Genus: Anatosaurus LULL & WRIGHT, 1942
Etymology: Latin, ant- (aanas), “duck”, and Greek, sauros, “lizard”: “duck
lizard”.
= Amatosaurus D. A. RUSSELL, 1982 (sic)
= Annatosaurus KRASSILOV, 1981 (sic)
= Anotosaurus COLBERT, 1962 (sic)
Species: longiceps (MARSH,
1890) LULL & WRIGHT,
1942
Etymology: Latin, longi, "long" and Latin, -ceps, "head." Long head.
= Edmontosaurus longiceps (MARSH, 1890)
= Trachodon longiceps MARSH, 1890
= Hadrosaurus longiceps (MARSH, 1890) NOPCSA, 1899
= Thespeius longiceps (MARSH, 1890) L.S. RUSSELL, 1930
= Anatotian longiceps (MARSH, 1890) OLSHEVSKY,
1991
Holotype: YPM 616
Locality: Niobrara County, Wyoming.
Horizon: Lance Formation.
Biostratigraphy:
Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.
Material: Right dentary without teeth.
_____________________________________________________________________________________
Species: annectens (MARSH, 1892) LULL & WRIGHT,
1942
Etymology: Latin annectens, "linking or joining."
= Claosaurus annectens MARSH, 1892
= Thespesius annectens (MARSH, 1892) GILMORE, 1924
= Trachodon annectens (MARSH, 1892) HATCHER, 1902
= Claosaruus annectans WILLISTON, 1898 (sic)
= Edmontosaurus annectens (MARSH, 1892) pro BRETT-SURMAN, 1975 (M.S. dissertation, unpublished)
= Edmontosaurus annectens (MARSH, 1892) McINTOSH, 1982
= Edmontosaurus annectens (MARSH, 1892)
BRETT-SURMAN
vide CHAPMAN & BRETT-SURMAN, 1990
Holotype: USNM 2414
Locality: Lightning Creek, Niobrara County, Wyoming.
(according to LULL & WRIGHT, 1942, T64W, R39N, just south of Cheyenne River
at Schneider Creek, Niobrara County, Wyoming)
Horizon: Lance Formation.
Biostratigraphy:
Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.
Material: Skull and skeleton.
From Marsh, 1892.
After Marsh
Plesiotype: YPM 2182
Locality: T64W, R38N, north of Lance Creek, Niobrara County, Wyoming.
Horizon: Lance Formation.
Biostratigraphy:
Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.
Material: Mounted skeleton.
Referred material:
BEECHER, 1902
Locality: Niobrara (Converse) County, Wyoming.
Horizon: Lance (Laramie) Formation.
Biostratigraphy:
Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.
Material:
Number: Not given: Skull and fragmentary skeleton.
Note: Pathology on the posteroventral right scapula. An unusual smooth-edged oval-shaped perforation.
BELL,2014
Locality:
Horizon:
Biostratigraphy:
Age: Late Cretaceous.
Material:
NMSG P 001: Nearly complete specimen. Note: With sin impressions in the throat area.
BOCHERENS, FIZET, CUIF, JAEGER, MICHARD & MARIOTTI, 1988
Locality:
Horizon: Lance Formation.
Biostratigraphy:
Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.
Material:
MNH-AMN 17: Skull and skeleton.
CAMPIONE & EVANS, 2011
Locality:
Horizon:
Biostratigraphy:
Age:
Material:
SM R4050: Skull and dentary.
DMNH 1493: Skull.
UMMP 20000: Skull and dentary.
NCSM 23119: Skull and dentary.
UCMP 128374: Skull and dentary.
ROM 57100: Skull and dentary.
CARPENTER, (1998) 2000
Locality: Dry Creek, Dawson County, Montana.
Horizon: Hell Creek Formation.
Biostratigraphy:
Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.
Material:
DMNH 1943: Nearly complete specimen.
Note: Pathologic left ilium, and caudal neural spines 13-16 pathologic, possible bitten by a tyrannosaurid.
DERSTLER, 1994
Locality: Wyoming.
Horizon: Lance Formation.
Biostratigraphy:
Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.
Material:
BHI: A nearly complete skeleton nicknamed Boggy Lipps.
DERSTLER, 1995
Locality: Dragons’ Grave, Niobrara County, Wyoming.
Horizon: Lance Formation.
Biostratigraphy:
Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.
Material:
Number: Not given: Edmontosaurus bone bed ranging from third-grown juveniles to large adults.
Note: Troodontid eggshells and juvenile bones found near by.
DONNAN, 1993
Locality: Ruth Mason Dinosaur Quarry, Faith, South Dakota.
Horizon: Hell Creek Formation.
Biostratigraphy:
Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.
Material:
BHI: A juvenile skeleton, and remains of a herd.
ANNE, WHITNEY, BROCKLEHURST, DONNELLY & ROTHSCHILD, 2022
TCM no number: Distal caudal with 1 distal indent.
TCM 2004.111.19: Mid-distal caudal with 1 proximal indet.
TCM 2004.111.50: Mid-distal caudal with 1 distal "mound" lesion.
TCM 2005.143.27: Mid-distal caudal with 3-4 proximal and 1 distal "button" lesion.
TCM 2005.143.29: Mid-distal caudal with 2 proximal "mound" lesion.
TCM 2006.63.45: Mid-distal caudal with 1 proximal and distal indent.
TCM 2007.207.82: Mid-distal caudal with 1 proximal "mound" lesion.
TCM 2007.207.92: Mid-distal caudal with porximal; distal indent.
TCM 2012.11.23: Distal caudal with 1 proximal and distal indet.
TCM 2012.11.26: Mid-distal caudal with 2 proximal "button" lesion.
TCM 2012.11.55: Mid-distal caudal with 1 proximal "mound" lesion.
TCM 2012.11.80: Mid caudal with 1 proximal "mound" lesion and 1 distal "button" lesion.
TCM 2013.22.55: Mid-distal caudal with 1 proximal "button" lesion.
TCM 2013.22.67: Mid-distal caudal with 1 proximal "button" lesion.
TCM 2014.111.52: Mid-distal caudal with 1 proximal "mound" lesion and 2 distal "button" lesion.
TCM 2014.111.101: Mid-distal caudal with 1 proximal "mound" lesion and 3 distal "button" lesion.
TCM 2014.111.119: Mid-distal caudal with 1 distal and 1 distal "button" lesion and distal indent.
TCM 2014.111.135: Mid-distal caudal with 1 proximal and 1 distal "button" lesion.
TCM 2015.16.125: Mid-distal caudal with 1 proximal indet.
TCM 2016.24.28: Mid-distal caudal with 1 distal indet.
TCM 2016.24.82: Mid-distal caudal with 1 proximal "mound" lesion.
TCM 2016.24.101: Mid-distal caudal with 1 distal "button" lesion and 1 proximal indet.
TCM 2017.121.24: Distal caudal with 1 proximal and distal indet.
DOUGHTY, 1985
Locality: South Dakota.
Horizon: Hell Creek Formation.
Biostratigraphy:
Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.
Material:
Ulster Museum: Skull and skeleton.
EIN, 1993Locality: Dragon’s Grave, on a remote Ranch in Wyoming.
Horizon: Lance Formation.Biostratigraphy:
Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.
Material:
Number: Not given: Several skeletons.
= cf. Edmontosaurus annectens FARKE & YIP, 2019
Locality: RAM locality V200029, McCone County, Montana.
Horizon: Hell Creek Formation.
Biostratigraphy:
Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.
Material:
RAM 9396: Nearly complete left femur.
HORNER, 1992
Locality:
Horizon:
Biostratigraphy:
Age:
Material:
MOR 003: Skull.
LIBKE, BELL, SOMERS & McKELLER, 2022
Locality: SMNH Locality 72F07-0023, in the Frenchman River valley, 215 miles southwest of Regina, Chambery Coulee, southeast of Eastend, near the North Dakota Border, Saskatchewan Province, Canada.
Horizon: Frenchman Formation.
Biostratigraphy:
Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.
Note: 28.3 m below the K-T boundary.Material:
RSM P2610.1: Fragmentary skeleton.
Note: With skin impressions.
LONGRICH, HORNER, ERICKSON & CURRIE, 2010
Locality: Montana.
Horizon: Hell Creek Formation.
Biostratigraphy:
Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.
Material:
AMNH 5041: Dentary. Note: Has toothmarks from a Tyrannosaurus rex.
LULL & WRIGHT, 1942Locality: Sec. 28, T27N, R49E, Montana.
Horizon: Lance Formation.
Biostratigraphy:
Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.
Material:
USNM 10289: Radius, ilium, cervical and caudal vertebrae.
Locality: Seven Mile Creek, 1/2 miles north of the Cheyenne River, in Weston County, just on the border of Niobrara County, Wyoming.Horizon: Lance Formation.
Biostratigraphy:
Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.
Material:
AMNH 5863: Skeleton.
Locality: 7 miles north of Sheldon's Ranch, Niobrara County, Wyoming.Horizon: Lance Formation.
Biostratigraphy:
Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.
Material:
USNM 6070: Section of caudal region covered by impression of skin.
USNM 4737: ("Burwell" specimen")
LYSON, HANKS & TREMAIN, 2003Locality: North Dakota.
Horizon:
Biostratigraphy:
Age: Late Cretaceous.
Material:
Number: Not given: Fragmentary specimen of a juvenile with skin impressions.
Note: Capped overlapping ovoid structus, a grooved scales and a 9cm by 10 cm trapezoid horn-like scale all from the caudal section.
McINTOSH, 1981
Locality: 2 or 3 miles north of Dogie Creek, Niobrara County, Wyoming.
Horizon: Lance Formation.
Biostratigraphy:
Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.
Material:
Number: Not given: Pelvis, many caudals, epidermis etc.
Locality: Lance Creek, Niobrara County, Wyoming.
Horizon: Lance Formation.
Biostratigraphy:
Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.
Material:
CM 243: 4 caudals and 2 phalanges.
CM 245: 2 caudals and 1 phalange.
CM 254: Left scapula.
CM 252: Left astragalus and distal end of tibia.
MERRELL, 1907Locality: Castle Butte, 7 miles northwest of Forsyth, Montana.
Horizon: Lance Formation.
Biostratigraphy:
Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.
Material:
USNM 3814: Skull, part of the fore limbs, and impressions of skin.
Locality: Wyoming.
Horizon:
Biostratigraphy:
Age: Late Cretaceous.
Material:
USNM 5370: Teeth.
MORRIS, 1971
Locality: Piney Buttes area, NE 1/4, SE. 36, T21N, R35E, south east of Fort Peck Reservoir, McCone County, Central Montana.
Horizon: Hell Creek Formation.
Biostratigraphy:
Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.
Material:
LACM 23502: Skull and skeleton.
LACM ?: Fragmentary skull and skeleton of a juvenile.
OSBORN, 1912
Locality: Niobrara (Converse) County, Wyoming.
Horizon: Lance Formation.
Biostratigraphy:
Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.
Material:
AMNH FARB 5060: Skull, skeleton and skin impression (some in the nasal area, PRIETO-MARQUEZ, & WAGNER, 2014).
Note: Mummy.
Locality:Horizon:
Biostratigraphy:
Age:
Material:
AMNH 5894: Caudal vertebrae and skin.
Note: = Trachodon mirabilis OSBORN, 1912
Locality:Horizon:
Biostratigraphy:
Age:
Material:
AMNH 5058: Skeleton with skin impressions (= Trachodon sp, OSBORN, 1912).
OSTROM, 1961Locality:
Horizon:
Biostratigraphy:
Age:
Material:
AMNH 5046:
PATCHUS & DERSTLER, 1998
Locality: N43°20.345’, W104°29.029’, Niobrara County, Wyoming.
Horizon: Lance Formation.
Biostratigraphy:
Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.
Material:
University of New Orleans: Tail, pelvis and posterior dorsal are well articulated with skin impressions.
SIVIERO, REGA, HAYES, COOPER, BRAND & CHADWICK, 2020
Locality: North and South Quarries, Niobrara County, Wyoming.
Horizon: Lance Formation.
Biostratigraphy:
Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.
Material:
Bonebeds with 3013 specimens, 96 with macroscopic bone abnormalities adn 55 with pathologies atributed to physical trauma.
Pre-Mortem Trauma:
HRS07948: Neural spine from a middle-distal caudal vertebra.
HRS02082: Neural spine from a middle caudal vertebra.
HRS00428: Middle-distal caudal vertebra.
HRS01106: Neural spine from a middle-distal caudal vertebra.
HRS00428: Distal caudal vertebra.
HRS00323, HRS05801, HRS00444, HRS07245: Distal caudal vertebrae.
HRS01295: Distal fragment of neural spine.
HRS000404: Neural spine from a distal caudal vertebra.
HRS01248: Neural spine fragment from a middle-distal caudal neural spine.
HRS14435: Middle caudal vertebra.
HRS00836: Middle caudal vertebra.
HRS00853: Chevron fragment.
HRS17086: Chevron fragment with the bifurcated end mostly complete.
HRS07257: Phalanx of the manus with apparent bone remodeling.
STEIN, 2021
Locality: Tooth Draw Deposit, Butte County, South Dakota.
Horizon: Hell Creek Formation.
Biostratigraphy:
Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.
Material:
TD-14-257: Fragmentary neonate right dentary.
TD-16-259: Partial fibual.
Note: With tooth marks and oblique irregular, pre-depositonal frature.
TD-13-037: Neonate humerus.
TDW-19-064: Adult humerus.
Number: Not given (528): Teeth, cranial elements, axial and appendicular elements.
TRIEBOLD & RUSSELL, 1995
Locality: “Sandy Quarry”, near Buffalo, South Dakota.
Horizon: Hell Creek Formation.
Biostratigraphy:
Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.
Material:
Number: Not given: 2 skeletons and isolated elements.
VERSLUYS, 1923
Locality: South Schneider Creek, Niobrara County, Wyoming.
Horizon: Lance Formation.
Biostratigraphy:
Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.
Material:
Anatosurus annectens Senckenberg-Museum Nr. 4036: Skull, mandible and fragmentary skeleton.
Note: Skeleton with skin impressions. (= Trachodon sp, OSBORN, 1912).
Anatosaurus annectens (after VERSLUYS, 1923), Senckenberg-Museum Nr. 4036.
WIELAND, 1925
Locality:?
Horizon:?
Biostratigraphy:
Age:
Material:
Number: Not given: Mummy with stomach contents of twigs of conifers and dicots, needles of Cunninghamites elegans and numerous seeds of fruits, all in macerated form.
WOSIK, GOODWIN & EVANS, 2017, 2019
Loclaity: Sandstone Basin Locality (UCMP locality V80092), Garfield County, northeastern Montana.
Horizon: Hell Creek Formation.
Biostratigraphy:
Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.
Material:
UCMP 235860: A nearly complete right dentary of a nestling.
UCMP 283755: An incomplete and edentulous posterior portion of a right dentry dental battery of a nestling.
UCMP 235857: An incomplete right surangular of a nestling.
UCMP 235859: An incomlpee right qudrate of a nestling.
UCMP 128181: fragmentary skeleton of a nestling.
ZHENG, FARKE & KIM, 2011
Locality: RAM V97048, McCone County, Montana.
Note: Detailed locality data are on file at the RAM and available to qualified researhcers upon request.
Horizon: Hell Creek Formation.
Biostratigraphy:
Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.
Material:
RAM 7150: A partial, mostly articulated skelton presearving the right hindlimb complete below the mid-shaft of the femur, right ischium, 77 caudal vertebrae, 23 chevrons and skin impression.
Private collection.
