Species: knopfleri SAMPSON, CARRANO & FORSTER, 2001
Etymology: In honor of Mark Knopfler, (singer/songwriter of Dire Straits) whose
music inspired the expeditions crews.
Holotype: UA 8680
Locality: MAD 93-18, near the village of Berivotra, Mahajanga Basin, Mahajana Province, northeastern Madagascar.
Horizon: Anebalemba Member, Maevarano Formation.
Biostratigraphy:
Age: Maastrichtian Stage, Uppermost Senonian Subepoch, Uppermost Gulf Epoch, Late Cretaceous.
Material: A well-preserved right dentary with several teeth.
After CARRANO, LOEWEN & SERTICH, 2011, Composite
Referred material:
CARRANO, SAMPSON & FORSTER, 2002
FMNH PR 2177, 2178, 2222, 2471, UA 9177: Left dentaries.
FMNH PR 2179: Right dentary.
UA 8682: Dentary.
FMNH PR 2124: Left splenial.
FMNH PR 3166 : Angular ( ?)
FMNH PR 2183 : Right maxilla.
FMNH PR 2165, 2180, 2226: Premaxillary/anterior dentary teeth.
FMNH PR 2164, 2181-82, 2198-2220-21, 2228: Posterior maxillary/dentary teeth.
FMNH PR 2139: Cervical vertebra 7.
FMNH PR 2140: Cervical vertebra 10.
FMNH PR 2141: Cervical vertebrae 8.
FMNH PR 2144: Dorsal vertera 4.
FMNH PR 2111, 2113-14, 2137-38, 2145, 2171, 2207, 2229, UA 8701: Dorsal vertebrae.
FMNH PR 2142: Sacral vertebra, posterior caudal centra.
FMNH PR 2133, 2204, 2230 : Proximal caudal vertebrae.
FMNH PR 2110, 2125-26, UA 8688, 8692 : Medial caudal vertebrae.
FMNH PR 2127-28, 2156-57, 2162-63, 2168, 2202-03, UA 8689-91, 8695-96, 8702-03 : Distal caudal vertebrae.
FMNH PR 3143, UA 8693-94 : Humeri.
FMNH PR 2132, 2227: Manual phalanges.
FMNH PR 2136:Right pedla phalanx II-2.
FMNH PR 2169: Unguals.
FMNH PR 2108: Left pubis.
FMNH PR 2109: Right pubis.
FMNH PR 2149-50, 2153, UA 8712: Femora.
FMNH PR 2205: Unidenfied pedal phalanx.
FMNH PR 2208, 2148, 2153, 2208, 2215, UA 8681, 8684: Right femora.
FMNH PR 2115, 2117, 2120, 2123: Left femora.
FMNH PR 2118, 2119: Proximal portion of left tibae.
FMNH PR 2112, 2121: Right tibia.
FMNH PR 2135: Left pedal ungual II/IV.
FMNH PR 2152: Distal 1/2 of a left tibia.
FMNH PR 2214: Left tibia, right metatarsal IV.
FMNH PR 2116: Rigiht ibia with partial fibulae and astragalocalcaneum.
FMNH PR 2122: Left tibia with partial fibulae and astragalocalcaneum.
FMNH PR 2235: Right calcaneum.
FMNH PR 2147, 2151, 2206: Right metatarsal II.
FMNH PR 2154: Left? metatarsal II.
FMNH PR 2146: Left? metatarsal III.
FMNH PR 2155: Left metatarsals III.
FMNH PR 2159: Right pedal phalanx III-1.
FMNH PR 2175: Distal portion of left metatarsal II.
FMNH PR 2214 : Left metatarsals IV.
FMNH PR 2234: Proximal portion of right metatarsal IV.
FMNH PR 2136, 2159, 2173, 2216-19, 2223, UA 8700, 8713-14: Pedal phalanges.
FMNH PR 2129, 2160, UA 9154: Left pedal phalanx II-1.
FMNH PR 2130: Left pedal phalanx IV-3.
FMNH PR 2131: Right pedal phalanx IV-3.
FMNH PR 2158: Left pedal phalanx IV-1.
FMNH PR 2167: Left pedal phalanx III-2.
FMNH PR 2172: Right pedal phalanx IV-1.
FMNH PR 2174: Right pedal phalanx IV-3.
FMNH PR 2134: Right pedal ungual.
FMNH PR 2135, 2236 : Unguals.
FMNH PR 2161: Right pedal phalanx II-1.
FMNH PR 2176: Left pedal phalanx III-2.
Note: Some specimens show paleopathologies (FARKE, 2004).
CARRANO, LOEWEN & SERTICH, 2011
FMNH PR 2143: Proximal 2/3rd of right humerus.
FMNH PR 2149, 2150: Proximal portion of left femur.
FMNH PR 2218: Left pedal phalanx III-1.
FMNH PR 2236: Left pedal ungal.
FMNH PR 2462: Axis.FMNH PR 2473: Right lacrimal.
FMNH PR 2463: Proximal 2/3 of a left pubis.
FMNH PR 2454: Left prearticular.
FMNH PR 2455: Left angular.
FMNH PR 2453: Left premaxilla.
FMNH PR 2456: Right postorbital.
FMNH PR 2457: Left postorbital.
FMNH PR 2459: Right metatarsal IV.
FMNH PR 2461: Proximal portion of a left pubis.
FMNH PR 2464: Cervical vertebra.
FMNH PR 2465: Cervical verterbra 5.
FMNHPR 2467: Posterior caudal vertebra.
FMNH PR 2468: Proximal 1/2 of left ischium.
FMNH PR 2469: Anterior caudal vertebra (2 elements).
FMNH PR 2470: Distal 2/3 of a left, right pubis.
FMNH PR 2472: Left? ilium.
FMNH PR 2477: Atlantal intercentrum.
FMNH PR 2478: Rib.
FMNH PR 2479: Pedal phalanx IV-3.
FMNH PR 2480: Left pedal phalanx III-2.
FMNH PR 2486: Pedal ungual II/IV.
FMNH PR 2490: Caudal centra.
FMNH PR 2491: Cervical 10.
FMNH PR 2492: Posterior caudal vertebra (3 elements).
FMNH PR 2493: Unidentified pedal phalanx.
FMNH PR 2495: Laterosphenoid?
UA 8710: Proixmal portion of right tibia.
UA 8711: Tibiae.
UA 8685: Left tibia.
UA 8686: Right pedal phalanx IV-4.
UA 8687: Right tibia.
UA 8689, 8690, 8691, 8695, 8696, 9139 9119, FMNH PR 2202, 2203: Posterior caudal.
UA 8692: Middle caudal centra.
UA 8693: Right humeral shaft.
UA 8694: Humeral shaft?
UA 8711: Right tibia.
UA 8712: Right femur.
UA 8702: Caudal centra.
UA 9090: Right postorbital.
UA 9095: Left frontal.
UA 9096: Right pedal phalanx IV-2.
UA 9098: Sacral vertebra 5-6, proximal 2/3 portion of a right fibula.
UA 9099: Proximal portion of right tibia.
UA 9100: Proximal 2/3 of a right pubis.
UA 9101: Left metatarsal IV.
UA 9102: Left metatarsal III and distal tarsal 3.
UA 9104: Proxial portion of left cervical rib 9.
UA 9106: Cervical vertebra 4.
UA 9107: Dorsal vertebra arch 4.
UA 9109: Unidentified vertebra.
UA 9111: Cervical vertbra 7.
UA 9112: Anterior caudal vertebra.
UA 9110, 9113, 9151: Caudal centrum.
UA 9115: Dorsal centra.
UA 9117: Pubis shaft?
UA 9118: Distal portion of left metatarsal I?
UA 9120, 9158: Middle/posterior caudal vertebra with rib.
UA 9121: Cervical vertebra 3.
UA 9122: Left metatarsal II.
UA 9127, 9140, 9148: Caudal centra.
UA 9130: Axis arch.
UA 9132: Proximal 1/2 of right fibula and unidentified pedal phalanx.
UA 9133: Proximal portion of a right ischium.
UA 9134: Proximal portion of a right fibula.
UA 9135: Left distal shaft of a femur.
UA 9136: Left scapula, shaft of a pubis.
UA 9142: Distal portion of left tibia.
UA 9143: Proiximal 1/2 of a right fibula.
UA 9144: Right pedal ungual.
UA 9146: Proximal portion of metacarpal IV?
UA 9149: Left prearticular.
UA 9150: Cervical vertebrae 9-10.
UA 9152: Middle chevron.
UA 9153: Proximal portion of left pubis.
UA 9155: Proximal portion of a dorsal rib.
UA 9156: Left pedal phalanx IV-1.
UA 9157: Left pedal ungual III-4.
UA 9159: Right coracoid.
UA 9160: Proximal portion of right dorsal rib 4, right scapulocoracoid.
UA 9162: Proximal part of a right pubis.
UA 9163: Distal portion of right metatarsal IV.
UA 9166: Right prearticular, articular.
ua 9168: Left ischium,
UA 9170: Right central and posterior porition of a right ilium, right femur.
UA 9171: Cervical vertebra 8.
UA 9172: Right ischium, right proximal 2/3 fibula.
UA 9173: Anterior caudal centrum.
UA 9174: Caudal arch.
UA 9103, 75, 76 9092, 1458, 9092, 9097: Posterior dorsal verebrae.
UA 9164: Dorsal arch.
UA 9165: Right proximal portion of humerus.
UA 9167: Right fibula.
UA 9169: Right proximal cervical rib 6.
UA 9178: Basioccipital.
UA 9190, 9182: Chevrons.
UA 9187: Caudal centra.
UA 9193: Proximal portion of left femur.
UA 9196: Chevron.
UA 9198: Left pedal ungual.
UA 8688, 9179, 9197: Middle caudal centra.
UA 9181: Cervical vertebra 8.
UA 9091, 9123, 9126, 9128: Posterior teeth.
UA 9094: Right pedal phalanx III-4.
UA 9774: Distal 1/4 of metatarsal II.
UA 9858: Middle/posterior caudal with rib.
UA 9859: Cervical centrum.
UA 9861, 9863: Proximal portions of ribs.
UA 9862: Proxial portion of dorsal rib 4.
CARRANO, SAMPSON & LOWEN, 2004
Number: Not given: Postorbital, frontal, qudrate, partial braincase, axis and intercentrum, additional cervicals and dorsals, ischia, fibulae, phalanges and unguals.
CARRANO, LOEWEN & SERTICH, 2011
Locality: "East of Mammal", near the village of Berivotra, Mahajanga Basin, Mahajana Province, northeastern Madagascar.
Horizon: Anebalemba Member, Maevarano Formation.
Biostratigraphy:
Age: Maastrichtian Stage, Uppermost Senonian Subepoch, Uppermost Gulf Epoch, Late Cretaceous.
Material:
UA 9145: Right dentary.
UA 9138: Distal 1/3 of left metatarsal II.
Locality: MAD 03-03, near the village of Berivotra, Mahajanga Basin, Mahajana Province, northeastern Madagascar.
Horizon: Anebalemba Member, Maevarano Formation.
Biostratigraphy:
Age: Maastrichtian Stage, Uppermost Senonian Subepoch, Uppermost Gulf Epoch, Late Cretaceous.
Material:
FMNH PR 2457: Braincase.
Locality: MAD 03-40, near the village of Berivotra, Mahajanga Basin, Mahajana Province, northeastern Madagascar.
Horizon: Anebalemba Member, Maevarano Formation.
Biostratigraphy:
Age: Maastrichtian Stage, Uppermost Senonian Subepoch, Uppermost Gulf Epoch, Late Cretaceous.
Material:
FMNH PR 2466: Axis.
Locality: MAD 03-66, near the village of Berivotra, Mahajanga Basin, Mahajana Province, northeastern Madagascar.
Horizon: Anebalemba Member, Maevarano Formation.
Biostratigraphy:
Age: Maastrichtian Stage, Uppermost Senonian Subepoch, Uppermost Gulf Epoch, Late Cretaceous.
Material:
FMNH PR 2460: Sacral vertebra 3-6.
Locality: MAD 05-04, near the village of Berivotra, Mahajanga Basin, Mahajana Province, northeastern Madagascar.
Horizon: Anebalemba Member, Maevarano Formation.
Biostratigraphy:
Age: Maastrichtian Stage, Uppermost Senonian Subepoch, Uppermost Gulf Epoch, Late Cretaceous.
Material:
UA 9190: Middle caudal centra.
Locality: MAD 05-42, near the village of Berivotra, Mahajanga Basin, Mahajana Province, northeastern Madagascar.
Horizon: Anebalemba Member, Maevarano Formation.
Biostratigraphy:
Age: Maastrichtian Stage, Uppermost Senonian Subepoch, Uppermost Gulf Epoch, Late Cretaceous.
Material:
FMNH PR 2458: Arch, posterior caudal centra, left scapula.
FMNH PR 2481: Cervical vertebrae 6, 7, cervical vertebra, Dorsal vertebrae 5-7, 11?, 12?, 13? 14?, dorsal centra 1-6, sacral centrua 1-6, anterior caudal vertebrae, anterior/middle caudal centrum, middle caudal centrum, middle/posterior centrum with rib, posterior caudal centra caudal centra, right dorsal rib, rib, fragment rib, anterior gastrale, chevron, left and right scapulae, left? coracoid, fragment of coracoid, right humerus, left and right ilia, left and right pubis, right femur, left tibia, right tibia, proximal2/3 of left fibula, right fibula and astragalus, left astragalus, distal 1/2 of left metatarsal II, right metatarsal II, left pedal phalanx III-1, and right pedal phalanx III-x.
FMNH PR 2482: Anterior/middle caudal centrum, posterior centrum.
FMNH PR 2484, 2458, 2610: Dorsal vertebrae.
FMNH PR 2485: Cervical 4, 2 cervical vertebrae, dorsal vertebra 2, 4, sacral centra 2, middle caudal centrum, posterior centrum, caudal centra, right cervical rib 4, cervical rib 6, proximal portion of right cervical rib 7, proximal portion of left dorsal rib, dorsal rib 4, left dorsal rib, fragmentary rib, chefron, left humerus, left ilium, right femur, tibia, fibula, left tibia, fibula, right metatarsal III, distal portion of left metatarsal IV, left pedal ungual II/IV and unidentified pedal phalanx.
FMNH PR 2496: Right frontal.
FMNH PR 2602, 2632, 2685, : Fragment of rib.
FMNH PR 2604: Left pedal phalanx II-2.
FMNH PR 2605: Right pedal phalanx IV-1.
FMNH PR 2606: Right scapula.
FMNH PR 2611: Unidentified pedal phalanx.
FMNH PR 2613: Right pedal phalanx III-x.
FMNH PR 1614: Right pedal phalanx IV-1.
FMNH PR 2616: Right pedal phalanx II-1.
FMNHPR 2617: Right pedal phlanx III-x.
FMNH PR 2630: Cervical vertebra.
FMNH PR 2634: Proximal portion of rib.
FMNH PR 2636: Dorsal vertebra 6.
FMNH PR 2667: Left pedal ungual II/IV.
FMNH PR 2681: Unidentified vertebra arch.
FMNH PR 2687: Right proximal portion of metatarsal III and distal tarsal 3.
FMNH PR 2672: Cervical right rib 10/dorsal 1.
FMNH PR 2673: Chefron.
FMNH PR 2677: Unidentified centra.
FMNH PR 2686: Left tibia,
FMNH PR 2699, 2700: Caudal centra.
FMNH PR 2163: Posterior caudal centra.
FMNH PR 2709: Right pedal phalanx II-1.
FMNH PR 2817: Left tibia.
FMNH PR 2818: Anterior tooth.
FMNH PR 2837: Dorsal centrum.
FMNH PR 2696: Posterior tooth.
FMNH PR 2481: Ceratobranchial, atlantal intercenturm, cervical vertebra 9.
Locality: MAD 07-16, near the village of Berivotra, Mahajanga Basin, Mahajana Province, northeastern Madagascar.
Horizon: Anebalemba Member, Maevarano Formation.
Biostratigraphy:
Age: Maastrichtian Stage, Uppermost Senonian Subepoch, Uppermost Gulf Epoch, Late Cretaceous.
Material:
UA 9189: Cervical 4.
Locality: MAD 07-17, near the village of Berivotra, Mahajanga Basin, Mahajana Province, northeastern Madagascar.
Horizon: Anebalemba Member, Maevarano Formation.
Biostratigraphy:
Age: Maastrichtian Stage, Uppermost Senonian Subepoch, Uppermost Gulf Epoch, Late Cretaceous.
Material:
UA 9184: Caudal vertebra.
Locality: MAD 93-01, near the village of Berivotra, Mahajanga Basin, Mahajana Province, northeastern Madagascar.
Horizon: Anebalemba Member, Maevarano Formation.
Biostratigraphy:
Age: Maastrichtian Stage, Uppermost Senonian Subepoch, Uppermost Gulf Epoch, Late Cretaceous.
Material:
FMNH PR 2201: Posterior tooth.
Locality: MAD 93-09, near the village of Berivotra, Mahajanga Basin, Mahajana Province, northeastern Madagascar.
Horizon: Anebalemba Member, Maevarano Formation.
Biostratigraphy:
Age: Maastrichtian Stage, Uppermost Senonian Subepoch, Uppermost Gulf Epoch, Late Cretaceous.
Material:
FMNH PR 2181: Posterior tooth (two elements).
Locality: MAD 93-26, near the village of Berivotra, Mahajanga Basin, Mahajana Province, northeastern Madagascar.
Horizon: Anebalemba Member, Maevarano Formation.
Biostratigraphy:
Age: Maastrichtian Stage, Uppermost Senonian Subepoch, Uppermost Gulf Epoch, Late Cretaceous.
Material:
UA 9192: Left pedal phalanx III-x.
Locality: MAD 93-30, near the village of Berivotra, Mahajanga Basin, Mahajana Province, northeastern Madagascar.
Horizon: Anebalemba Member, Maevarano Formation.
Biostratigraphy:
Age: Maastrichtian Stage, Uppermost Senonian Subepoch, Uppermost Gulf Epoch, Late Cretaceous.
Material:
FMNH PR 2133: Anterior caudal vertebra.
FMNH PR 2217: Left pedal phalanx II-2.
UA 8683: Distal portion of right metatarsal II.
Locality: MAD 93-34, near the village of Berivotra, Mahajanga Basin, Mahajana Province, northeastern Madagascar.
Horizon: Anebalemba Member, Maevarano Formation.
Biostratigraphy:
Age: Maastrichtian Stage, Uppermost Senonian Subepoch, Uppermost Gulf Epoch, Late Cretaceous.
Material:
UA 9188: Right pedal phalanx II-1.
UA 9186: Right pedal phalanx III-x.
Locality: MAD 93-35, near the village of Berivotra, Mahajanga Basin, Mahajana Province, northeastern Madagascar.
Horizon: Anebalemba Member, Maevarano Formation.
Biostratigraphy:
Age: Maastrichtian Stage, Uppermost Senonian Subepoch, Uppermost Gulf Epoch, Late Cretaceous.
Material:
FMNH PR 2220: Antrior tooth.
FMNH PR 2199, 9183, 9734: Posteior teeth.
UA 9137: Left pedal phalanx IV-2/3.
UA 9185: Right pedal phalanx VI-1.
UA 9199: Left pedal phalanx III-2/3.
UA 9773: Distal portion of undentified metatarsal.
Locality: MAD 93-38, near the village of Berivotra, Mahajanga Basin, Mahajana Province, northeastern Madagascar.
Horizon: Anebalemba Member, Maevarano Formation.
Biostratigraphy:
Age: Maastrichtian Stage, Uppermost Senonian Subepoch, Uppermost Gulf Epoch, Late Cretaceous.
Material:
FMNH PR 2223: Unidentifed pedal phalanx (2 elements).
Locality: MAD 93-72, near the village of Berivotra, Mahajanga Basin, Mahajana Province, northeastern Madagascar.
Horizon: Anebalemba Member, Maevarano Formation.
Biostratigraphy:
Age: Maastrichtian Stage, Uppermost Senonian Subepoch, Uppermost Gulf Epoch, Late Cretaceous.
Material:
FMNH PR 2679: Distal 1/2 left metatarsal II.
Locality: MAD 93-74, near the village of Berivotra, Mahajanga Basin, Mahajana Province, northeastern Madagascar.
Horizon: Anebalemba Member, Maevarano Formation.
Biostratigraphy:
Age: Maastrichtian Stage, Uppermost Senonian Subepoch, Uppermost Gulf Epoch, Late Cretaceous.
Material:
UA 8682: Unidentified pedal phalanx.
Locality: MAD 93-101, near the village of Berivotra, Mahajanga Basin, Mahajana Province, northeastern Madagascar.
Horizon: Anebalemba Member, Maevarano Formation.
Biostratigraphy:
Age: Maastrichtian Stage, Uppermost Senonian Subepoch, Uppermost Gulf Epoch, Late Cretaceous.
Material:
FMNH PR 2224: Manual phalanx.
Locality: MAD 95-14, near the village of Berivotra, Mahajanga Basin, Mahajana Province, northeastern Madagascar.
Horizon: Anebalemba Member, Maevarano Formation.
Biostratigraphy:
Age: Maastrichtian Stage, Uppermost Senonian Subepoch, Uppermost Gulf Epoch, Late Cretaceous.
Material:
UA 9194: Manual phalanx.
Locality: MAD 96-01, near the village of Berivotra, Mahajanga Basin, Mahajana Province, northeastern Madagascar.
Horizon: Anebalemba Member, Maevarano Formation.
Biostratigraphy:
Age: Maastrichtian Stage, Uppermost Senonian Subepoch, Uppermost Gulf Epoch, Late Cretaceous.
Material:
FMNH PR 2216: Left pedal phalanx IV-2.
UA 9195: Middle/posterior caudal vertebra with rib.
Locality: MAD 96-41, near the village of Berivotra, Mahajanga Basin, Mahajana Province, northeastern Madagascar.
Horizon: Anebalemba Member, Maevarano Formation.
Biostratigraphy:
Age: Maastrichtian Stage, Uppermost Senonian Subepoch, Uppermost Gulf Epoch, Late Cretaceous.
Material:
FMNH PR 2225: Manual phalanx.
Locality: MAD 98-26, near the village of Berivotra, Mahajanga Basin, Mahajana Province, northeastern Madagascar.
Horizon: Anebalemba Member, Maevarano Formation.
Biostratigraphy:
Age: Maastrichtian Stage, Uppermost Senonian Subepoch, Uppermost Gulf Epoch, Late Cretaceous.
Material:
FMNH PR 2221: Posterior tooth.
Locality: MAD 99-05, near the village of Berivotra, Mahajanga Basin, Mahajana Province, northeastern Madagascar.
Horizon: Anebalemba Member, Maevarano Formation.
Biostratigraphy:
Age: Maastrichtian Stage, Uppermost Senonian Subepoch, Uppermost Gulf Epoch, Late Cretaceous.
Material:
FMNH PR 2169: Manual ungual.
Locality: MAD 99-06, near the village of Berivotra, Mahajanga Basin, Mahajana Province, northeastern Madagascar.
Horizon: Anebalemba Member, Maevarano Formation.
Biostratigraphy:
Age: Maastrichtian Stage, Uppermost Senonian Subepoch, Uppermost Gulf Epoch, Late Cretaceous.
Material:
FMNH PR 2170: Posterior tooth.
Locality: MAD 99-30, near the village of Berivotra, Mahajanga Basin, Mahajana Province, northeastern Madagascar.
Horizon: Anebalemba Member, Maevarano Formation.
Biostratigraphy:
Age: Maastrichtian Stage, Uppermost Senonian Subepoch, Uppermost Gulf Epoch, Late Cretaceous.
Material:
FMNHPR 2173: Right pedal phalanx III-2.
Locality: MAD 99-33, near the village of Berivotra, Mahajanga Basin, Mahajana Province, northeastern Madagascar.
Horizon: Anebalemba Member, Maevarano Formation.
Biostratigraphy:
Age: Maastrichtian Stage, Uppermost Senonian Subepoch, Uppermost Gulf Epoch, Late Cretaceous.
Material:
FMNHPR 2218: Right pedal phalanx -2/3.
UA 9105: Unidentified metatarsal and rib fragment.
Locality: Not recorded, near the village of Berivotra, Mahajanga Basin, Mahajana Province, northeastern Madagascar.
Horizon: Anebalemba Member, Maevarano Formation.
Biostratigraphy:
Age: Maastrichtian Stage, Uppermost Senonian Subepoch, Uppermost Gulf Epoch, Late Cretaceous.
Material:
UA 8700: Right pedal phalanx III-1.
UA 9613: Distal portion of left tibia.
FANTI & THERRIEN, 2007
Locality: Berivotra, Mahajanga Basin, Mahajana Province, northeastern Madagascar.
Horizon: Anebalemba Member, Maevarano Formation.
Biostratigraphy:
Age: Maastrichtian Stage, Uppermost Senonian Subepoch, Uppermost Gulf Epoch, Late Cretaceous.
Material:
Morphotype 2
MSNM V5378: Lateral tooth.