Species: libratus LAMBE, 1914
Etymology: Latin, librare, "to balance."
= Albertosaurus libratus (LAMBE, 1914)
= Deinodon libratus (LAMBE, 1914) KUHN, 1965

Holotype: CMN (NMC) 2120

Locality: Quarry 36, 3 1/2 miles below the mouth of Berry Creek, on the south side of the Red Deer River, near prairie level, Dinosaur Provincial Park, Newell County, Alberta Province, Canada.

Horizon: Dinosaur Park Formation, upper Judith River Group.

Biostratigraphy:

Age: Judithian age, Middle Campanian Stage, middle Senonian subepoch, upper Gulf Epoch, Late Cretaceous.

Material: Almost complete skull and lower jaws, skeleton lacking cervicals and some dorsal vertebrae.
Note: Palaeopathology, bite makrs on the dentary (BROWN, CURRIE & THERRIEN, 2021), the right dorsal rib 3, gastralia 13 and 14, and left fibula have healed fractures, the left metatarsal IV is apparently diseased, with roughened exostoses on the lateral surface at midshaft and distally, the third phalanx of right pedal digit III is deformed and the claw of digit is ‘quite small and amorphous (MOLNAR, 1981)
Note: A furcula was found but BYRANT & RUSSELL disagree.

Referred material:

DALMAN & LUCAS, 2017

Locality: Dinosaur Provincial Park, Newell County, Alberta Province, Canada.

Horizon: Dinosaur Park Formation, upper Judith River Group.

Biostratigraphy:

Age: Judithian age, Middle Campanian Stage, middle Senonian subepoch, upper Gulf Epoch, Late Cretaceous.

Material:

FPDM-V8062: A complete right dentry with teeth.

YUN, 2021

SDNHM 35451: Distal metatarsal right II.

SDNHM 35452: Distal metatarsal left II.

SDNHM 35454: Distal metatarsal right III.

SDNHM 35455: Distal metatarsal right IV.

SDNHM 35453: Distal metatarsal II-1.

BROWN, CURRIE & THERRIEN, 2021

CM 12063: Right maxilla with bite marks.

CMN 57057: Right dentary of a juvenile with bite marks.

CMN 2248: Right dentary with tooth marks.

CMN 11315: Dentary tip of a juvenile with tooth marks.

ROM 436: Tip of dentary with tooth marks.

RTMP 1994.012.0602: FRagmentary skull and dentary with tooth marks.

RTMP 1998.093.0069: Fragmetn of dentary with tooth marks.

RTMP 2012.012.0082: Dentary fragment of a baby with tooth marks.

RTMP 1993.066.0001: Fragment of a left maxilla with tooth marks.

RTMP 1989.016.1782: Dentary fragment with tooth marks.

RTMP 1981.027.0087: Left dentary of a baby with tooth marks.

RTMP 2005.050.0021: Fragmentary dentary with tooth marks.RTMP 1981.010.0001: Left jubal with tooth marks.

RTMP 1996.005.007: Left denary with tooth marks.

RTMP 2010.005.0007: Left maxilla and dentary with tooth marks.

RTMP 2003.045.0084: Left dentary with tooth marks.

RTMP 2003.045.0076: Right tooth marks with tooth marks.

RTMP 1998.048.0001: Nasal with tooth marks.

RTMP 1985.062.0001: Back end of the the skull with tooth marks on the lower jugal.

RTMP 1967.009.0164: Left dentry with tooth marks.

RTMP 2017.012.0002: Right maxilla with tooth marks.

RTMP 1998.063.088: Right maxilla with tooth marks.

RTMP 200.45.0092: Left maxilla with tooth mark.

RTMP 1998.125.0001: Right maxilla with tooth marks.

RTMP 1983.036.0134: Right dentary with tooth marks.

RTMP 1996.036.0006: Right dentary with tooth marks.

RTMP 1987.025.0053: Left dentry with tooth marks.

RTMP 1994.143.0001: Fragmentary skull and dentary with tooth marks

RTMP 1982.028.0001: Left dentary with tooth marks.

RTMP 1999.055.0170: Left dentary with tooth marks.

RTMP 1995.025.0083: Left maxilla with tooth marks.

RTMP 1994.025.0006: Left maxilla with tooth marks.

RTMP 1997.012.0223: Right maxilla with tooth marks.

RTMP 2016.014.0001: Skull and dentary with tooth marks.

RTMP 1987.048.0004: Left dentary with tooth marks.

RTMP 2009.012.0014: Skull and dentary with tooth marks.

RTMP 1986.049.0029: Left dentary with tooth marks.

RTMP 1999.060.0040: Right dentary with tooth marks of a juvenile.

RTMP 2002.012.0101: Right dentary with tooth marks.

RTMP 1999.040.140: Left maxilla with tooth marks.

RTMP 1983.036.0100: Left maxilla of a juveinle with tooth marks.

RTMP 2010.121.0001: Right dentary with tooth marks.

RTMP 1996.025.0061: Left dentary with tooth marks.

RTMP 1993.036.138: Left dentary of a juvnile with tooth marks.

RTMP 2014.012.0006: Left maxilla with tooth marks.

RTMP 1998.093.012: Right surangular with tooth marks.

RTMP 1996.006.0013: Right dentary with tooth marks.

RTMP 2012.012.0029: Left dentary of a juvenile with tooth marks.

RTMP 2014.012.0040: Fragmentary dentary with tooth marks.

RTMP 1986.036.0314: Fragmentary left denary with tooth marks.

RTMP 1983.030.0001: Left Lacrimal with tooth mark.

UALVP 55605: Left maxilla with tooth marks.

UALVP 25937: Right dentary with tooth marks.

UALVP 49500: Fragmentary skull and dentary with tooth marks.

UALVP 52981: Fragmentary skull and dentary with tooth marks.

UALVP 5630: Left maxilla with tooth marks.

 

RUSSELL, 1970

Locality: 3 miles south of Steveville, Dinosaur Provincial Park, Newell County, Alberta Province, Canada.

Horizon: Dinosaur Park Formation, upper Judith River Group.

Biostratigraphy:

Age: Judithian age, Middle Campanian Stage, middle Senonian subepoch, upper Gulf Epoch, Late Cretaceous.

Material:

CMN (NMC) 8782: Fragmentary manus and pes, plus fragments of hind limbs of a juvenile.

 

Locality: Quarry unknown, Little Sandhill Creek Basin, Dinosaur Provincial Park, Newell County, Alberta Province, Canada.

Horizon: Dinosaur Park Formation, upper Judith River Group.

Biostratigraphy:

Age: Judithian age, Middle Campanian Stage, middle Senonian subepoch, upper Gulf Epoch, Late Cretaceous.

Material:

AMNH 5423: Skull and skeleton.

AMNH 5458: Skull and nearly complete skeleton.

 

Locality: Red Deer River, Alberta Province, Canada.

Horizon: Dinosaur Park Formation, upper Judith River Group.

Biostratigraphy:

Age: Judithian age, Middle Campanian Stage, middle Senonian subepoch, upper Gulf Epoch, Late Cretaceous.

Material:

FMNH PR308 (originally AMNH 5434, then sold to the Field Museum): Skull, skeleton lacking tail and hind feet.

After RUSSELL, 1970, FMNH 308

 

Locality: Near Steveville, Dinosaur Provincial Park, Newell County, Alberta Province, Canada.

Horizon: Dinosaur Park Formation, upper Judith River Group.

Biostratigraphy:

Age: Judithian age, Middle Campanian Stage, middle Senonian subepoch, upper Gulf Epoch, Late Cretaceous.

Material:

CMN (NMC) 2270: Maxilla.
Note: Juvenile/subadult.

CMN (NMC) 12063: Maxilla.
Note: Left maxilla with tooth marks (BROWN, CURRIE & THERRIEN).
Note: Juvenile/subadult.

CMN (NMC) 2250: Ilium.

CMN (NMC) 2193: Surangular.
Note: Juvenile/subadult.

CMN (NMC) 11814: Braincase.

ROM 683: Maxillae.
Note: Left maxillae with tooth marks (BROWN, CURRIE & THERRIEN).

ROM 436: Left premaxilla and maxilla.
Note: Juvenile/subadult.

RTMP 85.11.3, 86.144.1, 83.36.100: Fragmentary skulls.
Note: Juvenile/subadult.

RTMP 1983.036.0100: Maxilla Juvenile/subadult.
Note: Left maxilla with tooth marks (BROWN, CURRIE & THERRIEN).

RTMP 94.12.155: Juvenile dentary (CURRIE & DONG, 2001)

 

Locality: Quarry 029, Dinosaur Provincial Park, Newell County, Alberta Province, Canada.

Horizon: Dinosaur Park Formation, upper Judith River Group.

Biostratigraphy:

Age: Judithian age, Middle Campanian Stage, middle Senonian subepoch, upper Gulf Epoch, Late Cretaceous.

Material:

ROM 1247: Maxilla, and dentary.
Note: Juvenile/subadult with bite marks on the maxilla and left dentary and saurangular (BROWN, CURRIE & THERIEN, 2021).

 

Locality: Quarry 031, Dinosaur Provincial Park, Newell County, Alberta Province, Canada.

Horizon: Dinosaur Park Formation, upper Judith River Group.

Biostratigraphy:

Age: Judithian age, Middle Campanian Stage, middle Senonian subepoch, upper Gulf Epoch, Late Cretaceous.

Material:

ROM 1422: Fragmentary skull.
Note: Juvenile/subadult.

 

Locality: Quarry 218, Dinosaur Provincial Park, Newell County, Alberta Province, Canada.

Horizon: Dinosaur Park Formation, upper Judith River Group.

Biostratigraphy:

Age: Judithian age, Middle Campanian Stage, middle Senonian subepoch, upper Gulf Epoch, Late Cretaceous.

Material:

RTMP 94.12.155: Fragmentary skull.
Note: Juvenile/subadult.

 

Locality: East side of coulee, at Denhart, Alberta Province, Canada.

Horizon: Dinosaur Park Formation, upper Judith River Group.

Biostratigraphy:

Age: Judithian age, Middle Campanian Stage, middle Senonian subepoch, upper Gulf Epoch, Late Cretaceous.

Material:

ROM 4591: Nasals.
Note: Juvenile/subadult.

 

Locality: Quarry 21 of Sternberg 1950, Dinosaur Provincial Park, Newell County, Alberta Province, Canada.

Horizon: Dinosaur Park Formation, upper Judith River Group.

Biostratigraphy:

Age: Judithian age, Middle Campanian Stage, middle Senonian subepoch, upper Gulf Epoch, Late Cretaceous.

Material:

ROM 1237: Skeleton, lacking presacral vertebrae and right hind limb.

 

Locality: Quarry 48 of Sternberg 1950, Dinosaur Provincial Park, Newell County, Alberta Province, Canada.

Horizon: Dinosaur Park Formation, upper Judith River Group.

Biostratigraphy:

Age: Judithian age, Middle Campanian Stage, middle Senonian subepoch, upper Gulf Epoch, Late Cretaceous.

Material:

UALVP (UA) 10: Fragmentary skull, fragmentary dentary, several presacral vertebrae, ribs, right humerus and left metatarsus.
Note: Palaeopathologies,bite marks on maxillae, dentary, surangular,jugal, and angular (BROWN, CURRIE & THERRIEN, 2021), cervicals 7-8 (DISH), cervicals 5-6, right humerus, metatarsal IV, phalanx III-1 (osteomyelitits) (BELL, 2008).

 

 

Locality: Qurray 090, (Quarry 59 of Sternberg 1950), Little Sandhill Creek Basin, Dinosaur Provinical Park, Alberta Province, Canada.

Horizon: Dinosaur Park Formation, upper Judith River Group.

Biostratigraphy:

Age: Judithian age, Middle Campanian Stage, middle Senonian subepoch, upper Gulf Epoch, Late Cretaceous.

Material:

AMNH 5432: Left maxilla, 4 caudal vertebrae, fragments of pelvis, and left hind limb, left tibi-metatarsus and some phalanges of pes.
Note: Juvenile/subadult.

 

 

Locality: Quarry unknown, Dinosaur Provincial Park, Red Deer River, Newell County, Alberta Province, Canada.

Horizon: Dinosaur Park Formation, upper Judith River Group.

Biostratigraphy:

Age: Judithian age, Middle Campanian Stage, middle Senonian subepoch, upper Gulf Epoch, Late Cretaceous.

Material:

AMNH 5428: Nearly complete skeleton, except the tail, of which only three distal caudals, and the spine of the first caudals.

 

Locality: Quarry 74 (Quarry 50 of Sternberg 1950), Dinosaur Provincial Park, Newell County, Alberta Province, Canada.

Horizon: Dinosaur Park Formation, upper Judith River Group.

Biostratigraphy:

Age: Judithian age, Middle Campanian Stage, middle Senonian subepoch, upper Gulf Epoch, Late Cretaceous.

Material:

CMN (NMC) 11593: Pelvis, hind limbs and base of tail.

 

 

CURRIE, 2003

Locality: Bone bed 138 (BB 138), Dinosaur Provincal Park, approximately 50 km from Brooks, Alberta Province, Canada and Montana.

Horizon: Lowermost Oldman Formation, middle Judith River Group.

Biostratigraphy:

Age: Judithian age, Middle Campanian Stage, Senonian subepoch, Upper Gulf epoch, Late Cretaceous.

Material:

Material:

FMNH PR2211: Skeleton.

 

Locality: White Rock Coulee, 65 miles southeast of Dinosaur Provincial Park, Newell County, Alberta Province, Canada.

Horizon: Dinosaur Park Formation, upper Judith River Group.

Biostratigraphy:

Age: Judithian age, Middle Campanian Stage, middle Senonian subepoch, upper Gulf Epoch, Late Cretaceous.

Material:

RTMP 68.3.1: Hips and hind legs.

RTMP 83.30.1: Skull.

 

 

Locality: Sandy Point, 65 miles southeast of Dinosaur Provincial Park, Newell County, Alberta Province, Canada.

Horizon: Dinosaur Park Formation, upper Judith River Group.

Biostratigraphy:

Age: Judithian age, Middle Campanian Stage, middle Senonian subepoch, upper Gulf Epoch, Late Cretaceous.

Material:

RTMP 86.144.1: Fragmentary skeleton of a juvenile.

 

 

Locality: Badlands of the Red Deer River just west of Dinosaur Provincial Park, Newell County, Alberta Province, Canada.

Horizon: Dinosaur Park Formation, upper Judith River Group.

Biostratigraphy:

Age: Judithian age, Middle Campanian Stage, middle Senonian subepoch, upper Gulf Epoch, Late Cretaceous.

Material:

RTMP 73.30.1: Left hind leg, the pelvis and base of the tail.

 

 

Locality: Battle River, Dinosaur Provincial Park, Newell County, Alberta Province, Canada.

Horizon: Dinosaur Park Formation, upper Judith River Group.

Biostratigraphy:

Age: Judithian age, Middle Campanian Stage, middle Senonian subepoch, upper Gulf Epoch, Late Cretaceous.

Material:

RTMP 1991.163.0001: Skull and skeleton.
Note: With bite marks on the maxillae and dentary (BROWN, CURRIE & THERRIEN, 2021).

 

 

Locality: Quarry 238 (139?), Dinosaur Provincial Park, Newell County, Alberta Province, Canada.

Horizon: Dinosaur Park Formation, upper Judith River Group.

Biostratigraphy:

Age: Judithian age, Middle Campanian Stage, middle Senonian subepoch, upper Gulf Epoch, Late Cretaceous.

Material:

RTMP 1999.033.0001: Skull and skeskeleton.
Note: With bite marks on the maxillae and ventral side of surangular (BROWN, CURRIE & THERRIEN, 2021).

 

Locality: Quarry 219, Dinosaur Provincial Park, Newell County, Alberta Province, Canada.

Horizon: Dinosaur Park Formation, upper Judith River Group.

Biostratigraphy:

Age: Judithian age, Middle Campanian Stage, middle Senonian subepoch, upper Gulf Epoch, Late Cretaceous.

Material:

RTMP 1995.005.0001: Fragmentary skull and skeleton.
Note: With bite marks on the maxilla, dentary and nasal (BROWN, CURRIE & THERRIEN, 2021).

 

 

Locality: Quarry 234, Dinosaur Provincial Park, Newell County, Alberta Province, Canada.

Horizon: Dinosaur Park Formation, upper Judith River Group.

Biostratigraphy:

Age: Judithian age, Middle Campanian Stage, middle Senonian subepoch, upper Gulf Epoch, Late Cretaceous.

Material:

RTMP 2000.12.0011: Scattered skull.
Note: With bite marks on the maxillae and dentary (BROWN, CURRIE & THERRIEN, 2021).

 

CURRIE & RUSSELL, 2005

Locality: Quarry unknown, Dinosaur Provincial Park, Newell County, Alberta Province, Canada.

Horizon: Dinosaur Park Formation, upper Judith River Group.

Biostratigraphy:

Age: Judithian age, Middle Campanian Stage, middle Senonian subepoch, upper Gulf Epoch, Late Cretaceous.

Material:

AMNH 5358: Skull and skeleton.

 

Locality: U076, Dinosaur Provincial Park, Newell County, Alberta Province, Canada.

Horizon: Dinosaur Park Formation, upper Judith River Group.

Biostratigraphy:

Age: Judithian age, Middle Campanian Stage, middle Senonian subepoch, upper Gulf Epoch, Late Cretaceous.

Material:

Number: Not given: Fragmentary skeleton.

MAKOVICKY & CURRIE, 1998

Locality: Quarry 200, Dinosaur Provincial Park, Newell County, Alberta Province, Canada.

Horizon: Dinosaur Park Formation, upper Judith River Group.

Biostratigraphy:

Age: Judithian age, Middle Campanian Stage, middle Senonian subepoch, upper Gulf Epoch, Late Cretaceous.

Material:

RTMP 1991.036.0500: Fragmentary skeletons with furculae.
Note: Juvenile/subadult.
Note: Pathology, bite mark lesions on the skull, the shaft of the right humerus is swollen. The right fibula has a fracture, pedal phalanx II-2 is pathologic, a dislocated ungual of phalanx 4 of digit III. (TANKE & CURRIE, (1998) 2000)

Locality: Quarry 220, Dinosaur Provincial Park, Newell County, Alberta Province, Canada.

Horizon: Dinosaur Park Formation, upper Judith River Group.

Biostratigraphy:

Age: Judithian age, Middle Campanian Stage, middle Senonian subepoch, upper Gulf Epoch, Late Cretaceous.

Material:

RTMP 94.12.602: Fragmentary skeletons with furculae.
Note: Palaeopathology, midshaft of the right fibula had broken longitudinally and healed, several dorsal ribs from the middle of the sequence had well-healed fractures and a single gastralium had been broken and subsequently formed a pseudoarthrosis.

 

RYAN, RUSSELL, EBERTH & CURRIE, 2001

Locality: Centrosaurus bone bed (bone bed 43) Dinosaur Provinical Park, Alberta Province, Canada.

Horizon: Dinosaur Park Formation, upper Judith River Group.

Biostratigraphy:

Age: Judithian age, Middle Campanian Stage, middle Senonian subepoch, upper Gulf Epoch, Late Cretaceous.

Material:

Number: Not given:

 

THERRIEN, ZELENITSKY, TANAKA, VORIS, ERICKSON, CURRIE, DeBURH & KOBAYASHI, 2023

Locality: Quarry Q257, Dinosaur Provincial Park, Newell County, Alberta Province, Canada.

Horizon: Dinosaur Park Formation, upper Judith River Group, approximately 7 m below the stratigraphically lowest coal bed of teh Lethbridge Coal Zone.

Biostratigraphy: "Plateau Tuff" and 'LCZ" bentointes.

Age: Judithian age, Middle Campanian Stage, middle Senonian subepoch, upper Gulf Epoch, Late Cretaceous.

Material:

RTMP 2009.12.14: Skull and fragmentary skeleton of a juvenile.

Note: Stomach contents two caenagnathid Citipes; One femora, tibia, fibula, metatarsals and phalanges; Two tibae, fibulae, metatarsals and phalanges.

 

VORIS, ZELENITSKY, THERIEN & CURRIE, 2019

Locality: Dinosaur Provincial Park, Newell County, Alberta Province, Canada.

Horizon: Dinosaur Park Formation, upper Judith River Group.

Biostratigraphy:

Age: Judithian age, Middle Campanian Stage, middle Senonian subepoch, upper Gulf Epoch, Late Cretaceous.

Material:

RTMP 1994.143.1: Skull of a juvenile.

Note: This specimen was referred to Daspletosaurus.

 

= Laelaps incrassatus COPE, 1876 (nomen dubium)
Etymology: Latin, incrassatus, "flattened."
= Deinodon incrassatus (COPE, 1876) (nomen dubium)
= Dryptosaurus incrassatus (COPE, 1876) (nomen dubium)

Syntype: AMNH 3962

Locality: Montana.

Horizon: Judith River Formation.

Biostratigraphy:

Age: Judithian age, Campanian Stage, Senonian subepoch, Upper Gulf epoch, Late Cretaceous.

Material: 2 teeth.

Referred material:

COPE, 1876

Number: Not given: Dentary with teeth.

 

= Laelaps falculus COPE, 1876 (nomen dubium)
Etymology: Latin, falculus, diminutive of falx. falc—. "sickle."
= Deinodon falculus (COPE, 1876) (nomen dubium)
= Dromaeosaurus falculus (COPE, 1876) (nomen dubium)
= Dryptosaurus falculus (COPE, 1876) (nomen dubium)

Syntype: AMNH 3959

Locality: Montana.

Horizon: Judith River Formation.

Biostratigraphy:

Age: Judithian age, Campanian Stage, Senonian subepoch, Upper Gulf epoch, Late Cretaceous.

Material: Teeth.

 

 

= Laelaps hazenianus COPE, 1877 (nomen dubium)
Etymology: In honor of General Hazen, now in command at Fort Buford, Dakota, as a token of respect for his qualities as a man and his services in the interest of science.
= Deinodon hazenianus (COPE, 1877) (nomen dubium)
= Dryptosaurus hazenianus (COPE, 1877) (nomen dubium)

Syntype: AMNH 3957

Locality: Montana.

Horizon: Judith River Formation.

Biostratigraphy:

Age: Judithian age, Campanian Stage, Senonian subepoch, Upper Gulf epoch, Late Cretaceous.

Material: Teeth.

Note: Upper right tooth is a hadrosaur.

 

= Dryptosaurus kenabekides HAY, 1899
Etymology:

Lectotype: ANSP 9530, 9541, 9536, 9543

Locality: Missouri River, Fort Benton, Chouteau County, Montana.

Horizon: Judith River Formation.

Biostratigraphy:

Age: Judithian age, Campanian Stage, Senonian subepoch, Upper Gulf epoch, Late Cretaceous.

Material: Teeth.

 

= Gorgosaurus sternbergi MATTHEW & BROWN, 1922 (nomen nudum)
= Gorgosaurus sternbergi MATTHEW & BROWN, 1923
Etymology:In honor of C. H. Stermberg, who found the specimen.
= Albertosaurus sternbergi (MATTHEW & BROWN, 1923)
= Deinodon sternbergi (MATTHEW & BROWN, 1923) KUHN, 1965

Holotype: AMNH 5664

Locality: Quarry 006, (Quarry 54 of Sternberg 1950), Dinosaur Provincial Park, Newell County, Alberta Province, Canada.

Horizon: Dinosaur Park Formation, upper Judith River Group.

Biostratigraphy:

Age: Judithian age, Middle Campanian Stage, middle Senonian subepoch, upper Gulf Epoch, Late Cretaceous.

Material: Skull and skeleton missing caudal vertebrae beyond 24th, left radius, metacarpal I, and phalanx II-2, and left ribs.
Note: Juvenile/subadult and bite mark on right maxilla (BROWN, CURRIE & THERIEN, 2021).