Genus: Nanosaurus MARSH, 1877 (nomen dubium)
Etymology: Greek, nanos, “pigmy” and Greek, sauros, “lizard”: “Pigmy lizard”.
= Drinker BAKKER, GALTON, SIEGWARTH & FILLA, 1990
Etymology: In honor of Edward Drinker Cope.
= Nannosaurus NOPCSA, 1928 (sic)
= Othnielia GALTON, 1977
Etymology: In honor of Othniela Charles Marsh.
= Othnielosaurus GALTON, 2006
Etymology: In honor of Othniela Charles Marsh, and Greek, sauros, "lizard."
= Othneilia GALTON & POWELL, 1980 (sic)

Species: agilis MARSH, 1877
Etymology: Latin, agilis, "agile, nimble."

Holotype: YPM 1913

Locality: W 1/2, NE 1/4, sec. 28, T17S, R70W, Garden Park near Canon City, Fremont County, Colorado.

Horizon: Member? Morrison Formation.

Biostratigraphy: Zone?

Age: Age? Stage? Malm Epoch, Late Jurassic.

Material: Fragmentary dentary with impressions of teeth.

Paralectotypes.

YPM 1913b: Right ilium.

YPM 1913c: Femora.

YPM 1913d: Tibiae.

YPM 1913e: ?fibula.

YPM 1913f: Fragmentary long bone.

YPM 1913g: Fragmentary left tibia.

YPM 1913i: Fragmentary metatarsus.

YPM 1913m: ?Process of cranial bone, ?part of jugal, postorbital or squamosal.

YPM 1913n: ?Process of cranial bone, ?part of jugal, postorbital or squamosal.

YPM 1913p: Fragmentary left ilium.

YPM 1913q: Distal end of a small left tibia.

YPM 1913r: Anterior cervical rib.

YPM 1913rl: Anterior dorsal rib.

YPM 1913r9: Posterior cervical rib.

YPM 1913u: ?acetabular region of ilium.

YPM 1913w: ?Ischium.

A) Type jaw; B) 1913m; C) 1913n.

A) 1913b; B) 1913p; C) 1913u; D) 1913W.

A) 1913d; B) 1913f; C) 1913g; D) 1913e.

A) 1913i; B) 1913a; C) 1913r; D) 1913r9; E) 1913rl.

Referred material:

FOSTER, PAGNAC & HUNT-FOSTER, 2020

Locality: V9138, Little Houston Quarry, approximately 20 km west of Sundance in southern Crook County, just north of Little Houston Creek, Black Hills, northeast Wyoming.

Horizon: Member?, Morrison Formation.

Biostratigraphy: Zone?

Age: Age?, Stage?, Malm Epoch, Late Jurassic.

Material:

MWC 5822: Right dentary with 13 teeth.

JRF 95126: Anterior tooth.

JRF 95148: Lateral tooth.

JRF 9822: Iny femur mid-shaft.

JRF 95220: Anterior tooth.

SDSM 26913: Left femur.

SDSM 30490: Left femur of an adult.

SDSM 30494: Left femur.

SDSM 30496: Caudal vertebra.

SDSM 30502: Right humerus.

SDSM 30503: Right pubis.

GALTON, 1983

Locality: Reeds YPM Quarry 9, (Mammal Quarry), T22N, R77W, Sec. 12, Como Bluff West (west of Marshall Road), Albany County, Wyoming.

Horizon: Brushy Basin Member, Morrison Formation.

Biostratigraphy: Zone 3.

Age: Upper Part, Comobluffian age, Kimmeridgian Stage, uppermost Malm Epoch, Late Jurassic.

Material:

YPM 9523: Left dentary tooth.

YPM 9524: Left maxillary tooth.

= Nanosaurus rex MARSH, 1877
Etymology: Latin, rex, "the king."
= Laosaurus rex (MARSH, 1877) MARSH, 1896
= Othnielia rex (MARSH, 1877) GALTON, 1977

Holotype: YPM 1915

Locality: Como Bluff, Albany County, Wyoming.

Horizon: Member? Morrison Formation.

Biostratigraphy: Zone?

Age: Age?, Stage?, Malm Epoch, Late Jurassic.

Material: Left femur.

Reconstruction of femur; medial, laterial, posterior and anterior view.

Referred material:

GALTON, 1983

Locality: Reeds YPM Quarry 9, (Mammal Quarry), T22N, R77W, Sec. 12, Como Bluff West (west of Marshall Road), Albany County, Wyoming.

Horizon: Brushy Basin Member, Morrison Formation.

Biostratigraphy: Zone 3.

Age: Upper Part, Comobluffian age, Kimmeridgian Stage, uppermost Malm Epoch, Late Jurassic.

Material:

YPM 7451: Left maxillary tooth.

YPM 7452: Right dentary tooth.

Several very small dentary and maxillary teeth.

AMNH 11527, 11528: Teeth.
Note: These were refereed to Fabrosauridae in Prothro, 1981.

GALTON & JENSEN, 1973

Locality: Approximately 5 miles east of Willow Springs, Emery County, Utah.

Horizon: Upper or Brushy Basin Member, Morrison Formation.

Biostratigraphy: Zone 3.

Age: Comobluffian age, Kimmeridgian Stage, uppermost Malm Epoch, Late Jurassic.

Material:

BYU ESM-163R: Skeleton lacking skull, first cervicals, lower upper legs and caudals.

GLUT, 2006

Locality: Howe Ranch, Howe Quarry located at the very foot of the big Horn Mountains, near the original Cloverly Post Office, 25 miles northeast of Greybull, and 25 miles south of the Montana line, Big Horn County, Wyoming.

Horizon: Salt Wash Member, Morrison Formation.

Biostratigraphy: Zone 2.

Age: Comobluffian age, Kimmeridgian Stage, Uppermost Malm Epoch, Late Jurassic.

Material:

SMA 0010: Skull and skeleton.

Note: Nicknamed "Barbara".

Number: Not given: A second skull and skeleton.

 

PEIRCE, 2006

Locality: V9138, Little Houston Quarry, approximately 20 km west of Sundance in southern Crook Country, just north of Little Houston Creek, northeast Wyoming.

Horizon: Member? Morrison Formation.

Biostratigraphy: Zone?

Age: Age? Stage? Epoch? Late Jurassic.

Material:

MWC 5822: Lower jaw.

 

TURNER & PETERSON, 1999

Locality: DMNH Quarry 4 (Carpenter’s 1990 Quarry or Valley of Death 5 Quarry), Fremont County, Colorado.

Horizon: Lower Upper Brushy Basin Member, Morrison Formation.

Biostratigraphy: Upper zone 2.

Age: Comobluffian age, Kimmeridgian Stage, uppermost Malm Epoch, Late Jurassic.

Material:

Number: Not given:

 

= Laosaurus gracilis MARSH, 1878
Etymology: Latin, gracilis, "slender."

Holotype: YPM 1875

Locality: Reeds YPM Quarry 9, (Mammal Quarry), T22N, R77W, Sec. 12, Como Bluff West (west of Marshall Road), Albany County, Wyoming.

Horizon: Brushy Basin Member, Morrison Formation.

Biostratigraphy: Zone 3.

Age: Comobluffian age, Kimmeridgian Stage, uppermost Malm Epoch, Late Jurassic.

Material: 13 dorsal centra, a partial left lower leg, a few pieces from the right pes, lateral distal tarsal, distal ends of metatarsals I and IV.

Referred material:

GILMORE, 1909

Locality: Quarry, Como Bluff, Albany County, Wyoming.

Horizon: Member?, Morrison Formation.

Biostratigraphy: Zone?

Age: Age? Stage?, Malm Epoch, Late Jurassic.

Material:

USNM 5808: Right femur.

 

OSTROM & McINTOSH, 1966

Locality: Marsh-Felch’s YPM Quarry 1, Garden Park, Sec. 28, T17S, R70W, Canyon City, Fremont County, Colorado.

Horizon: Salt Wash Member, Morrison Formation.

Biostratigraphy: Zone 2.

Age: Comobluffian age, Kimmeridgian Stage, uppermost Malm Epoch, Late Jurassic.

Material:

Number: Not given:

= Laosaurus consors MARSH, 1894
Etymology:n Latan, consors, "spouse."
= Othnielia consors (MARSH, 1894) GALTON, 1977
= Othnielosaurus consors (MARSH, 1894) GALTON, 2006

Holotype: YPM 1882

Locality: Reed’s YPM Quarry 7, (Three Trees Quarry), T22N, R77W, Sec. 18, Como Bluff West (west of Marshall Road), Albany County, Wyoming.

Horizon: Brushy Basin Member, Morrison Formation.

Biostratigraphy: Zone 3.

Age: Comobluffian age, Kimmeridgian Stage, uppermost Malm Epoch, Late Jurassic.

Material: Articulated skeleton on a slab mount and parts of a skeleton and a few duplicated bones.

Referred material:

= Laosaurus ?consors OSTROM & McINTOSH, 1966

= Othnielia rex GALTON, 1983

Locality: Reed's Quarry 9 (Mammal Quarry), Como Bluff, (Sec. 12, T22N, R77W), Albany County, Wyoming.

Horizon: Upper or Brushy Basin member, Morrison Formation.

Biostratigraphy: Zone 3.

Age: Comobluffian age, Kimmeridgian Stage, uppermost Malm Epoch, Late Jurassic.

Material:

YPM 9522: Premaxillary tooth.

AMNH 14328: Premaxillary tooth.

AMNH 11526: Maxillary tooth.

 

Locality: Unknown, Aurora Quarry, Como Bluff?, Albany County, Wyoming.

Horizon: Member? Morrison Formation.

Biostratigraphy: Zone?

Age: Age?, Stage?, Malm Epoch, Late Jurassic.

Material:

Cope Collection AMNH 2372: An isolated left dentary tooth of a juvenile.

 

Locality: Marsh-Felch’s YPM Quarry 1, Garden Park, Sec. 28, T17S, R70W, Canyon City, Fremont County, Colorado.

Horizon: Salt Wash Member, Morrison Formation.

Biostratigraphy: Zone 2.

Age: Comobluffian age, Kimmeridgian Stage, uppermost Malm Epoch, Late Jurassic.

Material:

USNM 5829: Isolated left dentary.

 

BAKKER, GALTON, SIEGWARTh & FILLA, 1990

Locality: Felch Quarry, Oil Creek, Colorado.

Horizon: Member? Morrison Formation.

Biostratigraphy: Zone?

Age: Age?, Stage?, Malm Epoch, Late Jurassic.

Material:

UCM Field No. C-1-1984: Maxillary tooth.

 

FULLER, 2009

Locality: Dana Quarry, Ten Sleep, Washakie County, Wyoming.

Horizon: Member?, Morrison Formation.

Biostratigraphy: Zone?

Age: Age? Stage?, Malm Epoch, Late Jurassic.

Material:

Number: Not given: Skeletal remains.

 

LEVITT, 2007

Locality: Westphal Quarry, near Hyattville, Big Horn Basin, Big Horn County, Wyoming.

Horizon: Member? Morrison Formation.

Biostratigraphy: Zone?

Age: Age? Stage? Late Jurassic.

Material:

Number: Not given: Left dentary, teeth, right femur.

 

= Drinker nisti BAKKER, GALTON, SIEGWARTH & FILLA, 1990
Etymology: Nisti masculine third declension genitive form of “nist” in horror of the National Institute of Standards and Technology.

Holotype: CPS 106

Locality: Drinker nisti Quarry –L & Quarry U (Lower and Upper Big Nose quarry), west Como Bluff, sec. 16, tp 22N, R77W, Albany County, Wyoming.

Horizon: Breakfast Bench Beds, 17 meters below the Dakota Sandstone, Upper Brushy Basin Member, Late Morrison Formation.

Biostratigraphy: Zone 4.

Age: Upper Part, Comobluffian age, Tithonian Stage, uppermost Malm Epoch, Late Jurassic.

Material: Partial subadult skeleton with parts of upper and lower jaws, centra from all sections of the column, partial fore and hind limbs.

Caudal centra.

Ankle bones.

Top, fragmentary metatarsals, bottom, phalanges.

Referred material:

CPS 107: Very young individual, about 1/4 adult size, vertebral centra, proximal radius, distal metatarsals III and astragalus.

CPS 108: 1/2 grown individual, vertebral centra, distal humerus, phalange of hind foot.
Note: Lower level of Big Nose.

CPS 109: 1/2 grown individual, dorsal vertebral centrum.

 

Locality: Op-KAT, East Como Bluff, sec 5, tp 22N, R76W, Albany County, Wyoming.

Horizon: Breakfast Bench Beds, 18 m below Dakota Sandstone, Upper Brushy Basin Member, Late Morrison Formation.

Biostratigraphy: Zone 4.

Age: Upper Part, Comobluffian age, Tithonian Stage, uppermost Malm Epoch, Late Jurassic.

Material:

CPS 197: Fully adult individual, dentary tooth, caudal vertebrae, partial phalanges of pes.

CPS 198: 1/5 grown femur head.

 

Locality: Quarry 9, West Como Bluff, Wyoming.

Horizon: 30 m below the Dakota Sandstone, Upper Brushy Basin Member, Late Morrison Formation.

Biostratigraphy: Zone 4.

Age: Upper Part, Comobluffian age, Tithonian Stage, uppermost Malm Epoch, Late Jurassic.

Material:

USNM 5808: Juvenile femur.

YPM 9524: Anterior maxillary tooth.

 

BAKKER, 1997

Locality: Como Bluff, Wyoming.

Horizon: Upper Brushy Basin Member, Morrison Formation.

Biostratigraphy: Zone 4.

Age: Upper Part, Comobluffian age, Tithonian Stage, uppermost Malm Epoch, Late Jurassic.

Material:

TATE 545: Tooth.

 

TURNER & PETERSON, 1999

Locality: Two Thigh Quarry, Como Bluff, Como Bluff East (East of Marshall road), Albany County, Wyoming.

Horizon: Upper Brushy Basin Member, Morrison Formation.

Biostratigraphy: Zone 4.

Age: Upper Part, Comobluffian age, Tithonian Stage, uppermost Malm Epoch, Late Jurassic.

Material:

Number: Not given:

 

Locality: Ninemile Hill Site, Medicine Bow Anticline (also called Flat top Anticline), Carbon County, Wyoming.

Horizon: Upper Brushy Basin Member, Morrison Formation.

Biostratigraphy: Zone 4.

Age: Upper Part, Comobluffian age, Tithonian Stage, uppermost Malm Epoch, Late Jurassic.

Material:

Number: Not given:

 

= Drinker? Sp

Locality: Lynn Quarry, Medicine Bow Anticline (also called Flat top Anticline), Carbon County, Wyoming.

Horizon: Brushy Basin Member, Morrison Formation.

Biostratigraphy: Zone 3.

Age: Upper Part, Comobluffian age, Kimmeridgian Stage, uppermost Malm Epoch, Late Jurassic.

Material:

Number: Not given: