Genus: Dryosaurus MARSH, 1894
Etymology: Greek, dryos, “tree”, and Greek, sauros, “lizard”: “Tree lizard”.
= Dryopsaurus WEISHAMPEL & HEINRICH, 1992 (sic)
= Drysaurus GALTON, 1977 (sic)
Species: altus (MARSH, 1878) MARSH, 1894
Etymology: Latin, altus, "high."
= Laosaurus altus MARSH, 1878
Holotype: YPM 1876
Locality: Reed’s YPM Quarry 5, T22N, R77W, Sec. 17, Como Bluff, Albany County, Wyoming.
Horizon: Lower Upper Brushy Basin Member, Morrison Formation.
Biostratigraphy: Zone 2.
Age: Comobluffian age, Kimmeridgian Stage, uppermost Malm Epoch, Late Jurassic.
Material: Fragmentary skull with partial braincase and endocast.
Referred specimens
GALTON, & JENSEN, 1973
Locality: Sheetz’ Quarry 1, 5.5 miles southwest of Uravan, Montrose County, Colorado.
Horizon: Upper or Brushy Basin Member, Morrison Formation.
Biostratigraphy: Zone 3.
Age: Comobluffian age, Kimmeridgian Stage, uppermost Malm Epoch, Late Jurassic.
Material:
BYU ESM-171R : Vertebral fragments, fragment of left dentary with teeth, distal end of left humerus, proximal end of radius, distal end of left radius, proximal end of left femur, distal end of right femur, proximal end of left femur, distal end of left femur, proximal end of right tibia, distal end of left tibia, distal end of right tibia, distal end of left fibula, proximal surface of left astragalus.
Locality: YPM Quarry 13 1/2 of W. H. Reed at Como Bluff, Albany County, Wyoming.
Horizon: Member? Morrison Formation.
Biostratigraphy: Zone?
Age: Age?, Stage?, Malm Epoch, Late Jurassic.
Material:
YPM 1884: Fragmentary sacrum and caudal with damaged hind limbs.
Locality: Bone Cabin Quarry, 8 miles north of Como, Medicine Bow Anticline (also called Flat top Anticline), Albany County, Wyoming.Horizon: Salt Wash Member, Morrison Formation.
Biostratigraphy: Zone 2.
Age: Comobluffian age, Kimmeridgian Stage, uppermost Malm Epoch, Late Jurassic.
Material:
AMNH 834: Several fragmentary vertebrae, pectoral girdle, humerus, pelvic girdle and hind limbs.
Locality: Bone Cabin Quarry East (Eastward extension of old Bone Cabin Quarry), 8 miles north of Como, Medicine Bow Anticline (also called Flat top Anticline), Albany County, Wyoming.
Horizon: Salt Wash Member, Morrison Formation.
Biostratigraphy: Zone 2.
Age: Comobluffian age, Kimmeridgian Stage, uppermost Malm Epoch, Late Jurassic.
Material:
Number: Not given:
McINTOSH, 1981Locality: Elk Mountains, near Brown's Ranch, Johnson County, Wyoming.
Horizon: Member? Morrison Formation.
Biostratigraphy: Zone?
Age: Age?, Stage?, Malm Epoch, Late Jurassic.
Material:
CM 1949: 3 dorsals, about 28 caudals, right ilium, right femur, right tibia, right fibula, metatarsals.
Locality: Lily Park, Moffat County, Colorado.
Horizon: Member? Morrison Formation.
Biostratigraphy: Zone?
Age: Age?, Stage?, Malm Epoch, Late Jurassic.
Material:
CM 21786: About 10 dorsals, sacrum, 30 caudals, humeri, both hind limbs with the shafts of many of the elements incomplete, both pedes, other bones.
SHEPHERD, GALTON & JENSEN, 1978Locality: Bone Cabin Quarry East (Eastward extension of old Bone Cabin Quarry), 8 miles north of Como, Medicine Bow Anticline (also called Flat top Anticline), Albany County, Wyoming.
Horizon: Salt Wash Member, Morrison Formation.
Biostratigraphy: Zone 2.
Age: Comobluffian age, Kimmeridgian Stage, uppermost Malm Epoch, Late Jurassic.
Material:
Number: Not given:
Locality: Carnegie Quarry, Dinosaur National Monument, north of the little town of Jensen, Uintah County, Utah.
Horizon: Brushy Basin Member, Morrison Formation.
Biostratigraphy: Zone 3.
Age: Comobluffian age, Kimmeridgian Stage, uppermost Malm Epoch, Late Jurassic.
Material:
DNM 1016: Left ilium.
TURNER & PETERSON, 1999Locality: DMNH Quarry 6 (Small’s Quarry), DMNH Loc., 6111, 1992 Stegosaurus stenops Quarry, Garden Park, near Canon City, Colorado.
Horizon: Lower Upper Brushy Basin Member, Morrison Formation.
Biostratigraphy: Zone 2.
Age: Comobluffian age, Kimmeridgian Stage, uppermost Malm Epoch, Late Jurassic.
Material:
Number: Not given:
TURNER & PETERSON, 1999Locality: Upper Split Rock Site 2 (in upper sandstone bed), Mesa County, Western Colorado.
Horizon: Upper Brushy Basin Member, Morrison Formation.
Biostratigraphy: Zone 4.
Age: Upper Part, Comobluffian age, Tithonian Stage, uppermost Malm Epoch, Late Jurassic.
Material:
Number: Not given:
Locality: Reed’s YPM Quarry 5, T22N, R77W, Sec. 17, Como Bluff West (west of Marshall Road), Albany County, Wyoming.Horizon: Brushy Basin Member, Morrison Formation.
Biostratigraphy: Zone 3.
Age: Comobluffian age, Kimmeridgian Stage, uppermost Malm Epoch, Late Jurassic.
Material:
Number: Not given:
_____________________________________________________________________________________
Species: elderae CARPENTER & GALTON, 2018
Etymology: In honor of Ann Schaffer Elder (1958-2009), in recognition of her considerable assistance to Carpenter in his studies at the Carnegie Quarry in Dinosaur Natinoal Monument, Utah (Carpenter, 2013).
Holotype: CM 3392
Locality: Carnegie Quarry, Dinosaur National Monument, north of the little town of Jensen, Uintah County, Utah.
Horizon: Brushy Basin Member, Morrison Formation.
Biostratigraphy: Zone 3.
Age: Comobluffian age, Kimmeridgian Stage, uppermost Malm Epoch, Late Jurassic.
Material: Skull lacking tip of premaxilla, tip of quadrajugal, and tip of surangular, first five cervical vertebrae, seven anterior dorsal vertebrae with the proximal portions of the ribs of both sides in position, together with the right scapula, coracoid, and humerus, six posterior dorsal vertebrae articulated with the sacrum, and portions of the pelvic arch, proximal third of the right femur, proximal end of right tibia, one complete metatarsal and distal portions of two others, articulated left hind foot, with some limb and other identified bones.
Dryosaurus elderae (after CARPENTER, 1994) CM 3392
Referred material:
CM 11,337: Fragmentary skeleton of a juvenile. (Note: originally referred to Laosaurus gracilis MARSH, 1878, by GILMORE, 1925)
= Dryosaurus cf. D. altus CARPENTER, & LAMANNA, 2015
CM 87688: Braincase and 5 semi-articulated cervical vertebrae. Note: Orignally part of the type material of Uteodon aphanoectes.
CARPENTER & GALTON, 2018
DINO 4619: Left ilium.
DINO 1031: Right scapula.
GILMORE, 1925
Locality: Far east side of quarry face, Carnegie Quarry, Dinosaur National Monument, north of the little town of Jensen, Uintah County, Utah.
Horizon: Brushy Basin Member, Morrison Formation.
Biostratigraphy: Zone 3.
Age: Comobluffian age, Kimmeridgian Stage, uppermost Malm Epoch, Late Jurassic.
Material:
CM 11340: Skull lacking tip of snout and premaxilla, 9 (?) cervicals in articulation, 13 (?) partial and complete dorsals plus a gap for 3 more in the pectoral region, ribs, both scapulae, coracoids, sternal plates, and humeri, distal ends of both femora, shaft of an ischium, and proximal end of the right tibia and scapula of a baby.
Dryosaurus altus (after Carpenter, 1994), CM 11340 (juvenile)
