Genus: Camarasaurus COPE, 1877
Etymology: Greek, kamara, “chamber”, and Greek, sauros, “lizard”: “Chamber lizard”.
= Camacasaurus YOUNG, 1958 (sic)
= Camaeosuarus COPE, 1877 (sic)
= Camarasuarus BERMAN & McINTOSH, 1978 (sic)
= Camarosaurus OWEN, 1878 (sic)
= Camasaurus McINTOSH, 1977 (sic)
= Cathetosaurus JENSEN, 1988 (nomen nudum)
Etymology: Greek, kathetos, "perpendicular," referring to an ability to stand erect on its rear legs, and Greek, sauros, “lizard”: “Upright lizard”.
= Caulodon COPE, 1877 (nomen dubium)
Etymology: Greek, kaulos, “stalk”, and Greek, odon, “tooth”: “Stalk tooth”.
= Monosaurus BERMAN & McINTOSH, 1978 (sic)
= Morasaruus RIGGS, 1904 (sic)
= Morosaurus MARSH, 1878
Etymology: Greek, moros, “stupid” and Greek, sauros, “lizard”: Stupid lizard”.
= Mososaurus BAUR, 1884 (sic)
= Uintasaurus HOLLAND, 1919
Etymology: In reference to the Untia County, Utah and Greek, sauros, “lizard”: “Uinta lizard”.

Species: supremus COPE, 1877
Etymology: Latin, supremus, "above, upper, high."

Holotype: AMNH 5760

Locality: Cope’s Quarry 8 (about 500 feet southwest of Cope’s Nipple), Garden Park, Canon City, Fremont County, Colorado.
Note: Cope also referred it to Camarasaurus Quarry 2 and Osborn and Mook 1921, incorrectly called it Cope Quarry 1.

Horizon: Upper Brushy Basin Member, Morrison Formation.

Biostratigraphy: Zone 4.

Age: Upper Part, Comobluffian age, Tithonian Stage, Uppermost Malm Epoch, Late Jurassic.

Material: Cervical 10, 3 dorsal and 4 caudal vertebrae.

Cotype:

AMNH 5760": Dorsal vertebrae, dorsal centrum, right scapula, right coracoid, left pubis, portion of right ilium, caudal vertebrae 1-5, 7-9, 11-13, 15-18, 19-20, 22-23, 24, right scapula, left pubis, right tibia.

AMNH 5760': Caudal vertebrae 2-9, 12, 13, and centrum, dorsal vertebrae 1-4, 6-10, D-S, caudal vertebrae 1, 4-6,10-11, 14, 21, 27-29, 31-33, 35-38, chevrons 4, 8-9, 11, 13, 15-16, fragmentary ribs, right scapula, left ilium, ischia, right pubis, left tibia, right fibula.

AMNH 5761: Maxilla, proximal end of dentary and posterior part of skull, cervicals 6, 8-9, 11, dorsal neural spine, fragmentary dorsal vertebra, dorsal vertebrae 1, 3-6, 8-10, sacrum, ilia, pubes, and ischia, caudal vertebrae 1-8, 10-11, 14-18, 19, 22-25, 28-30, 34, 39, 42-44, chevron 1-2, 4-7, 10, sternal plates, fragmentary ribs, ventral ribs (gastralia), 3 left and 2 right scapulae, left and right coracoids, humeri, left metacarpal I, II, right tibia, right fibula, astragalus, right metatarsal III.
Note: Cervicals 11 and 12 are fused together and are pathologic in nature.

AMNH 5761a: Cervical vertebrae 1, 3, 5, 9-13, cervical ribs, dorsal vertebrae 1, 3-6, D-S, right ischium, left femur.

Referred material:

AMNH 5763: Cervical 7.

AMNH 5765: Caudal vertebra 15, left femur.

TIDWELL & CARPENTER, 1997

DMNH 27228: A well-preserved axis, a pubis, ischium, and anterior caudal vertebrae, and several other bones (33 in all).

BONAM, 2000

KUVP 129716: Fragmentary skeleton (with a calcaneum).

 

CARPENTER, 1988

Locality: Cope’s Quarry 1-7, at Cope’s Nipple or Saurian Hill, Fremont County, Colorado.

Horizon: Upper Brushy Basin Member, Morrison Formation.

Biostratigraphy: Zone 4.

Age: Upper Part, Comobluffian age, Tithonian Stage, Uppermost Malm Epoch, Late Jurassic.

Material:

Number: Not given:

 

GLUT, 2006

Locality: Waugh Ranch, near Hullett, Crook County, Wyoming.

Horizon: Member? Morrison Formation.

Biostratigraphy: Zone?

Age: Age? Stage?Upper Malm Epoch, Late Jurassic.

Material:

BHI 6200: Skull and skeleton.

 

LUCAS & HUNT, 1985

Locality: NE 1/4, SE 1/4, SW 1/4, Sec. 22, T15N, R1E, Sandoval County, New Mexico.

Horizon: Brushy Basin Member, Morrison Formation.

Biostratigraphy: Zone?

Age: Age? Stage?, Malm Epoch, Late Jurassic.

Material:

NMMNH P-21094: Right pubis, posterior dorsal vertebra, a middle dorsal vertebra, a right rib and various skull fragments.

 

RIGBY, 1982

Locality: San Ysidro Quarry or Camarasaurus quarry, San Ysidro, NE 1/4, SE. 1/4, SW 1/4, S22, T15N, R1E, Sandoval County, New Mexico.

Horizon: Brushy Basin Member, Morrison Formation.

Biostratigraphy: Zone 3.

Age: Upper Part, Comobluffian age, Kimmeridgian Stage, Uppermost Malm Epoch, Late Jurassic.

Material:

Number: Not given: Fragmentary skeleton.

 

SUNDELL, 2002, MARTIN, BURNHAM & LIM, 2002

Locality: Weston County, Wyoming.

Horizon: Member? Morrison Formation.

Biostratigraphy: Zone?

Age: Age? Stage?Upper Malm Epoch, Late Jurassic.

Material:

University of Kansas: 2 articulated adults and a sub adult specimen.
Note: Some skin impressions were found.

 

WHITE, 1978

Locality: Bone Cabin Quarry, 8 miles north of Como, Medicine Bow Anticline (also called Flat top Anticline), Albany County, Wyoming.

Horizon: Salt Wash Member, Morrison Formation.

Biostratigraphy: Zone 2.

Age: Comobluffian age, Kimmeridgian Stage, Uppermost Malm Epoch, Late Jurassic.

Material:

AMNH 467: Occiput lacking the paroccipital processes, braincase lacking the anterior portion, both pterygoids, right quadrate, part of left quadratojugal, both frontals, right postfrontal, both nasals, both jugals, both maxilla, and premaxilla, and both lower jaws.

MADSEN, McINTOSH & BERMAN, 1995

AMNH 607: Right dentary.

AMNH 611: Right maxilla.

AMNH 618: Both maxillae.

AMNH 657: Left dentary.

AMNH 673: Cranium.

AMNH 677: Right premaxilla.

AMNH 6126: Cranium.

Compsite skull.


= Amphicoelias latus COPE, 1877 (Juvenile?)
Etymology: Latin, latus, "wide."

Holotype: AMNH 5765

Locality: Garden Park, 8 miles north to northeast of Canyon City, Fremont County, Colorado.

Horizon: Member?, Morrison Formation.

Biostratigraphy: Zone?

Age: Age? Stage?, Malm Epoch, Late Jurassic.

Material: Right femur, 4 caudal vertebrae.

 


= Caulodon diversidens COPE, 1877 (nomen dubium)
Etymology: Turning in different directions.

Holotype: AMNH 5768

Locality: Garden Park, Canyon City, Fremont County, Colorado.

Horizon: Member?, Morrison Formation.

Biostratigraphy: Zone?

Age: Age? Stage?, Malm Epoch, Late Jurassic.

Material: 10 teeth.

 


= Caulodon leptoganus COPE, 1878 (nomen dubium)
Etymology:

Holotype: AMNH 5769

Locality: Garden Park, 8 miles north to northeast of Canyon City, Fremont County, Colorado.

Horizon: Member?, Morrison Formation.

Biostratigraphy: Zone?

Age: Age? Stage?, Malm Epoch, Late Jurassic.

Material: 2 teeth.

 

= Camarasaurus leptodirus COPE, 1879 (nomen dubium)
Etymology:

Holotype: AMNH 5763

Locality: Garden Park, 8 miles north to northeast of Canyon City, Fremont County, Colorado.

Horizon: Member?, Morrison Formation.

Biostratigraphy: Zone?

Age: Age? Stage?, Malm Epoch, Late Jurassic.

Material: 3 cervical vertebrae and other remains.

_____________________________________________________________________________________

Species: lentus (MARSH, 1889) MOOK, 1914
= Morosaurus lentus MARSH, 1889
Etymology: Latin, lentus, "tough."

Holotype: YPM 1910

Locality: Lakes' YPM Quarry 1A, (Big Canyon Quarry), T22N, R77W, Sec. 17, Como Bluff, Wyoming.
Locality: Reed’s YPM Quarry 1, T22N, R77W., sec. 10, Como Bluff West (west of Marshall Road), Albany County, Wyoming.

Horizon: Brushy Basin Member, Morrison Formation.

Biostratigraphy: Zone 3.

Age: Upper Part, Comobluffian age, Kimmeridgian Stage, Uppermost Malm Epoch, Late Jurassic.

Material: 4th cervical, 2nd caudal to 8th caudal, left scapula, both tibia, left hind foot, partial right coracoid, both humeri, left ischium, and left femur of a juvenile.

Referred material:

= Camarasaurus lentus MADSEN, McINTOSH & BERMAN, 1995

USNM 7759: Right premaxilla.

USNM 7944 1: Maxilla, incomplete supraoccipital, right exoccipital, right opisthotic, occipital condyle, incomplete right dentary not numbered but probably belonging to the same skull.

 

= Camarasaurus lentus GILMORE, 1925

Locality: Quarry face as exposed inside the Dinosaur National Monument Visitor Center, Carnegie Quarry, Vernal, Uintah County, Utah.

Horizon: Brushy Basin Member, Morrison Formation.

Biostratigraphy: Zone 3.

Age: Upper Part, Comobluffian age, Kimmeridgian Stage, Uppermost Malm Epoch, Late Jurassic.

Material:

CM 11338: Skull and skeleton of a juvenile.
Note: A possible patch of skin was found in-between some ribs.

Referred material at DNM.

CM 11373 (DNM 300 & 301) (It was transferred to the National Museum of Natural History, USNM 13786): Skull and skeleton, articulated and complete except for the tail.

CM 30743 (DNM 300): Cast of skull and mandible of the above specimen.

CM 3379 (DNM 130, part) (It was transferred to the National Museum of Natural History, USNM 15492, and used to complete the mounted specimen USNM 13786): 47 caudals complete to most of the tip of tail.

CM 11393 (DNM 240): Disarticulate skeleton of an adult animal consisting of the partial skull and mandible, 5+ cervicals, 10+ dorsals, sacrum, 25 caudals, many ribs, several chevrons, scapulae, coracoids, right humerus, right radius, right ulna, part of manus, ilia, pubes, ischia, femora, left fibula, calcaneum, 2 metatarsals, 3 phalanges, 2 claws. Skull later catalogued as CM 12020. 2 articulated cervicals were transferred to the University of Michigan V 16995)

CM 12020: Partial skull and mandible found among the bones of the skeleton DNM 240/Lk. It was initially given the catalogue number of the remainder of the skeleton, 11393, but was later recatalogued 12020, and a cast of it was used until recently to complete the headless skeleton of Apatosaurus louisae.

CM 21751 (DNM 201/A and B): Right premaxilla, both rami and mandible, many teeth of a large individual found in the same quarry as the above and quite near it, but in a distinctly higher stratum the usual bone layer in which the above specimens were found.

CM 11969 (DNM 325): Disarticulated skull only partially worked out consisting of the complete braincase, both postorbitals, both maxillae, left premaxilla, one pterygoid, both dentaries, both surangulars, right prearcticular and other elements not yet prepared and part of the neck.

CM 21732 (DNM 201/C, D., E., and F): A maxilla, dentary, quadratojugal and fragments of a poorly preserved skull, associated with but distinct from CM 21751.

CM 3381 (DNM 3): A large tooth.

CM 21702(DNM 39/60a): Right maxilla without teeth of a juvenile.

CM 21701 (DNM 39/65): Cervical.

CM 30760: 2 1/2 dorsals found with DNM 40)

CM 33955 (DNM 101): Dorsal vertebra.

CM 21733 (DNM 344): Several posterior dorsals articulated with the sacrum and right ilium. A few centra were transferred to the Everart Museum in Scranton, Pennsylvania.

CM 21712 (DNM 45): 13 articulated caudal vertebrae, the arches as yet unprepared.

CM 37001 (DNM 302, part): 2 neural arches of anterior caudals or sacrals.

CM 21716: 5 caudal centra (marked, apparently incorrectly as DNM 60).

CM 30769 (DNM 70/4): 2 caudals.

CM 30771 (DM 210): Caudal arch and spine of a juvenile.

CM 33896 (DNM 39, part): Caudal.

CM 33909 (DNM 232, part): Caudal.

CM 38342 (DNM 102, part): Caudal.

CM 33925 (DNM 210/2 and 6): Right scapula and coracoid.

CM 33929 (DNM 270/1): Right scapula, 3 cervicals.

CM 38320 (DNM 39/13): Large left humerus, more robust than usual, but probably to slender of Apatosaurus.

CM 21781 (DNM 45/21): Left humerus perhaps same individual as tail CM 21712.

CM 33972 (DNM 120, part): Right forelimb and foot.

CM 33971 (DNM 210/6): Head of right humerus, the remainder of the bone is on exhibit at the California Academy of Science, San Francisco, California.

CM 33979 (DNM 102, part): Right humerus, both ends.

CM 38334 (DNM ?39): Distal end of right humerus.

CM 33948 (DNM 232/23): Distal end of right humerus.

CM 33949 (DNM 232/17): Right radius.

CM 37003 (DNM 232 part): Metacarpal. Might belong to skeleton CM 11393 but found 20 feet away from it.

CM 21735 (DNM 147): Left ilium of a juvenile.

CM 33891 (DNM 130/10): Right ilium.

CM 21750: Left femur lacking the head. A part of head received the separate catalogue number 11971 (DNM 155.Fe).

CM 21759 (DNM 167) (originally catalogued as part of CM 9000): A small left femur.

CM 21772 (DNM 350/32): A left femur of a very small individual, found associated with Stegosaurus skeleton CM 11341. It perhaps belongs to this species. An equally small dorsal centrum and right coracoid found associated may belong to the same animal.

CM 33973 (DNM 210/20): Right femur, perhaps belongs to CM 11069. The proximal end of the bone is at the California Academy of Science, San Fransco, California.

CM 21777 (DNM 270, part): Left tibia, astragalus, metatarsal I, claw.

CM 21783 (DNM 150/45): Right tibia, found with a skeleton of Diplodocus but not belonging to it. The right fibula of the same limb is in the Denver Museum of Natural History where it was transferred with skeleton DNM 150.

CM 33956: Lower half of a small right tibia, with DNM 130/D.

CM 33927 (DNM 232/2): Right tibia.

CM 33958 (DNM 232, part): 2 claws.

Number: Not given (WHITE, 1958): Skull, cervical vertebrae and 6 dorsals with ribs.

CHATTERJEE & ZHONG, 2005

DINO 28: Disarticulated skull.

MADSEN, McINTOSH, & BERMAN, 1995

DNM 28: Greater part of a disarticulated skull and mandibles lacking the left lachrymal, palatines, vomers, and all medial elements of the mandibles except the left prearticular.

DNM 975: the “cliff skull” has been left in place on the quarry face, with the right side prepared in relief. All the external elements of the right side are articulated except the quadratojugal, which has been displaced ventrally, and the quadrate, which has not been found. The posterior and dorsal surfaces of the skull are exposed, showing the supraocciptial, exoccipitals, opisthotics, basiocipital, left quadrate, left quadratogual, parietals, frontals and nasals, in position. Many of the bones of the palate are missing , and those present have been more or less displaced. The right pterygoid and ectopterygoid have been displaced ventrally over the medial surface and the left mandible. The left pterygoid and right palatine have been removed and prepared. The articulated dentary, angular, and surangular of the right mandible, as well as the disarticulated right prearticular, angular and splenial have also been removed from the quarry and fully prepared.

Camarasaurus lentus (after MADSEN, McINTOSH & BERMAN, 1995), DINO 975.

DNM 1009: Probably includes the greater part of the skull that is exposed on the quarry face and referred to as the “hump skull.” Some elements displaced by compression against lower end of a right sauropod femur and probably lost. Exposed elements include the left articular, left prearticular, left angular, left surangular, left quadrate, left pteryoid, left ectopterygoid, right squamosal, right opisthotic, right prootic, right laterosphenoid, right pterygoid, supraocciptial, and right parietal.

DNM 32-37, 948-951, 953-971: Teeth.

DNM 4011: Right dentary.

DNM 3008: Partial, articulated skull questionably belonging to Camarasaurus consisting of right prefrontal, frontals, parietals, postoribitals, squamosals, laterospheonids, prootics, opisthotics, exoccipitals, and supraocciptial.

DNM 3699: Right premaxilla.

DNM 4257: Right maxilla.

BYU Carnegie Museum field no. 320/C given to BYU: Right maxilla.

MEYERS & HOOPS, 1993

DINO 3524: Left scapula.

DINO 4209: Left tibia of a juvenile.

DINO 42587: Right maxilla.

DINO 4269: Cervical vertebra.

DINO 4801-4806: Caudal vertebrae.

DINO 3818: Tibia.

DINO 4885: Phalange of a juvenile.

DINO: Skull and dentaries, unnumbered?

DINO 4395: Atlas.

DINO 4396-4403: Cervical vertebrae.

DINO 4406-4410: Dorsal ribs.

DINO 4411-4445: Caudal vertebrae.

DINO 4446-4458: Dorsal ribs.

DINO 4459: Scapula.

DINO 4461: Scapula.

DINO 4462: Coracoid.

DINO 4463: Humerus.

DINO 4464: Ulna.

DINO 4465: Radius.

DINO 4466-4470: Metacarpals.

DINO 4471-4473: Phalanges.

DINO 4511: Pubis.

DINO 4512, 4513: Ischium.

DINO 4515: Femur.

DINO 4516: Tibia.

DINO 4517: Fibula.

 

Locality: Jensen Quarry, near Dinosaur National Monument, Jensen, Utah.

Horizon: Member?, Morrison Formation.

Biostratigraphy: Zone?

Age: Age? Stage?, Malm Epoch, Late Jurassic.

Material:

BYU 9048: Complete braincase and portions of upper and lower jaws.

 

IKEJIRI, TIDWELL, & TREXLER, 2005

Locality: BS-Quarry & S-Quarry, Warm Springs Ranch, approximately 7 km southeast of the town of Thermopolis, Hot Springs County, Wyoming.

Horizon: Brushy Basin Member, Morrison Formation.

Biostratigraphy: Zone 3.

Age: Comobluffian age, Kimmeridgian Stage, Uppermost Malm Epoch, Late Jurassic.

Note: Just below the Tithonian-Kimmeridgian Stage Boundary, Malm Epoch, Late Jurassic.

Material:

WDC BS (no number): Frontal.

WDC BS-13: Left radius.

WDC BS-14: Sacrum.

WDC BS-101: Fibula.

WDC BS-104, 108: Sternal plates.

WDC BS-104, 179: Large scapulacoracoids.

WDC BS-152, 332, 192, 113, 263, 144, 10, 142, 141, 2 uncatalogued: Anterior caudals.

WDC BS-156: Right humerus.

WDC BS-184, 185, 270: 3 posterior dorsal vertebrae.

WDC BS-198, 255: Axis.

WDC BS-206, 255, 268, 177, 265, 210: Anterior cervical vertebrae.

WDC BS-206, 255, 268, 188, 159, 271, 346, 347: Proximal dorsals.

WDC BS-261, 259, 260, 252: Mid dorsal vertebrae.

WDC BS-208, 555, 382, 193: Posterior cervical vertebrae.

WDC BS-217: Right tibia.

WDC BS-199: Right fibula.

WDC BS-224: Chevron.

WDC BS-258: Atlas.

WDC BS-263: 2 fused caudals.

Note: Pathologic?

WDC BS-313: Ulna.

WDC BS-315: Left radius.

WDC BS-328: Postier dorsal vertebra.

WDC BS-388: Complete braincase.

WDC BS-15, 407: 2 partial nasals.

WDC BS-405, 435: 2 right quadrates.

WDC BS-189: A parital right maxilla.

IKEJIRI, 2005

WDC A: Jaw fragments, right quadrate, all disarticulated presacral vertebrae except dorsal vertebra no. 11 (?), sacrum, some anterior caudal verterae, scapulocoracoids, left humerus, left ulna, radius, ischium, pubes, femora and fibula.

WDC B: Braincase, jaw fragments, right quadrate, 4 anterior-mid-cervical vertebrae, 4 anterior and a few posterior dorsal vertebrae, left scapulocoracoid, left humerus, right ulna, right radius, pubes, ischia, and femora.

WDC specimens: Both sides of upper and lower jaws, frontal, lacrimal, jugal, many cranial elements, 2 sternal plates, 2 carpals, 8 metacarpals, many cervical and thoracic ribs, adn chevrons (including the first chevron).

IKEJIRI, WATKINS & GRAY, 2006

WDCB: BS-178: Anterior dorsal vertebra.

WDC A?:BS-224: First chevron.

WDC B:BS-724: Right humerus.

WDC B:GS-312: Left ulna..


= Camarasaurus lentus STOKES

Locality: Cleveland-Lloyd Dinosaur Quarry, approximately 20 miles south of Price, SE 1/4, SE 1/4, Sec. 21, T17S, R11E, SLBM. lat 39°19’00", Long. 110°41’15"W, San Rafael Swell Area, Carbon County, Utah.

Horizon: Brushy Basin Member, Morrison Formation.

Biostratigraphy: Zone 3.

Age: Upper Part, Comobluffian age, Kimmeridgian Stage, Uppermost Malm Epoch, Late Jurassic.

Material:

CEU 1694: Semi-articulated composite (?) partial skeleton, cervical vertebrae, sacrum, many caudal vertebrae, scapulae, limb elements adn pelvic bones.

(Note: This specimen may include more than one individual, IKEJIRI, 2005)

= Camarasaurus lentus CHURE, FIORILLO & JACOBSON, (1998) 2000

UUVP 5309: A partial pelvis and some limb bones.
Note: 3 areas of the ilium have multiple grooves from bite marks.

 

= Uintasaurus douglassi HOLLAND, 1919
Etymology: In honor of Mr. Earl Douglass.
= Camarasaurus douglassi (HOLLAND, 1919) OLSHEVSKY, 1978

Holotype: CM 11069

Locality: Quarry face as exposed inside the Dinosaur National Monument Visitor Center, Carnegie Quarry, Vernal, Uintah County, Utah.

Horizon: Brushy Basin Member, Morrison Formation.

Biostratigraphy: Zone 3.

Age: Upper Part, Comobluffian age, Kimmeridgian Stage, Uppermost Malm Epoch, Late Jurassic.

Material: 5 anterior cervical vertebrae. The last two nearly complete.

Referred material:

USNM 13786 (old CM 11373): A complete skeleton except for the tail.

Camarasaurus lentus (after MADSEN, McINTOSH & BERMAN, 1995), CM 13786.

 

= Camarasaurus annae ELLINGER, 1950
Etymology: In honor of Miss Polly Ann Ratliff, of Natchez, Mississippi.
= Camarosaurus annae ELLINGER, 1950 (sic)

Holotype: CM 8942 (DNM 130/2)

Locality: Quarry face as exposed inside the Dinosaur National Monument Visitor Center, Carnegie Quarry, Vernal, Uintah County, Utah.

Horizon: Brushy Basin Member, Morrison Formation.

Biostratigraphy: Zone 3.

Age: Upper Part, Comobluffian age, Kimmeridgian Stage, Uppermost Malm Epoch, Late Jurassic.

Material: A complete anterior dorsal vertebra.

Referred material;

DNM 130/2: Another dorsal of about the same size.

Note: These dorsals were found only 4 feet from the cervicals of CM 11069, and in all probability belong to the same individual, and these Camarasaurus vertebrae lay across an articulated series of Apatosaurus presacral vertebrae, CM 33916, many different limb and girdle elements belonging to several genera were found nearby and assigned to the same field number as the latter, DNM 210, but some may belong with this animal CM 33916.

_____________________________________________________________________________________

Species: grandis (MARSH, 1877) GILMORE, 1925
Etymology: Latin, grandis, "full-grown, grown up, large," in reference to large size of this taxon.
= Apatosaurus grandis MARSH, 1877
= Morosaurus grandis (MARSH, 1877) WILLISTON, 1898

Holotype: YPM 1901

Locality: Reed’s YPM Quarry 1, T22N, R77W., sec. 10, Como Bluff West (west of Marshall Road), Albany County, Wyoming.

Horizon: Brushy Basin Member, Morrison Formation.

Biostratigraphy: Zone 3.

Age: Upper Part, Comobluffian age, Kimmeridgian Stage, Uppermost Malm Epoch, Late Jurassic.

Material: Basioccipital, basisphenoid, left scapula, left coracoid, left humerus, left radius, left ulna, one carpal and metacarpal III, right scapulocoracoid, left sternal plate and both femora, several dorsal vertebrae, ribs, partial sacrum, articulated caudal vertebrae 1-27, and perhaps nine, disarticulated others,

Note: The ilium assigned to Marsh might belong to a smaller individual.

Paratypes: YPM 1905: Occiput and frontal region, most of braincase, both quadrates and parts of both quadratojugals, most of both ptergyoids, frontals, parietals, postorbitals, squamosals, quadrates, quadratojugals, pterygoids, left ectopterygoid, supraoccipital, exoccipitals, opisthotics, prootics, laterosphenoids, orbitosphenoids, basiocciptial, basisphenoid, left maxilla with accending process, most of both dentaries and surangulars, complete cervical series, most or all dorsal vertebra, incomplete sacrum, at least 12 caudal vertebrae and possibly others, ribs, scapulacoracoids, left humerus, right ulna, ?ilia, right ischium, femora, tibiae, fibulae, and greater parts of the pedes.

Camarasaurus grandis (after MADSEN, McINTOSH & BERMAN, 1995) Paratype: YPM 1905.

Note: ‘...Reed recovered a femur and caudal vertebrae from near the other specimens.. which probably belong to either YPM 1905 or an as yet unidentified individual’ (MADSEN, McINTOSH & BERMAN, 1995)

YPM 1903: Left pubis and right scapula.

Referred material:

McINTOSH, 1981

CM 21760: Cast of YPM 1901, 1905, 1903, and 1900.

 

FOSTER, 2005

Locality: Mygatt-Moore (M&M) Quarry, Mesa County, Western Colorado.

Horizon: Middle Brushy Basin Member, Morrison Formation.

Biostratigraphy: Zone 3.

Age: Comobluffian age, Kimmeridgian Stage, Uppermost Malm Epoch, Late Jurassic.

Material:

MWC 3630: Dorsal centrum of a juvenile.

 

Locality: Kings View Quarry, about 3 km south of the town of Fruita near the Colorado River, Mesa County, Western Colorado.

Horizon: Middle Brushy Basin Member, Morrison Formation.

Biostratigraphy: Zone 3.

Age: Comobluffian age, Kimmeridgian Stage, Uppermost Malm Epoch, Late Jurassic.

Material:

MWC 2538: 2 cervical vertebrae, 3 dorsal vertebrae, a cervical rib?, and a dorsal rib of a juvenile.


McINTOSH, MILES, CLOWARD & PARKER, 1996

Locality: East Camarasaurus Quarry, about 150 meters east of Bone Cabin Quarry, Albany County, northeast of Medicine Bow, Wyoming.

Horizon: Brushy Basin Member, Morrison Formation.

Biostratigraphy: Zone 3.

Age: Upper Part, Comobluffian age, Kimmeridgian Stage, Uppermost Malm Epoch, Late Jurassic.

Material:

Gunma Museum of Natural History, GMNH 101 (Western Paleontological Laboratories, Inc, 1995): Approximately 65% complete skeleton, consisting the large part of an articulated skeleton, which is in many ways quite unusual. Preserved are the skull and mandible, cervical vertebrae 1-8, dorsal neural arches 6-12, most of the dorsal ribs, the articulated sacrum lacking centra, a nearly complete set of caudal vertebrae lacking only the centra of numbers 1 and 2, 18 chevrons, part of the right scapula, left coracoid, left humerus and fragments of the right, partial left ulna, partial left manus, right ilium, pubes, ischia and nearly complete right hind limb and pes, head of left femur and part of left pes.

Note: Caudals 25, 26, 32, 33, 39, 40, 43, 46, 47, and possibly 50-53, are pathologic. Metatarsal II has a stress fracture (ROTHSCHILD & MOLNAR, 2005)

 


McWHINNEY, CARPENTER & ROTHSCHILD, 2001

Locality: Near Canon City, Colorado.

Horizon: Member? Morrison Formation.

Biostratigraphy: Zone?

Age:? Age? Stage? Upper Malm Epoch, Late Jurassic.

Material:

DMNH 2908: A nearly complete right humerus, associated dorsal vertebra and several caudal centra.
Note: The humerus shows a juxtacortial lesion pathology.

 

RIGGS, 1901

Locality: Quarry No. 12, Grand Valley, Near Fruita, Mesa County, Colorado.

Horizon: Member?, Morrison Formation.

Biostratigraphy: Zone?

Age: Age? Stage?, Malm Epoch, Late Jurassic.

Material:

Colombian Field Museum No. 6668: Right scapula, coracoid, humerus, ulna, radius and fore foot.

 


Locality: Grand River Valley, Colorado.

Horizon: Member?, Morrison Formation.

Biostratigraphy: Zone?

Age: Age? Stage?, Malm Epoch, Late Jurassic.

Material:

Colombian Field Museum No. 6670: Scapula-coracoid.

 


OSTROM & McINTOSH, 1966

Locality: Reed’s YPM Quarry 3, T22N, R77W, Sec. 11, Como Bluff West (west of Marshall Road), Albany County, Wyoming.

Horizon: Brushy Basin Member, Morrison Formation.

Biostratigraphy: Zone 3.

Age: Upper Part, Comobluffian age, Kimmeridgian Stage, Uppermost Malm Epoch, Late Jurassic.

Material:

YPM 1912: Postfrontal.

 

TURNER & PETERSON, 1999

Locality: DMNH Quarry 3 (Lindsey’s 1977 Quarry), Fremont County, Colorado.

Horizon: Member?, Morrison Formation.

Biostratigraphy: Zone 2.

Age: Comobluffian age, Kimmeridgian Stage, Uppermost Malm Epoch, Late Jurassic.

Material:

Number: Not given:

 

 

WILLISTON, 1898

Locality: Not known, but probably north of Lusk, Converse (now Niobrara) County, Wyoming.

Horizon: Member?, Morrison Formation.

Biostratigraphy: Zone?

Age: Age? Stage?, Malm Epoch, Late Jurassic.

Material:

KUVP 1354: Sacrum.

 


= Morosaurus impar MARSH, 1878
Etymology: Latin, impar, "odd."
= Camarasaurus impar (MARSH, 1878) STEEL, 1970

Holotype: YPM 1900

Locality: Reed’s YPM Quarry 1, T22N, R77W., sec. 10, Como Bluff West (west of Marshall Road), Albany County, Wyoming.

Horizon: Brushy Basin Member, Morrison Formation.

Biostratigraphy: Zone 3.

Age: Upper Part, Comobluffian age, Kimmeridgian Stage, Uppermost Malm Epoch, Late Jurassic.

Material: Sacrum. (Possibly incomplete, paired coracoids and ischia, several vertebrae and left femur, and might belong to YPM 1903; MADSEN, McINTOSH & BERMAN, 1995).

 

= Morosaurus robustus MARSH, 1878 (nomen dubium)
Etymology: Latin, robustus, "robusts, strong."
= Camarasaurus robustus (MARSH, 1878) WHITE, 1958 (nomen dubium)

Holotype: YPM 1902

Locality: Reed’s YPM Quarry 1, T22N, R77W., sec. 10, Como Bluff West (west of Marshall Road), Albany County, Wyoming.

Horizon: Brushy Basin Member, Morrison Formation.

Biostratigraphy: Zone 3.

Age: Upper Part, Comobluffian age, Kimmeridgian Stage, Uppermost Malm Epoch, Late Jurassic.

Material: Left ilium.

Note: Not Camarasaurus?

Referred material:

RIGGS, 1901

Locality: Freeze-out Hills, Wyoming.

Horizon: Member?, Morrison Formation.

Biostratigraphy: Zone?

Age: Age? Stage?, Malm Epoch, Late Jurassic.

Material:

KU 6285: Scapula-coracoid, tibia, ilium and a number of caudal vertebrae.

 


= Pleurocoelus montanus MARSH, 1896 (Juvenile)
Etymology: In reference to the Rocky Mountains.
= Astrodon montanus (MARSH, 1896) KUHN, 1939 (Juvenile)
= Astrodon (Pleurocelus) montanus (MARSH, 1896) KINGMAN, 2000

Holotype: YPM 1908

Locality: Reed’s YPM Quarry 1, T22N, R77W., sec. 10, Como Bluff West (west of Marshall Road), Albany County, Wyoming.

Horizon: Brushy Basin Member, Morrison Formation.

Biostratigraphy: Zone 3.

Age: Upper Part, Comobluffian age, Kimmeridgian Stage, Uppermost Malm Epoch, Late Jurassic.

Material: Complete left femur, fragmentary right femur, distal end of right humerus, anterior dorsal, dorsal centrum, another anterior dorsal centrum, and an anterior caudal centrum.


_____________________________________________________________________________________

Species: lewisi (JENSEN, 1988) MILLER, McINTOSH, STADTMAN & GILLETTE, 1992
Etymology: In honor of Mr. Arnold D. Lewis, stout companion of the trail, who patiently trained Jim Jensen in laboratory and field work.
= Cathetosaurus lewisi JENSEN, 1988)
= Cathetosaurus lewisi JENSEN, 1987 (nomen nudum)

Holotype: BYU 9047 (misprint of the specimen number as 9740, Jensen, 1988)

Locality: Dominguez-Jones Quarry, Pit 1, above confluence of Big and Little Dominguez Creeks, T14S, R98W, New Mexico Principal Meridian, Uncompahgre Plateau, Mesa County, Colorado.

Horizon: 10 m above the base of the Brushy Basin Member, Morrison Formation.

Biostratigraphy: Zone 2.

Age: Comobluffian age, Kimmeridgian Stage, Uppermost Malm Epoch, Late Jurassic.

Material: Atlas/axis, 12 cervical vertebrae with ribs, right humerus, radius, ulna, and partial manus, 12 dorsal vertebrae, 1 dorsosacral vertebra, 20 dorsal ribs, 4 sacral vertebrae with co-ossified ilium, pubis, ischia, 43 anterior caudal vertebrae and many chevrons.

Note: Ilium has teeth marks from a very large theropod.

Referred material:

TSCHOPP, WINGS, FRAUENFELDER, & ROTHSCHILED, 2016

Locality: Howe Stephens Quarry, Big Horn County, Wyoming.

Horizon: Member?, Lower Morrison Formation.

Biostratigraphy: Zone?

Age: Comobluffian age, Kimmeridgian Stage, uppermost Malm Epoch, Late Jurassic.

Material:

SMA 0002: Farily complete disarticulated skull and nearly complete articulated postcranial skeleton.

Note: With five different bone anomalies, foru of which are various kinds of overgrowths, on the manus and pes.