Genus: Thescelosaurus GILMORE, 1913
Etymology: Greek, theskelos, “wonderful, astonishing, surprising” and
Greek, sauros, “lizard”: “Astonishing lizard”.
= Bugenasaura GALTON, 1995
Etymology: Latin, bu prefix, "large," Latin, gena (feminine form), "cheek," Greek,
saura (feminine form), "lizard"; an allusion to the massive ridges on the maxilla and
dentary for the attachment of a structure functionally analogous to the cheeks
of mammals.
= Thescelesaurus NOPCSA, 1929 (sic)
= Thescolosaurus HUNT & LUCAS, 1992 (sic)
Species: neglectus GILMORE, 1913
Etymology: Latin, neglectus, "overlooked."
= Thescelesaurus neglectus NOPSCA, 1929 (sic)
Holotype: USNM 7757
Locality: Dogie Creek, Niobrara County, Wyoming.
Horizon: Lance Formation.
Biostratigraphy:
Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.
Material: Skeleton lacking skull and cervicals.
Paratype: USNM 7758
Locality: Lance Creek, Niobrara County, Wyoming.
Horizon: Lance Formation.
Biostratigraphy:
Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.
Material: A few cervical, dorsal, and caudal vertebrae, portions of both scapulae, ribs, bones of fore and hind feet, and portions of limb bones.
Referred material:
BOYD, 2014
Locality: Dewey County, South Dakota.
Horizon: Hell Creek Formation.
Biostratigraphy:
Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.
Material:
TLAM.BA.2014.027.0001: Relatively complete, slightly transversely crushed skull missing the supraorbitals, accessary supraorbitals, right postorbital, right quadratojugal, most of the right jugal (anterior-most process preserved), and the entire lower jaws, left antlantal neural arch, the atlantal interentrum, 8 dorsal vertebrae, the sacrodorsal, 3 sacral vertebrae, 43 caudals, 6 partial dorsal ribs, 9 chevrons, left scapulocoracoid, partial right and left pubes, right ischium, partial proximal right tibia, incomplete distal ends of right and left tibiae, partial right astragulus, right calcaneum, distal ends of right metatarsals III and IV, 8 pedal phalanges, and additional unidentified material.
BRAMAN & EBERTH, 1987
Locality: Alberta Province, Canada.
Horizon: Scollard Formation, Uppermost Edmonton Group.
Biostratigraphy:
Age: Lancian Age, uppermost Maastrichtian Stage, uppermost Senonian subepoch, Upper Upper Gulf Epoch, Late Cretaceous.
Material:
NMC 8537:
ESTES, 1964
Locality: UCMP V-5620, Lull Quarry, near the head of a tributary of Buck Creek, Powder River Basin, Niobrara County (Formerly Converse County), Wyoming.
Locality: UCMP V-5711, Bushy Tailed Blowout, Quarry area includes the floor of a small blowout approximately 10 yards northwest of the highest point on the ridge forming the northeastern side of the blowouts of loc. V-5003, S 1/2, Sec. 20, T37N, R64W, Lance Creek, Powder River Basin, Niobrara County, Wyoming.
Locality: UCMP V-5815, Hatcher Number 2, western side of a cross-stratified sandstone erosional remnant near the center of a large blowout on the crest of a ridge, the distance form the nearest point on the road, approximately 150 yards south of the quarry, to the Johnson ranch is 0.4 miles, Eastern Wyoming.
Horizon: Lance Formation.
Biostratigraphy:
Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.
Material:
Numbers: Not given: Isolated teeth.
FISHER, RUSSELL, STOSKOPF, BARRICK, HAMMER & KUZMITZ, 2000 (Willow)
Locality: Harding County, Northwestern South Dakota.
Horizon: Hell Creek Formation.
Biostratigraphy:
Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.
Material:
NCSM 15728: Skull and nearly complete skeleton.
Note: With heart impression?
Note: Peter Galton believes this specimen to be a Bugenasaura and not a new species of Thescelosaurus.
Thescelosaurus neglectus (modified from FISHER, et al., 2000), NCSM 15728
GALTON, 1997
Locality: 30 km east of Limas, Montana.
Horizon: Hell Creek Formation.
Biostratigraphy:
Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.
Material:
AMNH 5020: A toothless dentary.
GILMORE, 1915Locality: Deer Ears Buttes, Butte County, South Dakota.
Horizon: Lance Formation.
Biostratigraphy:
Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.
Material:
USNM 7760: Left scapula and coracoid.
Locality: "Beecher's Quarry," Niobrara County, Wyoming.Horizon: Lance Formation.
Biostratigraphy:
Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.
Material:
USNM 7761: Cervical centrum.
Locality: Niobrara County, Wyoming.
Horizon: Lance Formation.
Biostratigraphy:
Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.
Material:
USNM 8065: Proximal phalanx of digit III of pes.
Locality: Dawson County, Montana.
Horizon: Lance Formation.
Biostratigraphy:
Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.
Material:
AMNH 8016, 5031: Three skeletons.
LONGRICH, HORNER, ERICKSON & CURRIE, 2010
Locality: Montana.
Horizon: Hell Creek Formation.
Biostratigraphy:
Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.
Material:
MOR 1161: Femur: Note: Has toothmarks from a Tyrannosaurus rex.
LUPTON, GABRIEL & WEST, 1980Locality: Chris's Bone bed, McCone County, Montana.
Horizon: Hell Creek Formation.
Biostratigraphy:
Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.
Material:
MPM 4377: Left femur, terminal phalanx, left ischium.
McINTOSH, 1981
Locality: Unknown.
Horizon:
Biostratigraphy:
Age: Late Cretaceous.
Material:
CM 9900: Two left humeri.
Locality: Lance Creek, Niobrara County, Montana.Horizon: Lance Formation.
Biostratigraphy:
Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.
Material:
CM 246: Dorsal centrum.
Locality: Sheep Mountain, Carter County, Montana.Horizon: Lance Formation.
Biostratigraphy:
Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.
Material:
CM 21787: 2 humeri, 2 ulnae, 2 radii, fragments of ilium, metatarsal, 6 phalanges of several individuals.
CM 38327: 3 dorsals, 10 phalanges, 4 unguals.
MORRIS, 1976
Locality: LACM locality V 3152, NE /4 of NW /2, sec. 12, T21N, R42E, Garfield County, Montana.
Horizon: Hell Creek Formation.
Biostratigraphy:
Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.
Material:
LACM 33543: Partial skull including frontals, parietal, partial crainum, squamosal, ectopterygoids, jugals and incomplete lower jaw, vertebral column lacking atlas and caudals (axis, 6 cervicals, 20 dorsals, and 5 sacral Vertebrae), numerous ribs, left and right ilia, ischia, and pubes, several metapodials, and phalanges of the pes and manus.
Note: Axis and first few cervicals show 4 scutes. According to Galton, these scutes belong to a crocodile.
= Ornithomimus sp ?OSBORN, 1917
Locality: Montana.
Horizon: Hell Creek Formation.
Biostratigraphy:
Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.
Material:
AMNH 5891:
STEIN, 2021
Locality: Tooth Draw Deposit, Butte County, South Dakota.
Horizon: Hell Creek Formation.
Biostratigraphy:
Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.
Material:
TD-12-050: Left scapula.
TD-16-174: Right ilium.
TD-10-015: Adult right femur.
TD-13-159 (NCSM 28349): Neonate left femur.
Numbers: Not given: 80 centra and neural arches, 65 phalanges, unguals and metatarsals and teeth.
STERNBERG, C. M. 1924
Locality: Badlands in Sec. 15, T1, R5W, 3rd principal meridian, Rocky Creek, Saskatchewan Province, Canada.
Horizon: Lance Formation.
Biostratigraphy:
Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.
Material:
Number: Not given: Fragmentary skeleton.
WEISHAMPEL, WEISHAMPEL, 1983/RUSSELL, 1964Locality: Southern Alberta Province, Canada.
Horizon: Scollard Member, Paskapoo Formation.
Biostratigraphy:
Age: Lancian Age, uppermost Maastrichtian Stage, uppermost Senonian subepoch, Upper Upper Gulf Epoch, Late Cretaceous.
Material:
Number: Not given:
_____________________________________________________________________________________
Species: edmontonensis STERNBERG,
1940
Etymology: In reference to Edmonton.
Holotype: NMC 8537
Locality: SE 1/4, sec. 35, T33, R22, W of 4, about 8 miles northwest of Rumsey, Red Deer River, Starland County, Alberta Province, Canada.
Horizon: Scollard Formation, Uppermost Edmonton Group.
Biostratigraphy:
Age: Lancian Age, uppermost Maastrichtian Stage, uppermost Senonian subepoch, Upper Upper Gulf Epoch, Late Cretaceous.
Material: Fragmentary skull that includes a partial cranium, articulated column from first dorsal to 13th caudal with dorsal ribs, pelvic arch, hind limbs and most metatarsals, and disarticulated frontals, postorbital, mandibular ramus, teeth, cervical vertebrae and ribs, scapulae, coracoids, humeri, ungular phalanx of pes and fragments.
Referred material:
ESTES, BERBERIAN & MESZOLEY, 1969
Locality: Bug Creek Anthills, McCone County, Montana.
Horizon: Hell Creek Formation.
Biostratigraphy:
Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.
Material:
MCZ 3649: Teeth.
_____________________________________________________________________________________
Species: garbanii MORRIS, 1976 emend BOYD,
BROWN, SCHEETZ & CLARKE, 2009
= Thescelosaurus? garbanii MORRIS,
1976
Etymology: In honor of Mr. Harley Garbani, the discover of the specimen.
Holotype: LACM 33542
Locality: LACM V 3152, T21N, R42E, NE /2, NW /4, Sec. 22, Garfield County, Montana.
Horizon: Hell Creek Formation.
Biostratigraphy:
Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.
Material: 5 posterior cervical and 11 anterior dorsal vertebrae, left pes, tarsus, tibia, fibula, and distal part of femur.
Referred material:
TRIEBOLD, 1997
Locality: Sandy Site, Harding County, South Dakota.
Horizon: Hell Creek Formation.
Biostratigraphy:
Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.
Material:
Number: Not given:
_____________________________________________________________________________________
Species: infernalis (GALTON, 1995) emend BOYD, BROWN, SCHEETZ & CLARKE,
2009
= Bugenasaura infernalis GALTON, 1995
Etymology: Latin, infernalis, "belonging to the lower regions," an allusion to
the occurrence in the Hell Creek Formation.
Holotype: SDSM 7210
Locality: Olson Locality, SDSM number V729, Harding County, South Dakota.
Horizon: Hell Creek Formation.
Biostratigraphy:
Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.
Material: Left half of cranium, left dentary and fragment of right dentary, centra of 2 dorsal vertebrae, 2 dorsal vertebrae lacking transverse process and neural spines and 2 manual phalanges.
Referred material:
GALTON, 1997
Locality: Lusk, Niobrara County, Wyoming.
Horizon: Lance Formation.
Biostratigraphy:
Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.
Material:
YPM 8098: Right dentary tooth.
_____________________________________________________________________________________
Species: assiniboiensis BROWN, BOYD & RUSSELL, 2011
Etymology: Named for the District of Assiniboia, a regional administrative unit
of the North-West Territories, Canada, from 1882 to 1905 (located between 49° adn
51.97°N, and ~101.5 and ~111.5°W. The majority of this district became
the southrn portion of the modern province of Saskatchewan, with the westernmost
area becoming the easternmost portion of the province of Alberta. It encloses
the exposures of the Frenchman Formation. This district was named after the
Assinboine First Nations People.
= Thescelosaurus neglectus GALTON, 1989
Holotype: RSM P 1225.1
Locality: North side of the Frenchman River Valley on the north-west facing
side of a butte extending from the valley wall. LSD 11, Sec 2, T5, R 19, west
of the 3rd Meridian in southwestern
Saskatchewan Province, Canada.
Note: Was originally stated as from Northwest of Clarks Ranch, Frenchman
River Valley, south of Shanovan, LSD 7, Sec. 2,
T5,
R19,
W. 3rd, Map. Ref. 72F8, Saskatchewan Province, Canada.
Note: Exact Locailty
information is available from the RSM pon request.
Horizon: Frenchman Formation.
Biostratigraphy:
Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.
Material: Partial skeleton and fragmentary skull that includes a palpebral, incomplete frontals, parietal, squamosal, postorbital, pterygoid, dentary and cranium, and endocast, dorsal, sacral and caudal vertebral series, dorsal ribs, pelvic girdles, and hindlimbs.
