Procolophon zone, Lystrosaurus zone, WATSON, 1914, BROOM, 1906).

Age: Early Scythian Epoch, Early Triassic.

Material: Skull and lower jaw.

 

= Ptychognathus verticalis OWEN, 1876
Etymology:
= Lystrosaurus verticalis (OWEN, 1860)

Holotype: BMNH 36224

Locality: Rhenosterberg, Graaff Reinet District, Eastern Cape Province, South Africa.

Horizon: Katberg Formation, Beaufort Group.

Biostratigraphy: Lystrosaurus Assemblage zone (Lystrosaurus-Thrinaxodon zone, KEYSER, 1979, SACS, 1980, Procolophon zone, Lystrosaurus zone, WATSON, 1914, BROOM, 1906).

Age: Early Scythian Epoch, Early Triassic.

Material: Imperfect skull.

 

= Ptychognathus boopis OWEN, 1876
Etymology:
= Lystrosaurus boopis (OWEN, 1876)

Holotype: BMNH 36253

Locality: Rhenosterberg, Graaff Reinet District, Eastern Cape Province, South Africa.

Horizon: Katberg Formation, Beaufort Group.

Biostratigraphy: Lystrosaurus Assemblage zone (Lystrosaurus-Thrinaxodon zone, KEYSER, 1979, SACS, 1980, Procolophon zone, Lystrosaurus zone, WATSON, 1914, BROOM, 1906).

Age: Early Scythian Epoch, Early Triassic.

Material: Imperfect skull.

 

= Dicynodon copei SEELEY, 1889
Etymology: In honor of E. D. Cope.

Holotype: BMNH 47074

Locality: Cape Colony, South Africa.

Horizon:

Biostratigraphy:

Age: Early Triassic.

Material: Fragmentary skull.

Referred material:

BMNH 47073: Fragmentary skull.

 

= Dicynodon strigops BROOM, 1913
Etymology:
= Prolystrosaurus strigops (BROOM, 1913)

Holotype: AMNH 5581

Locality: Harrismith, Free (Orange Free) State, South Africa.

Horizon:

Biostratigraphy: Dicynodon Assemblage zone (formerly Dicynodon lacerticeps-Whaitsia zone of KEYSER, 1979, SACS, 1980, Daptocephalus zone of KITCHING, 1870, 1977, Upper Cistecephalus Zone of WATSON, 1914).

Age: Tatarian Age, Lopingian Subepoch, Zechstein Epoch, Late Permian.

Material: Fragmentary skull and fragmentary skeletons of 3 individuals.

 

= Lystrosaurus jeppei van HOEPEN, 1916
Etymology:

Holotype: Transvaal Museum 15

Locality: Harrismith, Free (Orange Free) State, South Africa.

Horizon: Katberg Formation, Beaufort Group.

Biostratigraphy: Lystrosaurus Assemblage zone (Lystrosaurus-Thrinaxodon zone, KEYSER, 1979, SACS, 1980, Procolophon zone, Lystrosaurus zone, WATSON, 1914, BROOM, 1906).

Age: Early Scythian Epoch, Early Triassic.

Material: Skull.

 

= Prolystrosaurus natalensis HAUGHTON, 1917
Etymology: In reference to Natal Province, South Africa.
= Lystrosaurus natalensis (HAUGHTON, 1917)

Holotype: SAM 3715

Locality: Loskop, KwaZulu Natal Province, South Africa.

Horizon: Katberg Formation, Beaufort Group.

Biostratigraphy: Lystrosaurus Assemblage zone (Lystrosaurus-Thrinaxodon zone, KEYSER, 1979, SACS, 1980, Procolophon zone, Lystrosaurus zone, WATSON, 1914, BROOM, 1906).

Age: Early Scythian Epoch, Early Triassic.

Material: Skull and lower jaw.

Referred material:

SAM 331g: Skull, lower jaw, and nearly complete skeleton.

 

JANENSCH, 1952

Locality: Bergville, KwaZulu Natal Province, South Africa.

Horizon: Katberg Formation, Beaufort Group.

Biostratigraphy: Lystrosaurus Assemblage zone (Lystrosaurus-Thrinaxodon zone, KEYSER, 1979, SACS, 1980, Procolophon zone, Lystrosaurus zone, WATSON, 1914, BROOM, 1906).

Age: Early Scythian Epoch, Early Triassic.

Material:

Number: Not given: Fragmentary lower jaw.

 

= Lystrosaurus rubidgei BROOM, 1940
Etymology: In honor of Dr. S. H. Rubidge.

Holotype: RC 46

Locality: Bethesda Road, Graaff Reinet District, Eastern Cape Province, South Africa.

Horizon: Katberg Formation, Beaufort Group.

Biostratigraphy: Lystrosaurus Assemblage zone (Lystrosaurus-Thrinaxodon zone, KEYSER, 1979, SACS, 1980, Procolophon zone, Lystrosaurus zone, WATSON, 1914, BROOM, 1906).

Age: Early Scythian Epoch, Early Triassic.

Material: Anterior 2/3rds of skull.

 

= Lystrosaurus bothai BROOM, 1949
Etymology:

Holotype: RC?

Locality: Rygerspoort, Cradock District, Eastern Cape Province, South Africa.

Horizon: Katberg Formation, Beaufort Group.

Biostratigraphy: Lystrosaurus Assemblage zone (Lystrosaurus-Thrinaxodon zone, KEYSER, 1979, SACS, 1980, Procolophon zone, Lystrosaurus zone, WATSON, 1914, BROOM, 1906).

Age: Early Scythian Epoch, Early Triassic.

Material: Skull.

 

= Lystrosaurus rajurkari TRIPATHI & SATSANGI, 1963
Etymology:

Holotype: GSI Type Number 17867

Locality: Talkuri, about 3 miles south east of Asansol, West Bengal State, India.

Horizon: Panchet Formation.

Biostratigraphy:

Age: Early Triassic.

Material: Fragmentary skull.

Referred material:

DAS GUPTA, 1922/COTTER, 1917

Locality: West Bengal State, India.

Horizon: Panchet beds.

Biostratigraphy:

Age: Early Triassic.

Material:

Number: Not given: Fragmentary remains.

= Lystrosaurus mccaigi TRIPATHI & SATASANGI, 1963

Locality: Iswarda, near Anasol, Raniganj Coalfield, West Bengal State, India.

Horizon: Upper Panchet Series.

Biostratigraphy:

Age: Early Triassic.

Material:

GSI 17866: Fragmentary skull.

 

_____________________________________________________________________________________

Species: curvatus (OWEN, 1876)
Etymology:
= Dicynodon curvatus OWEN, 1876

Holotype: BMNH R3792

Locality: Middle Beaufort Beds, Elandsberg, Cradock District, Eastern Cape Province, South Africa.

Horizon: Katberg Formation, Beaufort Group.

Biostratigraphy: Lystrosaurus Assemblage zone.

Age: Early Scythian Epoch, Early Triassic.

Material: Skull.

Modified from Colbert, 1974.

Referred material:

BOTHA-BRINK, 2017

Locality: 24m from the base of the hill Maanhaar on the farm Barendskraal, Eastern Cape Province, South Africa.

Horizon: Palingkloof Member, Balfour Formation.

Biostratigraphy: Lystrosaurus Assemblage zone.

Age: Early Scythian Epoch, Early Triassic.

Material:

NMQR 4001: Skull and fragmentary skeleton.

Note: Found in a burrow cast.

 

BOTHA-BRINK, HUTTENLOCKER, & MODESTO, 2014

Locality: Nooitgedacht 68, Bethulie District,  Eastern Cape Province, South Africa.

Horizon: Balfour Formation.

Biostratigraphy: Dicynodon Assemblage Zone.

Age: Tatarian Age, Lopingian Subepoch, Zechstein Epoch, Late Permian.

Material:

BP/1/3976: Skull.

 

CLUVER, 1971

Locality: South Africa.

Horizon: Katberg Formation, Beaufort Group.

Biostratigraphy: Lystrosaurus Assemblage zone.

Age: Early Scythian Epoch, Early Triassic.

Material:

Nat. Mus. No. C299: Skull.

SAM K1284: Skull and mandible with braincase.

SAM K90: Fragmentary skull and mandible.

SURKOV & BENTON, 2004

CAMZM T788: Fragmentary skull.

 

COLBERT, 1974

Locality: Shackleton-McGregor Glacier, Halfmoon Bluff, Seintinel Hill, Queen Maud Mountains, East Antarctica, Antarctica.

Horizon: Fremouv Formation.

Biostratigraphy:

Age: Anisian Stage, Lower Mid Triassic Epoch, Early Middle Triassic.

Material:

AMNH 9515: Left side of skull, with a sagittal section exposed in the rock.

 

COSGRIFF & HAMMER, 1981

Locality: Cumulus Hills, Shackleton Glacier, Queen Maud Mountains, East Antarctica, Antarctica.

Horizon: Fremouv Formation.

Biostratigraphy:

Age: Anisian Stage, Lower Mid Triassic Epoch, Early Middle Triassic.

Material:

WSU 0977: Fragmentary skull.

 

LYDEKKER, 1890

Locality: Cradock, northwest of Fort Beaufort, Eastern Cape Province, South Africa.

Horizon:

Biostratigraphy:

Age: Early Triassic.

Material:

BMNH R1655: Fragmentary skull.

 

RAY, 2005

Locality: Middleburg District, Eastern Cape Province, South Africa.

Horizon: Katberg Formation, Beaufort Group.

Biostratigraphy: Lystrosaurus Assemblage zone (Lystrosaurus-Thrinaxodon zone, KEYSER, 1979, SACS, 1980, Procolophon zone, Lystrosaurus zone, WATSON, 1914, BROOM, 1906).

Age: Early Scythian Epoch, Early Triassic.

Material:

SAM-PK-K1381: Skull and postcranials.

SAM-PK-K3, K1362: Skulls and lower jaws.

SAM-PK-K27: Skull, lower jaw and postcranials.

 

Locality: Bethulie, Free (Orange Free) State, South Africa.

Horizon: Katberg Formation, Beaufort Group.

Biostratigraphy: Lystrosaurus Assemblage zone (Lystrosaurus-Thrinaxodon zone, KEYSER, 1979, SACS, 1980, Procolophon zone, Lystrosaurus zone, WATSON, 1914, BROOM, 1906).

Age: Early Scythian Epoch, Early Triassic.

Material:

SAM-PK-K8014: Skull and lower jaw.

 

= Lystrosaurus wagneri van HOEPEN, 1916
Etymology:

Holotype: Transvaal Museum 18

Locality: Harrismith, Free (Orange Free) State, South Africa.

Horizon: Katberg Formation, Beaufort Group.

Biostratigraphy: Lystrosaurus Assemblage zone.

Age: Early Scythian Epoch, Early Triassic.

Material: Skull.

 

= Lystrosaurus jorisseni van HOEPEN, 1916
Etymology:

Holotype: Transvaal Museum 17

Locality: Harrismith, Free (Orange Free) State, South Africa.

Horizon: Katberg Formation, Beaufort Group.

Biostratigraphy: Lystrosaurus Assemblage zone.

Age: Early Scythian Epoch, Early Triassic.

Material: Compressed skull.

 

= Lystrosaurus theileri van HOEPEN, 1916
Etymology:

Holotype: Transval Museum 16

Locality: Harrismith, Free (Orange Free) State, South Africa.

Horizon: Katberg Formation, Beaufort Group.

Biostratigraphy: Lystrosaurus Assemblage zone.

Age: Early Scythian Epoch, Early Triassic.

Material: Damaged skull.

 

= Lystrosaurus platyceps (SEELEY, 1898)
Etymology: Greek, platy, "broad, flat, wide" and Latin, -ceps, "head."
= Mochlorhinus platyceps SEELEY, 1898

Holotype: Albany Museum AMG 454.

Locality: Bethulie, Free (Orange Free) State, South Africa.

Horizon: Katberg Formation, Beaufort Group.

Biostratigraphy: Lystrosaurus Assemblage zone (Lystrosaurus-Thrinaxodon zone, KEYSER, 1979, SACS, 1980, Procolophon zone, Lystrosaurus zone, WATSON, 1914, BROOM, 1906).

Age: Early Scythian Epoch, Early Triassic.

Material: Skull.

Referred material:

CLUVER, 1971

Locality: Cradock District, Eastern Cape Province, South Africa.

Horizon: Katberg Formation, Beaufort Group.

Biostratigraphy: Lystrosaurus Assemblage zone (Lystrosaurus-Thrinaxodon zone, KEYSER, 1979, SACS, 1980, Procolophon zone, Lystrosaurus zone, WATSON, 1914, BROOM, 1906).

Age: Early Scythian Epoch, Early Triassic.

Material:

SAM 706, TM R27: Skulls.

SAM-PK-7843: Skull.

 

RAY, 2005

Locality: Middleburg District, Eastern Cape Province, South Africa.

Horizon: Katberg Formation, Beaufort Group.

Biostratigraphy: Lystrosaurus Assemblage zone (Lystrosaurus-Thrinaxodon zone, KEYSER, 1979, SACS, 1980, Procolophon zone, Lystrosaurus zone, WATSON, 1914, BROOM, 1906).

Age: Early Scythian Epoch, Early Triassic.

Material:

SAM-PK-K4, K1389: Partial skulls.

= Lystrosaurus andersoni BROOM, 1907
Etymology:

Holotype: Natal Museum

Locality: Umkomazaan River, Western KwaZulu Natal Province, South Africa.

Horizon: Katberg Formation, Beaufort Group.

Biostratigraphy: Lystrosaurus Assemblage zone (Lystrosaurus-Thrinaxodon zone, KEYSER, 1979, SACS, 1980, Procolophon zone, Lystrosaurus zone, WATSON, 1914, BROOM, 1906).

Age: Early Scythian Epoch, Early Triassic.

Material: Skull.

= Lystrosaurus oviceps HAUGHTON, 1915
Etymology:

Holotype: SAM 641

Locality: "Tarka River, Cradock District", Eastern Cape Province, South Africa.

Horizon: Katberg Formation, Beaufort Group.

Biostratigraphy: Lystrosaurus Assemblage zone (Lystrosaurus-Thrinaxodon zone, KEYSER, 1979, SACS, 1980, Procolophon zone, Lystrosaurus zone, WATSON, 1914, BROOM, 1906).

Age: Early Scythian Epoch, Early Triassic.

Material: Fragmentary skull and dentary.

Modified from Colbert, 1974.

Referred material:

BRINK, 1951

Locality: Unknown, South Africa.

Horizon: Katberg Formation, Beaufort Group.

Biostratigraphy: Lystrosaurus Assemblage zone (Lystrosaurus-Thrinaxodon zone, KEYSER, 1979, SACS, 1980, Procolophon zone, Lystrosaurus zone, WATSON, 1914, BROOM, 1906).

Age: Early Scythian Epoch, Early Triassic.

Material:

BP M103: Skull and lower jaw.

NMB C299: Skull.

 

Locality: Vlakfontein, Middleburg District, Eastern Cape Province, South Africa.

Horizon: Katberg Formation, Beaufort Group.

Biostratigraphy: Lystrosaurus Assemblage zone (Lystrosaurus-Thrinaxodon zone, KEYSER, 1979, SACS, 1980, Procolophon zone, Lystrosaurus zone, WATSON, 1914, BROOM, 1906).

Age: Early Scythian Epoch, Early Triassic.

Material:

BP M108: Skull and lower jaw.

 

Locality: Edenvale, Middleburg District, Eastern Cape Province, South Africa.

Horizon: Katberg Formation, Beaufort Group.

Biostratigraphy: Lystrosaurus Assemblage zone (Lystrosaurus-Thrinaxodon zone, KEYSER, 1979, SACS, 1980, Procolophon zone, Lystrosaurus zone, WATSON, 1914, BROOM, 1906).

Age: Early Scythian Epoch, Early Triassic.

Material:

BP M110: Skull.

 

Locality: Honingkrans, Burgsersdorp, Albert District, Eastern Cape Province, South Africa.

Horizon: Katberg Formation, Beaufort Group.

Biostratigraphy: Lystrosaurus Assemblage zone (Lystrosaurus-Thrinaxodon zone, KEYSER, 1979, SACS, 1980, Procolophon zone, Lystrosaurus zone, WATSON, 1914, BROOM, 1906).

Age: Early Scythian Epoch, Early Triassic.

Material:

BP M114: Skull and lower jaw.

 

RAY, 2005

Locality: Harrismith, Free (Orange Free) State, South Africa.

Horizon: Katberg Formation, Beaufort Group.

Biostratigraphy: Lystrosaurus Assemblage zone (Lystrosaurus-Thrinaxodon zone, KEYSER, 1979, SACS, 1980, Procolophon zone, Lystrosaurus zone, WATSON, 1914, BROOM, 1906).

Age: Early Scythian Epoch, Early Triassic.

Material:

SAM-PK-1403: Partial skull and lower jaw.

 

Locality: Middleburg District, Eastern Cape Province, South Africa.

Horizon: Katberg Formation, Beaufort Group.

Biostratigraphy: Lystrosaurus Assemblage zone (Lystrosaurus-Thrinaxodon zone, KEYSER, 1979, SACS, 1980, Procolophon zone, Lystrosaurus zone, WATSON, 1914, BROOM, 1906).

Age: Early Scythian Epoch, Early Triassic.

Material:

SAM-PK-K1369: Skull and lower jaw.

 

Locality: Unknown, South Africa.

Horizon: Katberg Formation, Beaufort Group.

Biostratigraphy: Lystrosaurus Assemblage zone (Lystrosaurus-Thrinaxodon zone, KEYSER, 1979, SACS, 1980, Procolophon zone, Lystrosaurus zone, WATSON, 1914, BROOM, 1906).

Age: Early Scythian Epoch, Early Triassic.

Material:

SAM-PK-K1273: Skull and lower jaw.

 

= Ptychognathus pusillius OWEN, 1876
Etymology:

Holotype: BMNH R1691

Locality: Cape Colony, South Africa.

Horizon:

Biostratigraphy:

Age: Early Triassic.

Material: 6 lumbar vertebrae and a ilia.

 

= Lystrosaurus breyeri van HOEPEN, 1916
Etymology:

Holotype: Transvall Museum 19

Locality: Harrismith, Free (Orange Free) State, South Africa.

Horizon: Katberg Formation, Beaufort Group.

Biostratigraphy: Lystrosaurus Assemblage zone (Lystrosaurus-Thrinaxodon zone, KEYSER, 1979, SACS, 1980, Procolophon zone, Lystrosaurus zone, WATSON, 1914, BROOM, 1906).

Age: Early Scythian Epoch, Early Triassic.

Material: Skull.

_____________________________________________________________________________________

Species: mccaigi (SEELEY, 1898)
Etymology:
= Rhabdocephalus mccaigi SEELEY, 1898
= Lystrosaurus maccaigi BOTHA-BRINK, HUTTENLOCKER, & MODESTO, 2014 (sic)

Holotype: AMNH 404

Locality: Elandsberg, Cradock District, Eastern Cape Province, South Africa.

Horizon: Katberg Formation, Beaufort Group.

Biostratigraphy: Lystrosaurus Assemblage zone (Lystrosaurus-Thrinaxodon zone, KEYSER, 1979, SACS, 1980, Procolophon zone, Lystrosaurus zone, WATSON, 1914, BROOM, 1906).

Age: Early Scythian Epoch, Early Triassic.

Material: A partial skull.

Modified from Colbert, 1974.

Referred material:

Locality:

Horizon:

Biostratigraphy:

Age:

Material:

NMC 228:

 

 

Locality:

Horizon:

Biostratigraphy:

Age:

Material:

BP 380:

 

Locality: South Africa.

Horizon: Katberg Formation, Beaufort Group.

Biostratigraphy: Lystrosaurus Assemblage zone (Lystrosaurus-Thrinaxodon zone, KEYSER, 1979, SACS, 1980, Procolophon zone, Lystrosaurus zone, WATSON, 1914, BROOM, 1906).

Age: Early Scythian Epoch, Early Triassic.

Material:

SAM K116: Skull.

 

Locality:

Horizon:

Biostratigraphy:

Age:

Material:

BP 879:

BP 3972:

BP 4052:

BP 4053:

BP 4368:

 

Locality: Wilgerbosch, near New Bethesda, Graaff Reinet District, Eastern Cape Province, South Africa.

Horizon: Katberg Formation, Beaufort Group.

Biostratigraphy: Lystrosaurus Assemblage zone (Lystrosaurus-Thrinaxodon zone, KEYSER, 1979, SACS, 1980, Procolophon zone, Lystrosaurus zone, WATSON, 1914, BROOM, 1906).

Age: Early Scythian Epoch, Early Triassic.

Material:

UCMP 42707:

 

 

Locality: Harrismith, Free (Orange Free) State, South Africa.

Horizon: Katberg Formation, Beaufort Group.

Biostratigraphy: Lystrosaurus Assemblage zone (Lystrosaurus-Thrinaxodon zone, KEYSER, 1979, SACS, 1980, Procolophon zone, Lystrosaurus zone, WATSON, 1914, BROOM, 1906).

Age: Early Scythian Epoch, Early Triassic.

Material:

UCMP 124858:

 

ABDALA, GAETANO, SMITH, & RUBIDGE, 2019

Locality: Ripplemead Farm, near Nieu Bethesda, New Bethesda, Graaff Reinet District, Eastern Cape Province, South Africa.

Horizon: Elandsberg Member, Balfour Formation, Beaufort Group.

Biostratigraphy: Dicynodon Assemblage zone (formerly Dicynodon lacerticeps-Whaitsia zone of KEYSER, 1979, SACS, 1980, Daptocephalus zone of KITCHING, 1870, 1977, Upper Cistecephalus Zone of WATSON, 1914).

Age: Tatarian Age, Lopingian Subepoch, Zechstein Epoch, Late Permian.

Material:

Field number RS 440, SAM-PK-K10690: Skull and postcranial remains.

BOTHA-BRINK, HUTTENLOCKER, & MODESTO, 2014

Locality: Nooitgedacht 68, Bethulie District,  Eastern Cape Province, South Africa.

Horizon: Balfour Formation.

Biostratigraphy: Dicynodon Assemblage Zone.

Age: Tatarian Age, Lopingian Subepoch, Zechstein Epoch, Late Permian.

Material:

BP/1/3972: Skull.

NMQR 1637, 1639, 1640a, 1641, 1643, 1648, 1650, 1653, 3208, 3641, 3642, 3646, 3647, 3648, 3658, 3663, 3684a, 3687, 3689, 3690, 3693, 3694, 3695, 3699, 3700, 3702, 3703, 3705, 3706, 3708, 3711, 3712, 3713, 3919, 3922a, 3934, 3935, 3936, 3938, 3940: Skulls and associated skeletons in varying stagesof completeness and at various ontogenetic stages (Lowermost slopes of Loskop and Spitskop).

 

COLBERT,

Locality: Shenk Peak, Cumulus Hills of Shackleton Glacier area, 28 m about formation base, Queen Maud Mountains, East Antarctica, Antarctica.

Horizon: Fremouv Formation.

Biostratigraphy:

Age: Anisian Stage, Lower Mid Triassic Epoch, Early Middle Triassic.

Material:

WSU 0974: Front half of a large skull.

 

 

Locality: Thrinaxodon Col, Cumulus Hills of Shackleton Glacier, Queen Maud Mountains, East Antarctica, Antarctica.

Horizon: Fremouv Formation.

Biostratigraphy:

Age: Anisian Stage, Lower Mid Triassic Epoch, Early Middle Triassic.

Material:

AMNH 9505: Fragment of right maxilla with tusks and fragments of adjoining bones.

 

= Lystrosaurus putterilli van HOEPEN, 1915
Etymology:

Holotype: TM R44 (according to HAUGHTON & BRINK, 1954, TM 196)

Locality: Harrismith, Free (Orange Free) State, South Africa.

Horizon: Katberg Formation, Beaufort Group.

Biostratigraphy: Lystrosaurus Assemblage zone (Lystrosaurus-Thrinaxodon zone, KEYSER, 1979, SACS, 1980, Procolophon zone, Lystrosaurus zone, WATSON, 1914, BROOM, 1906).

Age: Early Scythian Epoch, Early Triassic.

Material: Skull and fragmentary skeleton.

Referred material:

TM R47: Skull.

 

= Lystrosaurus amphibius BRINK, 1951
Etymology:

Holotype: BP 136

Locality: Ripplemead, New Bethesda, Graaff Reinet District, Eastern Cape Province, South Africa.

Horizon: Katberg Formation, Beaufort Group.

Biostratigraphy: Lystrosaurus Assemblage zone (Lystrosaurus-Thrinaxodon zone, KEYSER, 1979, SACS, 1980, Procolophon zone, Lystrosaurus zone, WATSON, 1914, BROOM, 1906).

Age: Early Scythian Epoch, Early Triassic.

Material: Skull.

_____________________________________________________________________________________

Species:? seeleyi BROILI, 1907 (nomen dubium)
Etymology: In honor of H. G. Seeley.
= Dicynodon seeleyi (BROILI, 1907) (nomen dubium)

Holotype:

Locality:

Horizon:

Biostratigraphy:

Age:

Material:
_____________________________________________________________________________________

Species: hedini YOUNG, 1935
Etymology:

Holotype: Field No. 5000010, IVPP RV 3012

Locality: Hungshankou, east of the river, Talungkou, South Santai, Xinjiang Uygur Zizhiqu (Sinkang) Province, China.

Horizon: Jiucaiyuan Formation.

Biostratigraphy:

Age: Early Triassic.

Material: Skull and nearly complete skeleton.

Modified from Young, 1935

Referred material:

Locality: Hungshanting, Hotung, Tatungkou, South Santai, Fuyuan, Xinjiang Uygur Zizhiqu (Sinkang) Province, China.

Horizon: Jiucaiyuan Formation.

Biostratigraphy:

Age: Early Triassic.

Material:

Field No. 600067: Fragmentary skeleton.

 

GUBIN & SINITZA, 1993

Locality: The Canyon of Sain-Sar-Bulak Spring, 28 km westward from Noyan Soman, Mongolia.

Horizon: Dalin Shandakhuduk Formation.

Biostratigraphy:

Age: Early Triassic.

Material:

Number: Not given: 3 skeletons and 2 skulls.

 

YOUNG 1939

Locality: Xinjiang Uygur Zizhiqu (Sinkang) Province, China.

Horizon:

Biostratigraphy:

Age: Early Triassic.

Material:

Field no. 50.001: 2 distal part of radius, 2 femora, 2 tibiae and some other fragments.

Field no. 50.002: Distal part of a scapula.

 

Locality: Xinjiang Uygur Zizhiqu (Sinkang) Province, China.

Horizon:

Biostratigraphy:

Age: Early Triassic.

Material:

Field no. 500.010: Vertebrae, right coracoid and precoracoid, proximal part of left ilium, right ischium, proximal part of a right femur, proximal part of a tibia, phalanges.

= Dicynodon broomi YOUNG, 1939
Etymology: In honor of South African palaeontologist R. L. Broom
= Lystrosaurus broomi YOUNG, 1939

Holotype: No. 600065, IVPP RV 39060

Locality: Junggar and Turpan Basin, Xinjiang Uygur Zizhiqu (Sinkang) Province, China.

Horizon: Jiucaiyuan Formation, Early Triassic.

Material: An incomplete skull.

Lystrosaurus broomi modified from YOUNG, 1939

= Lystrosaurus latifrons SUN, 1973
Etymology: Latin, lati, "broad, wide" and Latin, frons, "brow, forehead."

Holotype: IVPP V3244

Locality: Turfan, south foot of the Tienshan mountains, north margin of the Turfan Basin, Xinjiang Uygur Zizhiqu (Sinkang) Province, China.

Horizon: Third fossiliferous stratum of Taoshuyuanzi.

Biostratigraphy:

Age: Early Triassic.

Material: Skull lacking tip of snout and right postorbital with left squamosal damaged.

Referred material:

IVPP V3247: Fragmentary skull.

= Lystrosaurus robustus SUN, 1973
Etymology: Latin, robustus, "robust."

Holotype: IVPP V3243

Locality: Turfan South foot of the Tienshan mountains, north margin of the Turfan Basin, Xinjiang Uygur Zizhiqu (Sinkang) Province, China.

Horizon: Third fossiliferous stratum of Taoshuyuanzi.

Biostratigraphy:

Age: Early Triassic.

Material: Skull and lower jaw.

Referred material:

IVPP V3246: Skull and lower jaw.

 

= Lystrosaurus shichanggouensis CHENG, 1986
Etymology: In reference to Shichanggou, Jimsar, Xinjiang Uygur Zizhiqu (Sinkang) Province, China.

Holotype: CAGS V 321

Locality: Shichanggou, Jimsar, Xinjiang Uygur Zizhiqu (Sinkang) Province, China.

Horizon: Jiucaiyuan Formation.

Biostratigraphy:

Age: Early Triassic.

Material: Skull, lower jaw and incomplete postcranial skeleton.

_____________________________________________________________________________________

Species: weidenreichi YOUNG, 1939
Etymology:

Holotype: Field Numbers 10.164-10.165, 1928, IVPP RV 39061

Locality: Locality 24, Sinkiang, Junggar Basin, Xinjiang Uygur Zizhiqu (Sinkang) Province, China.

Horizon: Jiucaiyuan Formation.

Biostratigraphy:

Age: Early Triassic.

Material: An incomplete postcranial skeleton, including vertebrae, ribs, a left scapula, 2 ilia, a left femur, tibia and fibula.

Referred material:

Locality: Xinjiang Uygur Zizhiqu (Sinkang) Province, China.

Horizon:

Biostratigraphy:

Age: Early Triassic.

Material:

Field no: Left squamosal, 8 vertebrae, incomplete sternum, ischium, pubis, 2 astragli, calcaneum, a number of phalanges and some undeterminable fragments.

 

Locality: Tatungkou, Fuyuanhsien, Xinjiang Uygur Zizhiqu (Sinkang) Province, China.

Horizon:

Biostratigraphy:

Age: Early Triassic.

Material:

Number: Not given: 5 vertebrae, several ribs, scapula, right coracoid and a fragment of precoracoid, sternum, left humerus, right pubis, left and right tibia, and a few phalanges.

 

Locality: Locality 14, Xinjiang Uygur Zizhiqu (Sinkang) Province, China.

Horizon:

Biostratigraphy:

Age: Early Triassic.

Material:

Field numbers: 10.135-36, 10.145, 10.161-62: 15 more or less complete vertebrae, parts of a pelvic girdle, a fragmentary femur, tibia, fibula, tarsali, metatarsalia and phalanges.

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Species: youngi SUN, 1964
Etymology: In honor of C. C. Young.

Holotype: IVPP V 8532, 63005-4

Locality: Tung-Hsiao-Lung-Kou, Jimusar, Xinjiang Uygur Zizhiqu (Sinkang) Province, China.

Horizon: Jiiucaiyuan Formation.

Biostratigraphy:

Age: Early Triassic.

Material: Skull and dentary.

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After Sun, 1964

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Species: georgi KALANDADZE, 1975
Etymology:

Holotype: PIN 3447/1

Locality: Astashikinskoyi I, Vetluga River, Gorkiy Province, European Russia.

Horizon: Ryabinian horizon.

Biostratigraphy:

Age: Indian Stage, Early Triassic.

Material: Anterior part of the skull, axis, 2 cervicals, 10 dorsals, 2 sacrals, 4 caudals, 9 proximal and some distal fragments of dorsal ribs, the most proximal part of the right scapula, a small fragment of the left coracoid, right humerus and ulna, most of the right ilium and all of the left, and the proximal part of the right ischium, right and left femora, both tibiae, the complete left fibula and distal part of the right fibula, and several scatterd bones of the foot--1 astragalus, 4 metapodials (2 of them partly destroyed), 1 phalanx, 3 pre-ungular phalanges, and 5 ungular phalanges.

Modified from KALANDADZE, 1975.
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