Genus: Carcharodontosaurus STROMER, 1931
Etymology: Carcharodon, “great White Shark” and Greek, sauros, “lizard”, Shark-toothed lizard.
= Carcharodonsaurus ROMER, 1966 (sic)
= Carchrodontosaurus WHITE, 1973 (sic)

Species: saharicus (DEPERET & SAVORNIN, 1925) STROMER, 1931
Etymology: In reference to the Sahara desert where the specimen was found.
= Megalosaurus saharicus DEPERET & SAVORNIN, 1925
= Dryptosaurus saharicus (DEPERET & SAVORNIN, 1925) DEPÉRET & SAVORNIN, 1928
= Megalosaurus africanus HUENE, 1956 (sic)

Holotype:

Locality: Timoun, Algeria.

Horizon:

Biostratigraphy:

Age: Late Aptian Stage, Middle Gallic Subepoch, Uppermost Early Cretaceous Epoch,-Cenomanian Stage, Middle Gallic Subepoch, Lowermost Gulf Epoch, Early Cretaceous.

Material: 2 teeth.
Note: Destroyed by a bombing raid on Munich by the British Royal Air Force, 1942, SERENO, 1996).

Referred material:

AMIOT, WANG, LECUYER, BUFFETAUT, BOUDAD, CAVIN, DING, FLUTEAU, KELLNER, TONG & ZHANG, 2010

Locality: Bou Laalou, Morocco.

Horizon: Kem Kem beds.

Biostratigraphy:

Age: Early Cenomanian Stage, Uppermiddle Gallic Subepoch, Lowermost Gulf Epoch, Middle Cretaceous.

Material:

M-BL-022, 023: Tooth enamel.

 

Locality: Khetitila Srhira, Morocco.

Horizon: Kem Kem beds.

Biostratigraphy:

Age: Early Cenomanian Stage, Uppermiddle Gallic Subepoch, Lowermost Gulf Epoch, Middle Cretaceous.

Material:

M-KS-013: Tooth bulk.

M-KS-023, 024, 028: Tooth enamel.

 

Locality: Chaaft, Morocco.

Horizon: Kem Kem beds.

Biostratigraphy:

Age: Early Cenomanian Stage, Uppermiddle Gallic Subepoch, Lowermost Gulf Epoch, Middle Cretaceous.

Material:

M-CH-012n, 013: Tooth enamel.

 

Locality: Takemout, Morocco.

Horizon: Kem Kem beds.

Biostratigraphy:

Age: Early Cenomanian Stage, Uppermiddle Gallic Subepoch, Lowermost Gulf Epoch, Middle Cretaceous.

Material:

M-TA-017, 019: Tooth enamel.

 

AZUMA, 2005

Locality:

Horizon:

Biostratigraphy:

Age:

Material:

FPDM-V6211: Fragmentary skull.

BENTON, BOUAZIZ, BUFFETAUT, MARTILL, OUAJA, SOUSSI & TRUEMAN, 2000/SCHLUTER & SCHWARZHANS, 1978

Locality: Tataouine region, southern Tunisia.

Horizon: Chenini Formation.

Biostratigraphy:

Age: Albian Stage, Middle Gallic Subepoch, Uppermost Early Cretaceous Epoch, Early Cretaceous.

Material:

Number: Not given: Teeth.

 

BOND & BROMLEY, 1970

Locality: Southwest of Gokwe, Zimbabwe.

Horizon: Gokwe Formation.

Biostratigraphy:

Age: Late Jurassic or Early Cretaceous.

Material:

Number: Not given: Teeth.

 

BOUAZIZ, BUFFETAUT, GHANMI, JAEGER, MARTIN, MAZIN & TONG, 1988

Locality: Locality RH 45, Foum Taahoune region, Southern Tunisia.

Horizon: Chenini Formation.

Biostratigraphy:

Age: Albian Stage, Middle Gallic Subepoch, Uppermost Early Cretaceous Epoch, Early Cretaceous.

Material:

Number: Not given: 4 teeth.

 

GOEDERT, AMIOT, BOUDAD, BUFFETAUT, FOUREL, GODEFROIT, KUSUHASHI, SUTEETHORN, TONG, WATABE, & lECUYER, 2016

Locality: Khetitila Shrira, Morocco.

Horizon: Ifezounae Formation.

Biostratigraphy:

Age: Cenomanian Stage, Uppermiddle Gallic Subepoch, Lowermost Gulf Epoch, Middle Cretaceous.

Material:

Mcb: Tooth.

 

JACOBS, WINKLER & GOMNI, 1996/GREIGERT et al. 1954, LAPPARENT, 1960

Locality: Marsa Matruth, Egypt.

Horizon: Baharija Formation.

Biostratigraphy:

Age: Cenomanian Stage, Uppermiddle Gallic Subepoch, Lowermost Gulf Epoch, Middle Cretaceous.

Material:

Number: Not given:

 

/WERNER, 1933

Locality: Northern Province, Sudan.

Horizon: Wadi Abu Hashim Member, Wadi Milk Formation.

Biostratigraphy:

Age: Cenomanian Stage, Upper Gallic subepoch, Lowest Gulf Epoch, Late Early to Early Middle Cretaceous.

Material:

Number: Not given:

 

/BUFFETAUT, 1989, 1989

Locality: Hammada du Guir, Morocco.

Horizon: Continental Red Beds.

Biostratigraphy:

Age: Albian Stage, Middle Gallic Subepoch, Uppermost Early Cretaceous Epoch,-Cenomanian Stage, Middle Gallic Subepoch, Lowermost Gulf Epoch, Early Cretaceous.

Material:

Number: Not given:

 

/LAPPARENT, 1960, MONBARON, 1978, BOUAZIZ et al, 1988

Locality: Tamenghest, Algeria.

Horizon: Continental Intercalaire.

Biostratigraphy:

Age: Albian Stage, Middle Gallic Subepoch, Uppermost Early Cretaceous Epoch, Early Cretaceous.

Material:

Number: Not given:

 

Locality: Wargla, Algeria.

Horizon: Continental Intercalaire.

Biostratigraphy:

Age: Albian Stage, Middle Gallic Subepoch, Uppermost Early Cretaceous Epoch, Early Cretaceous.

Material:

Number: Not given:

 

Locality: Adrar, Algeria.

Horizon: Continental Intercalaire.

Biostratigraphy:

Age: Albian Stage, Middle Gallic Subepoch, Uppermost Early Cretaceous Epoch, Early Cretaceous.

Material:

Number: Not given:

 

Locality: Agadez, Niger.

Horizon: Continental Intercalaire.

Biostratigraphy:

Age: Albian Stage, Middle Gallic Subepoch, Uppermost Early Cretaceous Epoch, Early Cretaceous.

Material:

Number: Not given:

Locality: Kasr-es-Souk, Morocco.

Horizon: Tegana Formation.

Biostratigraphy:

Age: Albian Stage, Middle Gallic Subepoch, Uppermost Early Cretaceous Epoch, Early Cretaceous.

Material:

Number: Not given:

 

Locality: Gharyan, Libya.

Horizon: Continental Intercalaire.

Biostratigraphy:

Age: Albian Stage, Middle Gallic Subepoch, Uppermost Early Cretaceous Epoch, Early Cretaceous.

Material:

Number: Not given:

 

Locality: Medenine, Tunsia.

Horizon: Chenini Formation.

Biostratigraphy:

Age: Aptian-Albian Stage, Middle Gallic Subepoch, Uppermost Early Cretaceous Epoch, Early Cretaceous.

Material:

Number: Not given:

 

LAPPARENT, 1951

Locality: Dahar, South Tunisia.

Horizon:

Biostratigraphy:

Age: Early Cretaceous.

Material:

Number: Not given: 9 teeth.

 

LAPPARENT, 1953

Locality: In Abangharit, 600 km south of Tamanrasset, 280 km east-north-east Algeria, In, Tamesna, Niger.

Horizon:

Biostratigraphy:

Age: Early Cretaceous.

Material:

Number: Not given: Vertebrae, 2 metatarsals, phalanges, 150 teeth.

Note: Tyrannosaurus size.

 

LAPPARENT, 1960

Locality: Timimou, Niger.

Horizon:

Biostratigraphy:

Age: Early Cretaceous.

Material:

Number: Not given: 12 teeth, caudal vertebrae.

 

Locality: Guermessa, Niger.

Horizon:

Biostratigraphy:

Age: Early Cretaceous.

Material:

Number: Not given: 5 teeth.

 

Locality: In Abangharit, Tamesna, Niger.

Horizon:

Biostratigraphy:

Age: Early Cretaceous.

Material:

Numbers: Not given: 137 teeth, caudal vertebrae.

 

 

Locality: El Rhaz, Niger.

Horizon:

Biostratigraphy:

Age: Aptian Stage, Middle Gallic Subepoch, Upper Early Cretaceous Epoch, Early Cretaceous.

Material:

Numbers: Not given: 2 teeth, fragmentary humerus, second phalange of digit II, ungual, distal portion of metatarsal , phalange of digit III?

 

Locality: Tedreft, Niger.

Horizon: Tiouraren Formation, Tegama Group.

Biostratigraphy:

Age: Aptian-Albian Stage, Middle Gallic Subepoch, Upper Early Cretaceous Epoch, Early Cretaceous.

Material:

Numbers: Not given: Fragmentary cervical vertebra, dorsal vertebrae, sacrum, 3 caudal vertebrae.

 

Locality: Aoulef, Niger.

Horizon:

Biostratigraphy:

Age: Early Cretaceous.

Material:

Numbers: Not given: Caudal vertebrae.

 

Locality: Mont Iguallala, Niger.

Horizon:

Biostratigraphy:

Age: Early Cretaceous.

Material:

Numbers: Not given: 2 caudal vertebrae.

 

 

Locality: Oued Baouet, Niger.

Horizon:

Biostratigraphy:

Age: Early Cretaceous.

Material:

Numbers: Not given: 2 caudal vertebrae.

 

Locality: 100 km SE of El Rhaz, Niger.

Horizon:

Biostratigraphy:

Age: Aptian Stage, Middle Gallic Subepoch, Upper Early Cretaceous Epoch, Early Cretaceous.

Material:

Numbers: Not given: 2 caudal vertebrae.

 

 

Locality: El Golea, Niger.

Horizon:

Biostratigraphy:

Age: Early Cretaceous.

Material:

M. Augieras personal collection : Caudal vertebrae.

 

LAVOCAT, 1954

Locality: Tabroumit, Morocco.

Horizon:

Biostratigraphy:

Age: Cenomanian Stage, Uppermiddle Gallic Subepoch, Lowermost Gulf Epoch, Middle Cretaceous.

Material:

Number: Not given: Ilium.

 

Locality: Gara Sba, Morocco.

Horizon:

Biostratigraphy:

Age: Cenomanian Stage, Uppermiddle Gallic Subepoch, Lowermost Gulf Epoch, Middle Cretaceous.

Material:

Number: Not given: Teeth.

 

RUSSELL, D. A., 1996

Locality: ‘Gres rouges infracenomaniens”, Tafilalt, Morocco.

Horizon:

Biostratigraphy:

Age: Albian Stage, Middle Gallic Subepoch, Uppermost Early Cretaceous Epoch, Early Cretaceous.

Material:

NMC 41859: Maxilla fragment.

NMC 41908, 41910, 41817, 41818, 41819: Isolated teeth.

NMC 50792: Cervical vertebra.

 

SERENO, BECK, DUTHEIL, GADO, LARSSON, LYON, MARCOT, RAUHUT, SADLEIR, SIDOR VARRICCHIO, WILSON & WILSON, 1998

Locality: Tenere Desert, Central Niger.

Horizon: Tegama Group, Elrhaz Formation.

Biostratigraphy:

Age: Aptian Stage, Middle Gallic Subepoch, Upper Early Cretaceous Epoch, Early Cretaceous.

Material:

NMC 41859: Maxillary fragment.

NMC 4198, 41910, 41817, 41818, 41819: Isolated teeth.

NMC 50792: Cervical vertebra.

 

Neotype: Ministere de l’Energie et des Mines, Rabat, Morocco, SGM-Din 1

SERENO, DUTHEIL, IAROCHENE, LARSSON, LYON, MAGWENE, SIDOR, VARRICCHIO & WILSON, 1996

Locality: Kem Kem region of southeastern Morocco.

Horizon: Kem Kem beds.

Biostratigraphy:

Age: Cenomanian Stage, Uppermiddle Gallic Subepoch, Lowermost Gulf Epoch, Middle Cretaceous.

Material: Skull missing the premaxilla and posterior end of skull. Approximately 1.6 m long.
Note: Pathology, maxilla and nasals has face-bite lesions (TANKE & CURRIE, (1998) 2000)

Ministere de l’Energie et des Mines, Rabat, Morocco, SGM-Din 3: Postaxial cervical vertebra.

AMIOT, BUFFETAUT, TONG, BOUDAD & KABIRI, 2004

M-TA-032, M-CH-003: Teeth.

 

 

STROMER, 1931

Locality: Ain Gedid am Westfuss des Gebel Harra, Egypt.
Note: Marsa Matruh, Baharije Basin, Egypt, North Africa (CHURE, 2000)

Horizon: Baharija Formation.

Biostratigraphy:

Age: ?Early Cenomanian Stage, Uppermiddle Gallic Subepoch, Lowermost Gulf Epoch, Middle Cretaceous.

Material:

Nr 1922 X 46: Teeth, left and right maxillae, nasals, cranium, fragmentary ilium, left ischium, left pubis, left femur, left fibula, dorsal vertebrae, episterpheus, chevron.

STROMER, 1934

Locality:

Horizon:

Biostratigraphy:

Age:

Material:

Nr 1912 VIII 68: Right ilium.

 

TAQUET, 1976

Locality: Kem-Kem, Morocco.

Horizon: Kem Kem Beds.

Biostratigraphy:

Age: Cenomanian Stage, Uppermiddle Gallic Subepoch, Lowermost Gulf Epoch, Middle Cretaceous.

Material:

Number: Not given: Teeth.

 

Locality: 34 Km southeast of El Rhaz, Gadoufaua, Niger.

Horizon:

Biostratigraphy:

Age: Early Cretaceous.

Material:

MNHN Td. 2250: Tooth.

 

Locality: 100 km Southeast of El Rhraz, Gadoufaua, Niger.

Horizon:

Biostratigraphy:

Age: Early Cretaceous.

Material:

MNHN Td. 2269: Teeth.

 

Locality: Gadoufaua, Niger.

Horizon:

Biostratigraphy:

Age: Aptian Stage, Middle Gallic Subepoch, Uppermost Early Cretaceous Epoch, Early Cretaceous.

Material:

Gad 8, Gad 5: Teeth.

 

Locality: Akarazeras, Niger.

Horizon:

Biostratigraphy:

Age: Early Cretaceous.

Material:

Number: Not given: Teeth.

 

____________________________________________________________________________________

Species: iguidensis BRUSATTE & SERENO, 2007
Etymology: In reference to Iguidi, Niger, where the specimen was found and Latin, ensis, "place."
= Carcharodontosaurus Nova BRUSATTE & SERENO, 2005

Holotype: MNN IGU2

Locality; Iguidi, west of In Abangharit, Agadez District, N17°56', E5°37", Niger.

Horizon: Echkar Formation, Tegama Group (Series).

Biostratigraphy:

Age: Cenomanian Stage, Upper Gallic subepoch, Lowest Gulf Epoch, Late Early to Early Middle Cretaceous.

Material: Left maxilla.

Carcharodontosaurus iguidensis (modified from BRUSATTE & SERENO, 2007), Holotype: MNN IGU2

Referred material:

MNN IGU3: Posterior skull roof and braincase including most of the frontal, parietal, supraoccipital, exoccipital-opisthotic, prootic, laterosphenoiid, orbitosphenoid, basiocciptial, basisphenoid-parasphenoid, and possibly sphenethmoid.

MNN IGU4: Mid section of a right lacrimal.

MNN IGU5: Anterior end of left dentary.

MNN IGU6: 6 through 10 teeth.

MNN IGU11: Mid cervical centrum.

Number: Not given: Teeth, and braincase.