Genus: Rhachiocephalus SEELEY, 1898
Etymology: Greek, rhachis, “spine, ridge”, Greek, kephale, “head”, + us: ridged head.
= Eocyclops BROOM, 1913
Etymology: Greek, eos, “dawn”, Greek, kyklops, “round eye”: dawn cyclops.
= Kitchingia BROOM & GEORGE, 1950
Etymology: In honor of the Kitching family (James, Croonie, and Croonie’s sons), who collected fossils for Broom.
= Neomegacyclops BROOM, 1931
Etymology: Greek, neos, “new”, and Megacyclops: New Megacyclops.
= Pelorocyclops van HOEPEN, 1934
Etymology:
= Platycyclops BROOM, 1932
Etymology: Greek, platys, “broad, flat”, Greek, kyklops, “round eye”: Flat cyclops.
= Platypodosaurus OWEN, 1880
Etymology: Greek, platys, “broad, flat”, Greek, pod- (pous), “foot”, Greek, sauros, “lizard”: Broad foot lizard.

Species: magnus (OWEN, 1876) SEELEY, 1898
Etymology: Latin, magnus, "great, large."
= Udenodon magnus OWEN, 1876
= Platypodosaurus magnus (OWEN, 1876)
= Oudenodon magnus (OWEN, 1876)

Holotype: BMNH 36252

Locality: Brak river, near Beaufort West District, Western Cape Province, South Africa.

Horizon:

Biostratigraphy: Cistecephalus zone (formerly Auacephalodon-Cistecephalus zone KEYSER, 1979, SACS, 1980, Aulacephalodon baini zone KEYSER & SMITH, 1977-78, Upper Cistecephalus Zone, KITCHING, 1970, 77, Lower Cistecephalus zone WATSON, 1914).

Age: Tatarian Age, Lopingian Subepoch, Zechstein Epoch, Late Permian.

Material: Fragmentary skull.

Referred material:

BMNH 36237: Fragmentary mandible.

 

DAY & RUBIDGE, 2019

Locality: Nobelsfontien 248, south of Biesiespoort station, Victoria West District, North Cape West Province, South Africa.

Horizon: Hoedemaker Member, Teekloof Formation, Beaufort Group.

Biostratigraphy: Upper Tropidostoma Zone.

Age: Tatarian Age, Lopingian Subepoch, Zechstein Epoch, Late Permian.

Material:

BP/1/8149:

 

HUENE, 1942

Locality: Ruhuhu, Tanzania, East Africa.

Horizon:

Biostratigraphy:

Age:

Material:

K 15: Skull.

K 100: Skull of a juvenile.

 


BRINK, 1982

Locality: Waterfalls Valley, Graaff Reinet District, Eastern Cape Province, South Africa.

Horizon:

Biostratigraphy: Cistecephalus zone (formerly Auacephalodon-Cistecephalus zone KEYSER, 1979, SACS, 1980, Aulacephalodon baini zone KEYSER & SMITH, 1977-78, Upper Cistecephalus Zone, KITCHING, 1970, 77, Lower Cistecephalus zone WATSON, 1914).

Age: Tatarian Age, Lopingian Subepoch, Zechstein Epoch, Late Permian.

Material:

GSO R58: Skull with part of lower jaw.

 

 

Locality: Knoppiesfontein, Bethulie District, Free (Orange Free) State, South Africa.

Horizon:

Biostratigraphy: Cistecephalus zone (formerly Auacephalodon-Cistecephalus zone KEYSER, 1979, SACS, 1980, Aulacephalodon baini zone KEYSER & SMITH, 1977-78, Upper Cistecephalus Zone, KITCHING, 1970, 77, Lower Cistecephalus zone WATSON, 1914).

Age: Tatarian Age, Lopingian Subepoch, Zechstein Epoch, Late Permian.

Material:

BP F815: Weathered skull.

 

Locality: Aasvoelokrans, Murraysburg District, Western Cape Province, South Africa.

Horizon:

Biostratigraphy: Cistecephalus zone (formerly Auacephalodon-Cistecephalus zone KEYSER, 1979, SACS, 1980, Aulacephalodon baini zone KEYSER & SMITH, 1977-78, Upper Cistecephalus Zone, KITCHING, 1970, 77, Lower Cistecephalus zone WATSON, 1914).

Age: Tatarian Age, Lopingian Subepoch, Zechstein Epoch, Late Permian.

Material:

BP F1512: Skull.

 


Locality: Klipplaat, Richmond District, Free (Orange Free) State, South Africa.

Horizon:

Biostratigraphy: Cistecephalus zone (formerly Auacephalodon-Cistecephalus zone KEYSER, 1979, SACS, 1980, Aulacephalodon baini zone KEYSER & SMITH, 1977-78, Upper Cistecephalus Zone, KITCHING, 1970, 77, Lower Cistecephalus zone WATSON, 1914).

Age: Tatarian Age, Lopingian Subepoch, Zechstein Epoch, Late Permian.

Material:

BP F2458: Complete skull.

 


Locality: Beeldhouersfontein, Murryasburg District, Republic of South Africa.

Horizon:

Biostratigraphy: Cistecephalus zone (formerly Auacephalodon-Cistecephalus zone KEYSER, 1979, SACS, 1980, Aulacephalodon baini zone KEYSER & SMITH, 1977-78, Upper Cistecephalus Zone, KITCHING, 1970, 77, Lower Cistecephalus zone WATSON, 1914).

Age: Tatarian Age, Lopingian Subepoch, Zechstein Epoch, Late Permian.

Material:

BP F2889: Almost complete skull.

 

 

Locality: Agtertang, Colesberg District, Northern Cape Province, South Africa.

Horizon:

Biostratigraphy: Cistecephalus zone (formerly Auacephalodon-Cistecephalus zone KEYSER, 1979, SACS, 1980, Aulacephalodon baini zone KEYSER & SMITH, 1977-78, Upper Cistecephalus Zone, KITCHING, 1970, 77, Lower Cistecephalus zone WATSON, 1914).

Age: Tatarian Age, Lopingian Subepoch, Zechstein Epoch, Late Permian.

Material:

NMB VDK47: Skull lacking left zyogomatic arch and occiput.

NMB VDK48: Skull.

NMB VDK50: Skull lacking arches.

 

 

Locality: T’samekoms, Colesberg District, Northern Cape Province, South Africa.

Horizon:

Biostratigraphy: Cistecephalus zone (formerly Auacephalodon-Cistecephalus zone KEYSER, 1979, SACS, 1980, Aulacephalodon baini zone KEYSER & SMITH, 1977-78, Upper Cistecephalus Zone, KITCHING, 1970, 77, Lower Cistecephalus zone WATSON, 1914).

Age: Tatarian Age, Lopingian Subepoch, Zechstein Epoch, Late Permian.

Material:

NMB VDK73: Skull lacking right zygomatic arch.

 


= Platypodosaurus robustus OWEN, 1880
Etymology: Latin, robustus, "robust."

Holotype: BMNH 50121

Locality: Catelomds, Claremont, South Africa.

Horizon:

Biostratigraphy:

Age: Late Permian.

Material: Fragmentary skeleton.

Referred material:

BMNH 50122: Right scapula with the terminal phalangeals of the four digits of the manus.

BMNH 50123: Proximal end of right femur.

BMNH 50124: Fragmentary tibia.

BMNH 50125: Presternum?

BMNH 50126: Fragmentary vertebra.

 


LYDEKKER, 1890

Locality: Near Fort Beaufort, Beaufort West District, Western Cape Province, South Africa.

Horizon:

Biostratigraphy:

Age: Late Permian.

Material:

BMNH 47090: Sacrum and fragmentary pelvis and 10 caudal vertebrae.

 


= Rhachiocephalus giganteus (van HOEPEN, 1934)
Etymology: Latin, giganteus, "huge, gigantic."
= Pelorocyclops giganteus van HOEPEN, 1934

Holotype: National Museum

Locality: South Africa.

Horizon:

Biostratigraphy: Cistecephalus zone (formerly Auacephalodon-Cistecephalus zone KEYSER, 1979, SACS, 1980, Aulacephalodon baini zone KEYSER & SMITH, 1977-78, Upper Cistecephalus Zone, KITCHING, 1970, 77, Lower Cistecephalus zone WATSON, 1914).

Age: Tatarian Age, Lopingian Subepoch, Zechstein Epoch, Late Permian.

Material: Skull.

 


= Rhachiocephalus angusticeps (BROOM, 1937)
Etymology: Latin, angustus, "narrow" and Latin, -ceps, "head."
= Platypodosaurus angusticeps BROOM, 1937

Holotype: Transvaal Museum 1487

Locality: A few miles east of Mazelspoort, Orange Free State, South Africa.

Horizon:

Biostratigraphy: Cistecephalus zone (formerly Auacephalodon-Cistecephalus zone KEYSER, 1979, SACS, 1980, Aulacephalodon baini zone KEYSER & SMITH, 1977-78, Upper Cistecephalus Zone, KITCHING, 1970, 77, Lower Cistecephalus zone WATSON, 1914).

Age: Tatarian Age, Lopingian Subepoch, Zechstein Epoch, Late Permian.

Material: Skull missing posterior end.

 

= Dicynodon scheepersi BROOM, 1948
Etymology:

Holotype: RC 95

Locality: Third Dirft, Adendorp, Graaff Reinet District, Eastern Cape Province, South Africa.

Horizon:

Biostratigraphy: Cistecephalus zone (formerly Auacephalodon-Cistecephalus zone KEYSER, 1979, SACS, 1980, Aulacephalodon baini zone KEYSER & SMITH, 1977-78, Upper Cistecephalus Zone, KITCHING, 1970, 77, Lower Cistecephalus zone WATSON, 1914).

Age: Tatarian Age, Lopingian Subepoch, Zechstein Epoch, Late Permian.

Material: Skull.

= Kitchingia planifrons BROOM & GEORGE, 1950
Etymology: Latin, plani, “flat’ and Latin, frons, "brow, forehead."

Holotype: BP 124

Locality: Brandkraal, Murraysburg District, Western Cape Province, South Africa.

Horizon:

Biostratigraphy: Cistecephalus zone (formerly Auacephalodon-Cistecephalus zone KEYSER, 1979, SACS, 1980, Aulacephalodon baini zone KEYSER & SMITH, 1977-78, Upper Cistecephalus Zone, KITCHING, 1970, 77, Lower Cistecephalus zone WATSON, 1914).

Age: Tatarian Age, Lopingian Subepoch, Zechstein Epoch, Late Permian.

Material: A large skull.

_____________________________________________________________________________________

Species: longus (BROOM, 1913)
Etymology: Latin, longus, "long."
= Eocyclops longus BROOM, 1913

Holotype: AMNH

Locality: Grootvlei (part of Paardekraal), Beaufort West District, Western Cape Province, South Africa.

Horizon:

Biostratigraphy: Cistecephalus zone (formerly Auacephalodon-Cistecephalus zone KEYSER, 1979, SACS, 1980, Aulacephalodon baini zone KEYSER & SMITH, 1977-78, Upper Cistecephalus Zone, KITCHING, 1970, 77, Lower Cistecephalus zone WATSON, 1914).

Age: Tatarian Age, Lopingian Subepoch, Zechstein Epoch, Late Permian.

Material: Skull.

Referred material:

Locality: Graaff Reinet District, Eastern Cape Province, South Africa.

Horizon:

Biostratigraphy:

Age: Late Permian.

Material:

SAM 3425: Skull.

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Species: whaitsi (BROOM, 1931)
Etymology: In honor of Rev. John H. Whaits (1870-1947).
= Megacyclops whaitsi BROOM, 1931
= Neomegacyclops whaitsi (BROOM, 1931)
= Eocyclops longus HAUGHTON, 1917 non BROOM, 1913

Holotype: SAM 3425

Locality: Graaff Reinet District, Eastern Cape Province, South Africa.

Horizon: Hoedemaker member, Middle Teekloof Formation.

Biostratigraphy: Tropidostoma zone (Tropidostoma-Endothidon Assemblage zone KEYSER, 1979, SACS, 1980, Tropidostoma microtrema zone KEYSER & SMITH, 1977-78, Lower Cistecephalus zone KITCHING, 1970, 1977, Endothiodon zone, WATSON, 1914).

Age: Tatarian Age, Lopingian Subepoch, Zechstein Epoch, Late Permian.

Material: Skull.
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Species: rugosus (HAUGHTON, 1932)
Etymology: Latin, rugosus, "wrinkled."
= Megacyclops rugosus HAUGHTON, 1932
= Neomegacyclops rugosus (HAUGHTON, 1932)
= Platypodosaurus rugosus (HAUGHTON, 1932)

Holotype: SAM 10637 (Stockely number S266)

Locality: B4, Katumba Viwilli, Tanganyika, Tanzania, East Africa.

Horizon:

Biostratigraphy: Cistecephalus zone (formerly Auacephalodon-Cistecephalus zone KEYSER, 1979, SACS, 1980, Aulacephalodon baini zone KEYSER & SMITH, 1977-78, Upper Cistecephalus Zone, KITCHING, 1970, 77, Lower Cistecephalus zone WATSON, 1914).

Age: Tatarian Age, Lopingian Subepoch, Zechstein Epoch, Late Permian.

Material: Snout, lower jaw, femur.
Note: Rhachiocephalus per GAY & CRUICKSHANK, 1999.
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Species: haughtoni (BROOM, 1932)
Etymology: In honor of Sidney Henry Haughton (1888-1982), British Paleontologist
= Platycyclops haughtoni BROOM, 1932

Holotype: SAM 8750

Locality: Doornplass, Graaff Reinet District, Eastern Cape Province, South Africa.

Horizon:

Biostratigraphy: Cistecephalus zone (formerly Auacephalodon-Cistecephalus zone KEYSER, 1979, SACS, 1980, Aulacephalodon baini zone KEYSER & SMITH, 1977-78, Upper Cistecephalus Zone, KITCHING, 1970, 77, Lower Cistecephalus zone WATSON, 1914).

Age: Tatarian Age, Lopingian Subepoch, Zechstein Epoch, Late Permian.

Material: Skull.

Referred material:

= Platycyclops latirhinus BROOM, 1940
Etymology: Latin, lati, "broad, wide" and Greek, rhinus, “nose, snout.”

Holotype: RC 25

Locality: Wimbledon, Graaff Reinet District, Eastern Cape Province, South Africa.

Horizon:

Biostratigraphy: Cistecephalus zone (formerly Auacephalodon-Cistecephalus zone KEYSER, 1979, SACS, 1980, Aulacephalodon baini zone KEYSER & SMITH, 1977-78, Upper Cistecephalus Zone, KITCHING, 1970, 77, Lower Cistecephalus zone WATSON, 1914).

Age: Tatarian Age, Lopingian Subepoch, Zechstein Epoch, Late Permian.

Material: Anterior 2/3rds of skull.

 

= Platycyclops richardi BROOM, 1948
Etymology:

Holotype: RC 66

Locality: Woodcliffe, Graaff Reinet District, Eastern Cape Province, South Africa.

Horizon:

Biostratigraphy: Cistecephalus zone (formerly Auacephalodon-Cistecephalus zone KEYSER, 1979, SACS, 1980, Aulacephalodon baini zone KEYSER & SMITH, 1977-78, Upper Cistecephalus Zone, KITCHING, 1970, 77, Lower Cistecephalus zone WATSON, 1914).

Age: Tatarian Age, Lopingian Subepoch, Zechstein Epoch, Late Permian.

Material: Anterior 2/3rds of a skull.

 


= Platycyclops rubidgei BROOM, 1948
Etymology: In honor of Dr. S. H. Rubidge.

Holotype: RC 26

Locality: Grootdriefontein, Murraysburg District, Western Cape Province, South Africa.

Horizon:

Biostratigraphy: Cistecephalus zone (formerly Auacephalodon-Cistecephalus zone KEYSER, 1979, SACS, 1980, Aulacephalodon baini zone KEYSER & SMITH, 1977-78, Upper Cistecephalus Zone, KITCHING, 1970, 77, Lower Cistecephalus zone WATSON, 1914).

Age: Tatarian Age, Lopingian Subepoch, Zechstein Epoch, Late Permian.

Material: Skull.

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Species: usiliensis (HUENE, 1942) CLUVER & KING, 1983
Etymology: In reference to Usili Mountain, Ngaka, Ruhuhu River, Tanzania, Tanganyika, East Africa.
= Megacyclops usiliensis HUENE, 1942
= Neomegacyclops usiliensis (HUENE, 1942) BOOSTRA, 1953

Holotype: GPIT U 31

Locality: Usili Mountain, Ngaka, Ruhuhu River, Tanzania, Tanganyika, East Africa.

Horizon: Kawinga Formation.

Biostratigraphy: Cistecephalus zone (formerly Auacephalodon-Cistecephalus zone KEYSER, 1979, SACS, 1980, Aulacephalodon baini zone KEYSER & SMITH, 1977-78, Upper Cistecephalus Zone, KITCHING, 1970, 77, Lower Cistecephalus zone WATSON, 1914).

Age: Tatarian Age, Lopingian Subepoch, Zechstein Epoch, Late Permian.

Material: Skull missing posterior end.
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Species: wesselsi (BROOM, 1948)
Etymology:
= Pelanomodon wesselsi BROOM, 1948

Holotype: RC 65

Locality: Murraysburg District, Western Cape Province, South Africa.

Horizon:

Biostratigraphy: Cistecephalus zone (formerly Auacephalodon-Cistecephalus zone KEYSER, 1979, SACS, 1980, Aulacephalodon baini zone KEYSER & SMITH, 1977-78, Upper Cistecephalus Zone, KITCHING, 1970, 77, Lower Cistecephalus zone WATSON, 1914).

Age: Tatarian Age, Lopingian Subepoch, Zechstein Epoch, Late Permian.

Material: Skull.
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Species: cyclops BOONSTRA, 1953
Etymology:
= Neomegacyclops cyclops BOONSTRA, 1953

Holotype: SAM 11707

Locality: Ruhuhu Coalfield region, Ngaka-Kingori Hill area, Tanzania, Tanganyika, East Africa.

Horizon:

Biostratigraphy: Cistecephalus zone (formerly Auacephalodon-Cistecephalus zone KEYSER, 1979, SACS, 1980, Aulacephalodon baini zone KEYSER & SMITH, 1977-78, Upper Cistecephalus Zone, KITCHING, 1970, 77, Lower Cistecephalus zone WATSON, 1914).

Age: Tatarian Age, Lopingian Subepoch, Zechstein Epoch, Late Permian.

Material: Fragmentary skull.

Dorsal and lateral view of the pineal region afer Boonstra, 1953.

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Species: behemoth MAISCH, 2005
Etymology:

Holotype: GPIT K 15

Locality: Kingori Mountain, about 1000 m west of the northern peak, Ngaka area, Ruhuhu Basin, Southwestern Tanzania.

Horizon: Kawinga Formation.

Biostratigraphy:

Age: Tatarian Age, Lopingian Subepoch, Zechstein Epoch, Late Permian.

Material: Skull.

Referred material:

GPIT K 15 B: Fragmenary skullcomprising most of the interorbital skull roof and the pineal region.