Genus: Anatosaurus LULL & WRIGHT, 1942
Etymology: Latin, ant- (aanas), “duck”, and Greek, sauros, “lizard”: “duck
lizard”.
= Amatosaurus D. A. RUSSELL, 1982 (sic)
= Annatosaurus KRASSILOV, 1981 (sic)
= Anotosaurus COLBERT, 1962 (sic)
Species: longiceps (MARSH,
1890) LULL & WRIGHT,
1942
Etymology: Latin, longi, "long" and Latin, -ceps, "head." Long head.
= Edmontosaurus longiceps (MARSH, 1890)
= Trachodon longiceps MARSH, 1890
= Hadrosaurus longiceps (MARSH, 1890) NOPCSA, 1899
= Thespeius longiceps (MARSH, 1890) L.S. RUSSELL, 1930
= Anatotian longiceps (MARSH, 1890) OLSHEVSKY,
1991
Holotype: YPM 616
Locality: Niobrara County, Wyoming.
Horizon: Lance Formation.
Biostratigraphy:
Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.
Material: Right dentary without teeth.
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Species: annectens (MARSH, 1892) LULL & WRIGHT,
1942
Etymology: Latin annectens, "linking or joining."
= Claosaurus annectens MARSH, 1892
= Thespesius annectens (MARSH, 1892)
= Trachodon annectens (MARSH, 1892)
= Claosaruus annectans WILLISTON, 1898 (sic)
= Edmontosaurus annectens (MARSH, 1892)
BRETT-SURMAN
vide CHAPMAN & BRETT-SURMAN, 1990
Holotype: USNM 2414
Locality: Lightning Creek, Niobrara County, Wyoming.
(according to LULL & WRIGHT, 1942, T64W, R39N, just south of Cheyenne River
at Schneider Creek, Niobrara County, Wyoming)
Horizon: Lance Formation.
Biostratigraphy:
Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.
Material: Skull and skeleton.
After Marsh
Plesiotype: YPM 2182
Locality: T64W, R38N, north of Lance Creek, Niobrara County, Wyoming.
Horizon: Lance Formation.
Biostratigraphy:
Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.
Material: Mounted skeleton.
Referred material:
BEECHER, 1902
Locality: Niobrara (Converse) County, Wyoming.
Horizon: Lance (Laramie) Formation.
Biostratigraphy:
Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.
Material:
Number: Not given: Skull and fragmentary skeleton.
Note: Pathology on the posteroventral right scapula. An unusual smooth-edged oval-shaped perforation.
BELL,2014
Locality:
Horizon:
Biostratigraphy:
Age: Late Cretaceous.
Material:
NMSG P 001: Nearly complete specimen. Note: With sin impressions in the throat area.
BOCHERENS, FIZET, CUIF, JAEGER, MICHARD & MARIOTTI, 1988
Locality:
Horizon: Lance Formation.
Biostratigraphy:
Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.
Material:
MNH-AMN 17: Skull and skeleton.
CAMPIONE & EVANS, 2011
Locality:
Horizon:
Biostratigraphy:
Age:
Material:
SM R4050: Skull and dentary.
DMNH 1493: Skull.
UMMP 20000: Skull and dentary.
NCSM 23119: Skull and dentary.
UCMP 128374: Skull and dentary.
ROM 57100: Skull and dentary.
CARPENTER, (1998) 2000
Locality: Dry Creek, Dawson County, Montana.
Horizon: Hell Creek Formation.
Biostratigraphy:
Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.
Material:
DMNH 1943: Nearly complete specimen.
Note: Pathologic left ilium, and caudal neural spines 13-16 pathologic, possible bitten by a tyrannosaurid.
DERSTLER, 1994
Locality: Wyoming.
Horizon: Lance Formation.
Biostratigraphy:
Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.
Material:
BHI: A nearly complete skeleton nicknamed Boggy Lipps.
DERSTLER, 1995
Locality: Dragons’ Grave, Niobrara County, Wyoming.
Horizon: Lance Formation.
Biostratigraphy:
Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.
Material:
Number: Not given: Edmontosaurus bone bed ranging from third-grown juveniles to large adults.
Note: Troodontid eggshells and juvenile bones found near by.
DONNAN, 1993
Locality: Ruth Mason Dinosaur Quarry, Faith, South Dakota.
Horizon: Hell Creek Formation.
Biostratigraphy:
Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.
Material:
BHI: A juvenile skeleton, and remains of a herd.
DOUGHTY, 1985
Locality: South Dakota.
Horizon: Hell Creek Formation.
Biostratigraphy:
Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.
Material:
Ulster Museum: Skull and skeleton.
EIN, 1993Locality: Dragon’s Grave, on a remote Ranch in Wyoming.
Horizon: Lance Formation.Biostratigraphy:
Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.
Material:
Number: Not given: Several skeletons.
= cf. Edmontosaurus annectens FARKE & YIP, 2019
Locality: RAM locality V200029, McCone County, Montana.
Horizon: Hell Creek Formation.
Biostratigraphy:
Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.
Material:
RAM 9396: Nearly complete left femur.
HORNER, 1992
Locality:
Horizon:
Biostratigraphy:
Age:
Material:
MOR 003: Skull.
LIBKE, BELL, SOMERS & McKELLER, 2022
Locality: SMNH Locality 72F07-0023, in the Frenchman River valley, 215 miles southwest of Regina, Chambery Coulee, southeast of Eastend, near the North Dakota Border, Saskatchewan Province, Canada.
Horizon: Frenchman Formation.
Biostratigraphy:
Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.
Note: 28.3 m below the K-T boundary.Material:
RSM P2610.1: Fragmentary skeleton.
Note: With skin impressions.
LONGRICH, HORNER, ERICKSON & CURRIE, 2010
Locality: Montana.
Horizon: Hell Creek Formation.
Biostratigraphy:
Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.
Material:
AMNH 5041: Dentary. Note: Has toothmarks from a Tyrannosaurus rex.
LULL & WRIGHT, 1942Locality: Sec. 28, T27N, R49E, Montana.
Horizon: Lance Formation.
Biostratigraphy:
Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.
Material:
USNM 10289: Radius, ilium, cervical and caudal vertebrae.
Locality: Seven Mile Creek, 1/2 miles north of the Cheyenne River, in Weston County, just on the border of Niobrara County, Wyoming.Horizon: Lance Formation.
Biostratigraphy:
Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.
Material:
AMNH 5863: Skeleton.
Locality: 7 miles north of Sheldon's Ranch, Niobrara County, Wyoming.Horizon: Lance Formation.
Biostratigraphy:
Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.
Material:
USNM 6070: Section of caudal region covered by impression of skin.
USNM 4737: ("Burwell" specimen")
LYSON, HANKS & TREMAIN, 2003Locality: North Dakota.
Horizon:
Biostratigraphy:
Age: Late Cretaceous.
Material:
Number: Not given: Fragmentary specimen of a juvenile with skin impressions.
Note: Capped overlapping ovoid structus, a grooved scales and a 9cm by 10 cm trapezoid horn-like scale all from the caudal section.
McINTOSH, 1981
Locality: 2 or 3 miles north of Dogie Creek, Niobrara County, Wyoming.
Horizon: Lance Formation.
Biostratigraphy:
Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.
Material:
Number: Not given: Pelvis, many caudals, epidermis etc.
Locality: Lance Creek, Niobrara County, Wyoming.
Horizon: Lance Formation.
Biostratigraphy:
Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.
Material:
CM 243: 4 caudals and 2 phalanges.
CM 245: 2 caudals and 1 phalange.
CM 254: Left scapula.
CM 252: Left astragalus and distal end of tibia.
MERRELL, 1907Locality: Castle Butte, 7 miles northwest of Forsyth, Montana.
Horizon: Lance Formation.
Biostratigraphy:
Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.
Material:
USNM 3814: Skull, part of the fore limbs, and impressions of skin.
Locality: Wyoming.
Horizon:
Biostratigraphy:
Age: Late Cretaceous.
Material:
USNM 5370: Teeth.
MORRIS, 1971
Locality: Piney Buttes area, NE 1/4, SE. 36, T21N, R35E, south east of Fort Peck Reservoir, McCone County, Central Montana.
Horizon: Hell Creek Formation.
Biostratigraphy:
Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.
Material:
LACM 23502: Skull and skeleton.
LACM ?: Fragmentary skull and skeleton of a juvenile.
OSBORN, 1912
Locality: Niobrara (Converse) County, Wyoming.
Horizon: Lance Formation.
Biostratigraphy:
Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.
Material:
AMNH FARB 5060: Skull, skeleton and skin impression (some in the nasal area, PRIETO-MARQUEZ, & WAGNER, 2014).
Note: Mummy.
Locality:Horizon:
Biostratigraphy:
Age:
Material:
AMNH 5894: Caudal vertebrae and skin.
Note: = Trachodon mirabilis OSBORN, 1912
Locality:Horizon:
Biostratigraphy:
Age:
Material:
AMNH 5058: Skeleton with skin impressions (= Trachodon sp, OSBORN, 1912).
OSTROM, 1961Locality:
Horizon:
Biostratigraphy:
Age:
Material:
AMNH 5046:
PATCHUS & DERSTLER, 1998
Locality: N43°20.345’, W104°29.029’, Niobrara County, Wyoming.
Horizon: Lance Formation.
Biostratigraphy:
Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.
Material:
University of New Orleans: Tail, pelvis and posterior dorsal are well articulated with skin impressions.
SIVIERO, REGA, HAYES, COOPER, BRAND & CHADWICK, 2020
Locality: North and South Quarries, Niobrara County, Wyoming.
Horizon: Lance Formation.
Biostratigraphy:
Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.
Material:
Bonebeds with 3013 specimens, 96 with macroscopic bone abnormalities adn 55 with pathologies atributed to physical trauma.
Pre-Mortem Trauma:
HRS07948: Neural spine from a middle-distal caudal vertebra.
HRS02082: Neural spine from a middle caudal vertebra.
HRS00428: Middle-distal caudal vertebra.
HRS01106: Neural spine from a middle-distal caudal vertebra.
HRS00428: Distal caudal vertebra.
HRS00323, HRS05801, HRS00444, HRS07245: Distal caudal vertebrae.
HRS01295: Distal fragment of neural spine.
HRS000404: Neural spine from a distal caudal vertebra.
HRS01248: Neural spine fragment from a middle-distal caudal neural spine.
HRS14435: Middle caudal vertebra.
HRS00836: Middle caudal vertebra.
HRS00853: Chevron fragment.
HRS17086: Chevron fragment with the bifurcated end mostly complete.
HRS07257: Phalanx of the manus with apparent bone remodeling.
STEIN, 2021
Locality: Tooth Draw Deposit, Butte County, South Dakota.
Horizon: Hell Creek Formation.
Biostratigraphy:
Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.
Material:
TD-14-257: Fragmentary neonate right dentary.
TD-16-259: Partial fibual.
Note: With tooth marks and oblique irregular, pre-depositonal frature.
TD-13-037: Neonate humerus.
TDW-19-064: Adult humerus.
Number: Not given (528): Teeth, cranial elements, axial and appendicular elements.
TRIEBOLD & RUSSELL, 1995
Locality: “Sandy Quarry”, near Buffalo, South Dakota.
Horizon: Hell Creek Formation.
Biostratigraphy:
Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.
Material:
Number: Not given: 2 skeletons and isolated elements.
VERSLUYS, 1923
Locality: South Schneider Creek, Niobrara County, Wyoming.
Horizon: Lance Formation.
Biostratigraphy:
Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.
Material:
Senckenberg-Museum Nr. 4036: Skull, mandible and fragmentary skeleton.
Note: Skeleton with skin impressions. (= Trachodon sp, OSBORN, 1912).
Anatosaurus annectens (after VERSLUYS, 1923), Senckenberg-Museum Nr. 4036.
WIELAND, 1925
Locality:?
Horizon:?
Biostratigraphy:
Age:
Material:
Number: Not given: Mummy with stomach contents of twigs of conifers and dicots, needles of Cunninghamites elegans and numerous seeds of fruits, all in macerated form.
WOSIK, GOODWIN & EVANS, 2017, 2019
Loclaity: Sandstone Basin Locality (UCMP locality V80092), Garfield County, northeastern Montana.
Horizon: Hell Creek Formation.
Biostratigraphy:
Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.
Material:
UCMP 235860: A nearly complete right dentary of a nestling.
UCMP 283755: An incomplete and edentulous posterior portion of a right dentry dental battery of a nestling.
UCMP 235857: An incomplete right surangular of a nestling.
UCMP 235859: An incomlpee right qudrate of a nestling.
UCMP 128181: fragmentary skeleton of a nestling.
ZHENG, FARKE & KIM, 2011
Locality: RAM V97048, McCone County, Montana.
Note: Detailed locality data are on file at the RAM and available to qualified researhcers upon request.
Horizon: Hell Creek Formation.
Biostratigraphy:
Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.
Material:
RAM 7150: A partial, mostly articulated skelton presearving the right hindlimb complete below the mid-shaft of the femur, right ischium, 77 caudal vertebrae, 23 chevrons and skin impression.
Private collection.
= Claosaurus affinis WIELAND, 1903 (nomen dubium)
Etymology: Latein, ad + finis, “boundary."
= Trachodon affinis (WIELAND, 1903) (nomen dubium)Holotype: YPM 3219
Locality: West side of Cheyenne River, nearly opposite Corral Draw, near Red Shirt, Custer County, South Dakota.
Horizon: Pierre Formation.
Biostratigraphy:
Age: Campanian Stage, Senonian Subepoch, Gulf Epoch, Late Cretaceous.
Material: Proximal phalanx digits, II, IV and third phalanx digit IV.
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Species: saskatchewanensis (STERNBERG,
1926) LULL & WRIGHT, 1942
Etymology: In reference to Saskatchewan
Province, Canada.
= Thespesius saskatchewanensis STERNBERG,
1926
= Trachodon saskatchewanensis (STERNBREG,
1926) KUHN, 1936
= Edmontosaurus saskatchewanensis (STERNBERG,
1926)
BRETT-SURMAN
vide CHAPMAN & BRETT-SURMAN,
1990
Holotype: NMC 8509
Locality: Rocky Creek, sec. 22, T1N, R5, west of 3rd meridian, Saskatchewan Province, Canada.
Horizon: Lance Formation.
Biostratigraphy:
Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.
Material: An almost complete skull, 16 dorsal vertebrae, 37 caudal centra, fragmentary chevrons, left scapula and coracoid, left femur, and part of the right, right tibia with astragalus and calcaneum, both ischia and some fragmentary ribs.
Referred material:
HORNER, 1992
Locality:
Horizon:
Biostratigraphy:
Age:
Material:
UCM 20829:
