Species: armatus MARSH, 1877
Etymology: Latin, armatura, 'arms, equipment'.

Holotype: YPM 1850

Locality: Lakes' Yale Quarry 5, Morrison, Jefferson County, Colorado.

Horizon: Upper Brushy Basin Member, Morrison Formation.

Biostratigraphy: Zone 4.

Age: Comobluffian age, Upper Tithonian Stage, Uppermost Malm Epoch, Late Jurassic.

Material: Skull and skeleton that is still in a plaster jacket.

Skeleton and reconstruction.

Plate.

Referred specimens:

USNM 4936: Posterior half of a skull with braincase.

 

= cf. Stegosaurus armatus BILLION-BRYANT, MAZIN, & POUECH, 2010, BRASSEY, MAIDMENT & BARRETT, 2015

Locality: Red Canyon Ranch, near Shell, Big Horn County, Wyoming.

Horizon: Member? Morrison Formation.

Biostratigraphy: Zone?

Age: Age?, Stage?, Malm Epoch, Late Jurassic.

Material:

NHMUK R36730 (old SMA 0173-DS-RCR2003-02): Fragmentary specimen. (Nicknamed "Sarah/Sophie ")

 

HAYASHI, CARPENTER, WATABE & McWHINNEY, 2012

Locality: Bone Cabin Quarry, 8 miles north of Como, Medicine Bow Anticline (also called Flat top Anticline), Albany County, Wyoming.

Horizon: Salt Wash Member, Morrison Formation.

Biostratigraphy: Zone 2.

Age: Comobluffian age, Kimmeridgian Stage, uppermost Malm Epoch, Late Jurassic.

Material:

NSM-PV 20380: Fragmentary skeleton of an young adult.

Locality: Reed's YPM Quarry 13, T22N, R76W, Sec. 4 or 5, Como Bluff, East (East of Marshall road), Albany County, Wyoming.

Horizon: Salt Wash Member, Morrison Formation.

Biostratigraphy: Zone 2.

Age: Comobluffian age, Kimmeridgian Stage, uppermost Malm Epoch, Late Jurassic.

Material:

DMNH 1483: Fragmentary skeleton of an old adult.

 

 

TURNER & PETERSON, 1999

Locality: Lake’s quarries, 1, 5, 8, Jefferson County, Colorado.

Horizon: Brushy Basin Member, Morrison Formation.

Biostratigraphy: Zone 4.

Age: Upper Part, Comobluffian age, Kimmeridgian Stage, uppermost Malm Epoch, Late Jurassic.

Material:

Number: Not given:

_____________________________________________________________________________________
Species: stenops MARSH, 1887
Etymology:
= Diracodon stenops (MARSH, 1887) BAKKER, 1986

Holotype: USNM 4934

Locality: Marsh-Felch’s YPM Quarry 1, Garden Park, Sec. 28, T17S, R70W, Canyon City, Fremont County, Colorado.

Horizon: Salt Wash Member, Morrison Formation.

Biostratigraphy: Zone 2.

Age: Comobluffian age, Kimmeridgian Stage, uppermost Malm Epoch, Late Jurassic.

Material: Skull, lower jaws, atlas, axis, 25 presacral vertebrae, 3 sacral vertebrae, 22 caudal vertebrae, 5 cervical ribs, 12 thoracic ribs (right), 13 thoracic ribs (left), 4 chevron, left scapula, left coracoid, 2 humeri, 2 ulnae, 2 radii, 4 left metacarpals (II, III, I, V), 2 left proximal row digits III, IV, left carpus, right radial, 3 right metacarpals, 5 right phalanges, 2 femora, right tibia, right fibula, right astragalus, right calcaneum, 2 ilia, 2 ischia, 2 pubes, 17 dermal plates, 1 dermal spine and numerous dermal ossicles.

Stegosaurus hyoids.

Ventral view of the pelvis.

Scales of Stegosaurus stenops. A-B) holotype USNM 4934, A) dorsal view, B) ventral view, after Gilmore 1914, C-D) Specimen from Bone Cabin Quarry (after Glut 2003).

Stegosaurus stenops (after Marsh via Ostrom and McIntosh 1966).

Stegosaurus stenops tail spikes after Gilmore, 1914.

Tail spikes of Stegosaurus stenops. A) V-shaped tail spikes, B) horizontal tail spikes, C) caudal (tail) vertebrae in lateral view, D), Tail spikes over caudal vertebrae, and E) dorsal view of tail spikes.

Referred specimens:

GILMORE, 1914

USNM 2112: Left coracoid.

USNM 2274: Posterior portion of a skull.

USNM 4280: Right metatarsals I-III, right proximal phalanx digit I, right proximal phalanges (Digits II and III0. ungual phalanx Digit III.

USNM 4714: 25 caudal vertebrae, one with chevron, 6 dermal plates, 4 dermal spines.

 

USNM 4929: Left and right radius, left and right ulna, left humerus, left scapula, five bones of left fore foot.

USNM 6135: Dorsal vertebrae, 1-4, 6-9, 11-13, sacrals, caudal centrum, 36 caudal vertebra, 4 caudal spines, 13 chevrons, dermal plates 15-19? right and left ilia, right and left pubes, right and left ischia, 2nd dorsal rib, 6th dorsal rib, 7th left and right dorsal ribs, 15th dorsal rib and right sternal bone, 3 dermal plates, tail dermal spines.

USNM 6629: Dermal spine, 13 caudal vertebrae.

USNM 6645: Occiput and brain case, with a few other elements.

USNM 7401: Right ulna and right radius. Juvenile.

USNM 7349: Left pes metatarsals I-III, left digit II proximal phalanx.

USNM 7361: Right scapulocoracoid.

USNM 7366: Left ungual phalanx digit II and right pes.

USNM 7369: Right ungual phalanx of digit I.

USNM 7371: Left scapula.

USNM 7383: Dermal plate 3.

USNM 7389: Left fibula.

USNM 7397: Right calcaneum.

USNM 7411: Right coracoid, 1st left dorsal rib, 2nd right dorsal rib, 5th right dorsal rib.

USNM 7584: Posterior nucal plate, dermal plates 6-12.

USNM 7612: Left proximal phalanx digit I.

USNM 7615: Dermal ossicles, dermal plates 1-2.

USNM 7731: 3rd dorsal rib, 8th left dorsal rib, 9th left dorsal rib.

USNM 7736: Left ungual phalanx digit II.

USNM 7764: Right metacarpal I.

USNM 7782: Left ungual phalanx digit I.

USNM 7809: Left 6th dorsal rib.

YPM 1856: Atlas, axis, 3rd cervical vertebra, Anterior cervical vertebra, mid-cervical vertebra, and posterior cervical vertebra, anterior dorsal vertebrae, 1st and 2nd caudal vertebrae, right pubis, and tibia and astragalus.

 

Locality: Reed's YPM Quarry 13, T22N, R76W, Sec. 4 or 5, Como Bluff, East (East of Marshall road), Albany County, Wyoming.

Horizon: Salt Wash Member, Morrison Formation.

Biostratigraphy: Zone 2.

Age: Comobluffian age, Kimmeridgian Stage, uppermost Malm Epoch, Late Jurassic.

Material:

YPM 4834: Sacrum and endosacral cast.

GILMORE, 1918

USNM 8612: A composite skeleton comprised of the following;

USNM 736: Left proximal digit I.

USNM 2112: Right coracoid.

USNM 4280: Right metatarsals I-III, right proximal phalanx digit I, right proxmials digits II-III, right ungual phalanx digit III.

USNM 4714: Dermal plate no. 13.

USNM 4929: Left scapula, humeri, radii, ulnae, left ulnare, left metacarpals I, IV, V.

USNM 4935: Skull, right dentary, atlas, axis.

USNM 6531: 1-4, 6-9, 11-13 dorsal vertebrae, sacral, 5-45 caudal vertebrae, dermal plates no. 16-19, dermal spines 1-4, left femur, ilia, pubes, ischia, right second dorsal rib, left sixth dorsal rib, left and right dorsal rib seven, left dorsal rib eight, right dorsal rib fifteen and right sternal.

USNM 7348: 10 cervical vertebrae.

USNM 7349: Left metatarsals I-III, left proximal digit II.

USNM 7361: Right scapula, right coracoid.

USNM 7411: Left third cervical rib, left dorsal rib, right fifth dorsal rib.

USNM 7764: Right metacarpal I.

USNM 7366: Right ungual phalanx digit II.

USNM 7369: Right ungual phalanx digit I.

USNM 7380: Right femur, left tibia, right fibula.

USNM 7383: Dermal plate no. 3.

USNM 7397: Right calcaneum.

USNM 7389: Left fibula.

USNM 7584: Dermal plate no. 6-12.

USNM 7612: Left proximal phalanx, digit I.

USNM 7615: Dermal plates no. 1-2, 4.

USNM 7731: Left second dorsal rib, left dorsal rib eight and right dorsal rib nine.

USNM 7736: Left ungual phalanx digit II.

USNM 7782: Left ungual phalanx digit I.

USNM 7809: Left sixth dorsal rib.

Note: The rest of the skeleton is plaster.

 

ARMSTRONG, AVERETT, AVERETT, McREYNOLDS & WOLNY, 1987/BOLLAN, 1991

Locality: Bollan’s Site, Rabbit Valley, Mesa County, Western Colorado.

Horizon: Lower Part, Brushy Basin Member, Morrison Formation.

Biostratigraphy: Zone 2.

Age: Comobluffian age, Kimmeridgian Stage, uppermost Malm Epoch, Late Jurassic.

Material:

Number: Not given: 2 complete dorsal plates, 1 caudal plate, 1 cervical plate, 4 articulated tail spikes, a fifth smaller tail spike, 7 caudal vertebrae, front limb bones, hind limb bones, 1 complete manus, parts of the ilium and the ischium, several ribs, dermal ossicles, 47 vertebrae, and 12 neural spines.

Note: Bollan Stegosaur.

 

HARRIS & CARPENTER, 1996, CARPENTER, 1998

Locality: DMNH Quarry 6 (Small’s Quarry), DMNH Loc., 6111, 1992 Stegosaurus stenops Quarry, Garden Park, near Canon City, Fremont County, Colorado.

Horizon: Lower Upper Brushy Basin Member, Morrison Formation.

Biostratigraphy: Zone 2.

Age: Comobluffian age, Kimmeridgian Stage, uppermost Malm Epoch, Late Jurassic.

Material:

DMNH 2818: Skull, skeleton with skin impressions.

DMNH 2212: Broken tail spike.
Note: 2 pathologic tail spikes (McWHITNEY, ROTHSCHILDS & CARPENTER, 2001,CARPENTER, SANDERS, McWHINNEY & WOOD, 2005)

 

HOLT, 1990

Locality: Field Museum Quarry, Grand Junction, Colorado.

Horizon: Member? Morrison Formation.

Biostratigraphy: Zone?

Age: Age?, Stage?, Malm Epoch, Late Jurassic.

Material:

Numbers: Not given: 31 caudal vertebrae, 4 spines, 2 dermal plates, femur and a number of ribs.

 

LEADER & SMALL, 1999

Locality: Wolf Creek Site, Near Massadona, Moffat County, Colorado.

Horizon: Brushy Basin Member, Morrison Formation.

Biostratigraphy: Zone?

Age: Age?, Stage?, Malm Epoch, Late Jurassic.

Material:

DMNH: Disarticulated bones.

 

MADSEN, 1976

Locality: Cleveland-Lloyd Dinosaur Quarry, approximately 20 miles south of Price, SE 1/4, SE 1/4, Sec. 21, T17S, R11E, SLBM. lat 39°19’00", Long. 110°41’15"W, San Rafael Swell Area, Carbon County, Utah.

Horizon: Brushy Basin Member, Morrison Formation.

Biostratigraphy: Zone 3.

Age: Upper Part, Comobluffian age, Kimmeridgian Stage, uppermost Malm Epoch, Late Jurassic.

Material:

Number: Not given: At least 2 individuals.

 

TURNER & PETERSON, 1999

Locality: DMNH Quarry 2 (Kessler’s 1937 Quarry), Fremont County, Colorado.

Horizon: Lower Part, Brushy Basin Member, Morrison Formation.

Biostratigraphy: Zone 2.

Age: Comobluffian age, Kimmeridgian Stage, uppermost Malm Epoch, Late Jurassic.

Material:

Number: Not given:

 

= Diracodon laticeps MARSH, 1881

Locality: Reed's YPM Quarry 13, T22N, R76W, Sec. 4 or 5, Como Bluff, East (East of Marshall road), Albany County, Wyoming.

Horizon: Salt Wash Member, Morrison Formation.

Biostratigraphy: Zone 2.

Age: Comobluffian age, Kimmeridgian Stage, uppermost Malm Epoch, Late Jurassic.

Pleisotype:

USNM 4288: 23 caudal vertebrae, 4 dermal spines, one plate.

 

= Stegosaurus cf. S. stenops

Locality: Kenny’s Stegosaurus Site, Garden Park area, Canon City, Fremont County, Colorado.

Horizon: Lower Part, Brushy Basin Member, Morrison Formation.

Biostratigraphy: Zone 2.

Age: Comobluffian age, Kimmeridgian Stage, uppermost Malm Epoch, Late Jurassic.

Material:

Number: Not given:

 

= Stegosaurus stenops (?) TURNER & PETERSON, 1999

Locality: Lower Split Rock Site 1 (in lower sandstone bed), Mesa County, Western Colorado.

Horizon: Brushy Basin Member, Morrison Formation.

Biostratigraphy: Zone 3.

Age: Upper Part, Comobluffian age, Kimmeridgian Stage, uppermost Malm Epoch, Late Jurassic.

Material:

Number: Not given:

 

Locality: Bone Cabin Quarry 8 miles north of Como, Medicine Bow Anticline (also called Flat top Anticline), Albany County, Wyoming.

Horizon: Salt Wash Member, Morrison Formation.

Biostratigraphy: Zone 2.

Age: Comobluffian age, Kimmeridgian Stage, uppermost Malm Epoch, Late Jurassic.

Material:

Number: Not given:

 

Locality: Bone Cabin Quarry East (Eastward extension of old Bone Cabin Quarry), 8 miles north of Como, Medicine Bow Anticline (also called Flat top Anticline), Albany County, Wyoming.

Horizon: Salt Wash Member, Morrison Formation.

Biostratigraphy: Zone 2.

Age: Comobluffian age, Kimmeridgian Stage, uppermost Malm Epoch, Late Jurassic.

Material:

Number: Not given:

 

Locality: Meilyn Quarry, Medicine Bow Anticline (also called Flat top Anticline), Carbon County, Wyoming.

Horizon: Lower Salt Wash Member, Morrison Formation.

Biostratigraphy: Zone 1.

Age: Lower Part, Comobluffian age, Kimmeridgian Stage, uppermost Malm Epoch, Late Jurassic.

Material:

Number: Not given:

 

Locality: Weege Boys Quarry, Medicine Bow Anticline (also called Flat top Anticline), Carbon County, Wyoming.

Horizon: Lower Part, Brushy Basin Member, Morrison Formation.

Biostratigraphy: Zone 2.

Age: Comobluffian age, Kimmeridgian Stage, uppermost Malm Epoch, Late Jurassic.

Material:

Number: Not given:

_____________________________________________________________________________________

Species: ungulatus MARSH, 1879
Etymology:
= Stegosaurus angulatus SAUVAGE, 1880 (sic)

Holotype: YPM 1853

Locality: Reed’s YPM Quarry 12, Como Bluff, T. 22 N., R. 77 W., Sec. 18, Como Bluff West (west of Marshall Road), Carbon County, Wyoming.

Horizon: Upper Brushy Basin Member, Morrison Formation.

Biostratigraphy: Zone 3.

Age: Upper Part, Comobluffian age, Kimmeridgian Stage, uppermost Malm Epoch, Late Jurassic.

Material: Dorsal vertebrae, 3rd, 5th, and other caudal vertebrae, right humerus, left ulna, left radius, scapula and coracoid, left ischium, left femur, left tibia, fibula, astragalus and calcaneum, metacarpal, caudal spines, 3 neck plates, a dorsal plate, sacral plate?

Note: The mounted specimen at Yale is as follows (CARPENTER & GALTON, 2001)…
YPM 1853: Posterior part of skull roof, braincase, quadrates (replaced with cast of skull USNM 4934 in the late 1980s), posterior dorsals, caudals, ribs, right humerus, both femora, left tibia and fibula, right fibula, pes phalanges, plates and paired spikes.
Stegosaurus duplex (YPM 1858), cervicals, dorsals, dorsal ribs, sacrum, distal caudals, ilia, left pubis and ischium.
Stegosaurus sulcatus (YPM 1854) right scapula and coracoid (cast) and right ulna.
Stegosaurus sulcatus (YPM 1859) right carpus, right metacarpals and phalanges.
Stegosaurus sp (YPM), right radius, left carpus, metacarpals and phalanges.

Stegosaurus ungulatus (after Marsh via Ostrom and McIntosh 1966).

Stegosaurus ungulatus tail spikes after Gilmore, 1914.

Referred specimens:

Locality: Reed's YPM Quarry 13, T22N, R76W, Sec. 4 or 5, Como Bluff, East (East of Marshall road), Albany County, Wyoming.

Horizon: Salt Wash Member, Morrison Formation.

Biostratigraphy: Zone 2.

Age: Comobluffian age, Kimmeridgian Stage, uppermost Malm Epoch, Late Jurassic.

Material:

YPM 1857: Sacrum, ilium? endosacral cast, 3rd and 5th caudal vertebrae.

USNM 6099: 21 caudal vertebrae, 4 spines, 2 dermal plates.

 

Locality:

Horizon: Member?, Morrison Formation.

Biostratigraphy: Zone ?

Age: Age?, Stage?, Malm Epoch, Late Jurassic.

Material:

USNM 7348: Cervical vertebrae.

USNM 6646: 2 ungual phalanges, part of radius, right femur, portion of ilium, dorsal centrum, caudal centrum, 3 caudal vertebrae, tibia, calcaneum, and right astragalus, carpal bone, 2 dorsal vertebrae, distal end of right fibula, right humerus, chevron, top of caudal spine, fragment of plate, tall dermal spine, dorsal process fragment, dorsal neural process fragment.
Note: Pathologic dermal spike (McWHITNEY, ROTHSCHILDS & CARPENTER, 2001, CARPENTER, SANDERS, McWHINNEY & WOOD, 2005)

TMM 42175-2: Caudal vertebrae.

TMM 42175-3: Caudal spine.

TMM 42175-4: Anterior scutes.

TMM 42175-5: Anterior scutes.

 

Locality: ?

Horizon: Member? Morrison Formation.

Biostratigraphy: Zone?

Age: Age?, Stage?, Malm Epoch, Late Jurassic.

Material:

YPM 1854: Left ulna, tubercular spine.

 

ESCASO, ORTEGA, DANTAS, MALAFAIA, PIMENTEL, PEREDA-SUBERBIOLA, SANZ, KULLBERG, KULLBERG & BARRIGA, 2007

Locality: Casal Novo Locality, Batalha, Portugal.

Horizon:

Biostratigraphy:

Age: Kimmeridgian-Tithonian Stage, Middle-Upper Malm Epoch, Late Jurassic.

Material:

LHNB(CN) 1-01: Tooth.

LHNB(CN) 1-02: Midcaudal vertebra.

LHNB(CN) 1-03: Anteriro caudal vertebra.

LHNB(CN) 1-04: Distal part of an anterior caudal vertebra neural spine.

LHNB(CN) 1-05: Right tibia, fibula, astragalus and calcaneum.

LHNB(CN) 1-06: Anterior dorsal vertebra (3rd?).

LHNB(CN) 1-07: Cervical plate.

LHNB(CN) 1-10: Midposterior dorsal vertebra.

LHNB(CN) 1-13: Third cervical centra.

LHNB(CN) 1-16: Fourth or fifth cervical vertebra.

LHNB(CN) 1-17: Axis.

 

TURNER & PETERSON, 1999

Locality: Reeds YPM Quarry 11, Como Bluff West (west of Marshall Road), Albany County, Wyoming.

Horizon: Brushy Basin Member, Morrison Formation.

Biostratigraphy: Zone 3.

Age: Upper Part, Comobluffian age, Kimmeridgian Stage, uppermost Malm Epoch, Late Jurassic.

Material:

Number: Not given:

 

= Stegosaurus duplex MARSH, 1887
Etymology:

Holotype: YPM 1858

Locality: Reeds YPM Quarry 11, T. 22 N., R. 77 W., Sec. 12, Como Bluff West (west of Marshall Road), Albany County, Wyoming.

Horizon: Brushy Basin Member, Morrison Formation.

Biostratigraphy: Zone 3.

Age: Upper Part, Comobluffian age, Kimmeridgian Stage, uppermost Malm Epoch,

Material: Fourth (?) cervical vertebra, dorsal vertebrae, thoracic rib, sacrum and right and left ilia, caudal vertebrae, right and left pubis, right and left ischium, left femur, left tibia, fibula, astragalus, and calcaneum.

_____________________________________________________________________________________

Species: ? affinis MARSH 1881 (nomen nudum)
Etymology:

Holotype: Uncatalogued.

Locality: Reed's YPM Quarry 13, T22N, R76W, Sec. 4 or 5, Como Bluff, East (East of Marshall road), Albany County, Wyoming.

Horizon: Salt Wash Member, Morrison Formation.

Biostratigraphy: Zone 2.

Age: Comobluffian age, Kimmeridgian Stage, uppermost Malm Epoch, Late Jurassic.

Material: Pubis.

_____________________________________________________________________________________
Species: sulcatus MARSH, 1887
Etymology: Latin, sulcatus, "to make furrowed." In reference to the furrows on the caudal spines.

Holotype: USNM 4937

Locality: Reed's YPM Quarry 13, T22N, R76W, Sec. 4 or 5, Como Bluff, East (East of Marshall road), Albany County, Wyoming.

Horizon: Salt Wash Member, Morrison Formation.

Biostratigraphy: Zone 2.

Age: Comobluffian age, Kimmeridgian Stage, uppermost Malm Epoch, Late Jurassic.

Material: 4 dermal spines, 2 dermal plates (one pertaining to the neck, the other to the mid-dorsal region), right humerus, right ulna, two radii, both fore feet, portions of both scapulae, parts of femora, fibula, ischium, and caudal vertebrae.

Feet of Stegosaurus sulcatus, A). hand (manus), B) pes (hind foot), and C) claw in dorsal and lateral views.

Stegosaurus sulcatus spikes after Gilmore, 1914,

Referred specimens:

YPM 1854: Scapula-coracoid, left ulna.

YPM 1859: Ulnare, metacarpal I, III, IV, phalangeal elements, unguals.

YPM 4835: Left radius.

YPM 4836: Metatarsal I or II, II or III, III or IV.

 

BAKKER, 1990

Locality: Reed's YPM Quarry 13, T22N, R76W, Sec. 4 or 5, Como Bluff, East (East of Marshall road), Albany County, Wyoming.

Horizon: Salt Wash Member, Morrison Formation.

Biostratigraphy: Zone 2.

Age: Comobluffian age, Kimmeridgian Stage, uppermost Malm Epoch, Late Jurassic.

Material:

USNM 6646: Fragmentary skeleton.

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