Species: bucklandii RITGEN, 1826
Etymology: In honor of W. Buckland.
= Megalosaurus bucklandi MANTELL, 1827 (sic)

Lectotype: OUM J13505

Locality: Stonesfield near Woodstock, about 12 miles to the north west of Oxford, (SP 387171), Oxfordshire County, England, Southern United Kingdom.

Horizon: Unknown Horizon, Taynton Limestone Formation (old Stonesfield Member, Sharps Hill Formation).

Biostratigraphy: Procerites progracilis ammonite zone.

Age: Middle Bathonian Stage, Middle Dogger Epoch, Middle Jurassic.

Material: Fragmentary mandibular ramus with teeth.

Referred material:

Skull imperfectly known, with very large orbital fenestrae containing sclerotic plates, 4 premaxillary teeth, 12 possibly more maxillary teeth, 11 or 12 cervical vertebrae, femur.

OUM J12142 (= OUM J.13505): Dentary.

OUM J13561: Right femur.

OUM J13565: Left ischium.

OUM J.13577: Dorsal vertebra.

OUM J. 13576: Sacrum.

OUM J.13578: Proximal caudal.

OUM J. 13579: Caudal vertebra.
Note: Paleopathology, cervical rib shows an unusual swelling at the base of the capitular process that appears to be a healed fracture (TANKE & ROTHSCHILD, 1999).

OUM J. 13581, J. 13585: Ribs.

OUM J.29881: Ilium.

OUM J.13563: Right pubis.

OUM J.13565: Ischium.

OUM J.13567: Left pubis.

OUM J.13572: Metatarsal.

OUM J.13574: Right scapula and coracoid.

OUM J.13575: Left humerus.

OUM J.13506, J.13559: 2 maxilla.

OUM J13568: Left tibia.

OUM J13572: Left metatarsal II.

OUM J13573: Left metatarsal II.

OUM J13569: Right metatarsal III.

OUM J29753: Proximal right femur.

OUM J29754: Proximal right femur.

OUM J29758: Distal right tibia.

OUM J29761: Left humerus.

OUM J29765: Right scapula.

OUM J29797: Crushed pubis.

OUM J29872: Left ischium.

OUM J29881: Right ilium.

OUM J29882: Right ilium.

OUM J29883: Right ilium.

OUM J29884: Left ilium.

OUM J.29889: Fragmentary right scapula.

OUM J29813: Articular region of a small right mandible.

OUM J12142: Partial mandibular ramus with teeth.

BENSON, 2009

NHM R1099, CAMSM Woodwardian Collection D.11.35c, OUMNH J13574, J.29891, J.2988: Scapulocoracoids.

SPAMER, DAESCHLER & VOSTREYS-SHAPIRO,1995

ANSP 9910: Cast of original.

WALKER, 1964

OUM J.13574: Scapulocoracoid.

OUM J 13560: Right ilium.

Top after Galton & Jensen, 1979, Bottom after Day & Barrett, 2004.

 

DAY & BARRETT, 2004

Morph A

OUM J29802: Right femur.

BMNH 31804: Left femur.

BMNH 31806: Femur.

SDM 44.24: Femur.

MNHN 9630: Left femur.

 

Morph B

OUM J.13561: Femur.

OUM J29803: Right femur.

BMNH 31808: Left femur.

SDM 44.23: Femur.

Composite skeleton.

BENSON, 2009

Localtiy: Uknown, England, Southern United Kingdom

Horizon:

Biostratigraphy:

Age: Middle Jurassic.

Material:

NHM R9303: Maxilla.

NHM R8304, R9305: 2 dentaries.

= Metriacanthosaurus reynoldsi WELLES, POWELL & PICKERING, 1995 (as per BENSON, 2010)

Locality: New Park Quarry, Longborough, near Stow-on-the-Wold, (SP 176282), Gloucestershire County, England, Southern United Kingdom

Horizon: Chipping Norton Member, Chipping Norton Formation.

Biostratigraphy: Zigzagiceras zigzag zone.

Age: Lower Bathonian Stage, Middle Dogger Epoch, Middle Jurassic.

Material:

OUM J29800: Right scapula.

SDM 44.1: Maxilla.

SDM 44.23, 44.24: Femora.

BENTON & SPENCER, 1995

SDM 4-10, 12-5, 18, 21, 23-6:

NHMUK (old BMNH) R9666-76, R.9678-9, R.9681-7, R.9689-700:

LOMAX & TAMURA, 2014

NHMUK OR8305: Left dentary fragment.

STGCM 1944.5: Middle caudal vertebra.

STGCM 1944.7: Proximal caudal vertebra.

STGCM 1944.23: Rifht femur.

= Megalosaurus bucklandi CLARK & BLOWS, 2023

STGCM 1944.1: Fragmentary left maxilla with teeth.

STGGM 1944.10: Dorsal vertebra.

STGCM 1944.15: Right coracoid.

NHMUK (old BMNH) R9674: Dorsal vertebra.

NHMUK (old BMNH) R8172: Fragmentary dorsal vertebra.

NHMUK (old BMNH) R9676: Mid caudal vertebrta.

STGCM 9144.23, 24: Right femora.

 

 

BENTON & SPENCER, 1995

Locality: Shipton-on-Cherwell Quarry, North-West Corner, (SP 475178), Oxfordshire County, England, Southern United Kingdom.

Horizon:

Biostratigraphy:

Age: Upper Bathonian Stage, Dogger Epoch, Middle Jurassic.

Material:

OUM J.13598, J.29773, J.13882, J.29765: Scapula and isolated teeth.

 

CLARK & BLOWS, 2023

Locality: Oakham Quarry, Gloucestershire County, England, Southern United Kingdom

Horizon: Chipping Norton Member, Chipping Norton Formation.

Biostratigraphy: Zigzagiceras zigzag zone.

Age: Lower Bathonian Stage, Middle Dogger Epoch, Middle Jurassic.

Material:

STGCM 1944.2, 3: Teeth.

STGCM 1944.19: Fragment of right ilium.

NHMYUK R9679:Fragment of sacrum.

 

HUENE, 1926/LYDEKKER, 1888

Locality?

Horizon:

Biostratigraphy:

Age:

Material:

BMNH 2301: Cast of a portion of the right ramus and mandible.

BMNH 47963, 31834, 42024, R234, 41305 & Oxford Mus: A number of single found teeth.

BMNH 31813: 2 dorsal vertebrae

BMNH R285: Fragmentary small cervical vertebra.

BMNH 47169: Dorsal centrum.

BMNH 28957: 2 sacrals

BMNH R700: 4 anterior caudals, of a juvenile.

BMNH 36585: Left ulna, 1 sacral rib.

BMNH 2581: 1 incomplete caudal vertebra.

BMNH 378303, 44097, 44097a, 31824-5: 5 single ribs.

BMNH R1099: Left scapula and coracoid.

BMNH 31810: Right coracoid.

BMNH 40131: 1 damaged left coracoid.

BMNH R283: Fragmentary right ilium.

BMNH: 1 ulna.

BMNH R1100, R1101: 2 right ilia.

BMNH R1100: Right ilium, after Day & Barrett, 2004.

BMNH 31811: Left ilium, after Day & Barrett, 2004.

BMNH 25582: Ischium.

BMNH 28301: Fragmentary ischium.

BMNH 31809: Proximal end of a left tibia.

BMNH R1103: 1 damaged right tibia.

BMNH R1102: Left tibia.

BMNH R1104: Medial metatarsal.

BMNH 31932: 1 phalanx.

BMNH 40125a: An imperfect lateral metatarsal.
Note: Probably from Stonesfield although labeled Purbeck.

BMNH: Some more bones.

WELLES & PICKERING, 1999

BMNH 31828: Right jugal.

Marlborough Collection; Partial right mandible.
Note: Now lost.

BMNH R1098, 28957: Fragmentary sacrum.

BMNH 25581: Proximal caudal.

BMNH 25582: Proximal left pubis.

BMNH 31810: Right coracoid.

BMNH 31932: Pedal phalanx.

J. Phillips Collection, now lost: pedal ungual.

BMNH uncatalouged: Left tibia.

GSM IGS 3887: Sacrum.

RCS 72: Lost right femur.

RCS 73: Lost femur shaft.

RCS 74: Lost distal tibia.

GPIT 18392: Fibula.

PHILLIPS, 1871

Oxford Museum: A great number of single bones, amongst them: jaws, cervical and other vertebrae, haemapophyhses, scapula-coracoid, humerus, all parts of the pelvis, femur, the 3 metatarsals belonging together.

 

 

/ LYDEKKER, 1888

Locality: Huntsman’s Quarry, Naunton, (SP 126253), Gloucestershire County, England, Southern United Kingdom.

Horizon: ‘Cotswold slates’, Eyeford Member.

Biostratigraphy:

Age: Middle Bathonian Stage, Dogger Epoch, Middle Jurassic.

Material:

BMNH 28608: Tooth.

BENTON & SPENCER, 1995

BMNH R2635: Tooth.

GLCRM T.70-1, G.72-3, T.74-6: Limb bones and ribs.

 

/HUXLEY, 1869

Locality: Locality not mentioned, perhaps near Stonesfield or Sherbourne, Dorset County, England, Southern United Kingdom.

Horizon: Probably Stonesfield Slates.

Biostratigraphy:

Age: Bathonian Stage, Dogger Epoch, Middle Jurassic.

Material:

Number: Not given: Left maxilla.

 

BUFFETAUT, CUNY & Le LEOUFF, 1991/MERCIER, 1937

Locality: Les Aucrais, Calvados Department, Normandy, France.

Horizon:

Biostratigraphy:

Age: Lower Bathonian Stage, Dogger Epoch, Middle Jurassic.

Material:

Number: Not given: Tooth.

 

DELAIR, 1973

Locality: Stanton, Wiltshire County, England, Southern United Kingdom.

Horizon: Forest Marble.

Biostratigraphy:

Age: Upper Bathonian Stage, Dogger Epoch, Middle Jurassic.

Material:

BMNH 39476: Tooth.

 

Locality: Swindon, Wiltshire County, England, Southern United Kingdom.

Horizon: Kimmeridge Clay.

Biostratigraphy:

Age: Kimmeridgian Stage, Middle Malm Epoch, Late Jurassic.

Material:

Number: Not given: Unspecified remains.

 

FREEMAN, 1979

Locality: Old Cement Works Quarry, (Washford Quarry), Kirtlington, Oxfordshire County, England, Southern United Kingdom. Ordnance Survey Grid Reference SP 495200.

Horizon: Kirtlington Mammal bed, near base of the Forest Marble.

Biostratigraphy: Oppelia aspidioides Zone.

Age: Upper Bathonian Stage, upper middle Dogger Epoch, Middle Jurassic.

Material:

Number: Not given: Teeth.

Note: Maybe belongs to a coelurosaur.

 

GALTON & MOLNAR, 2005

Locality: Stonesfield Slate, Oxfordshire County, England, Southern United Kingdom.

Horizon:

Age: Upper Bathonian Stage, upper middle Dogger Epoch, Middle Jurassic.

Material:

OUMNH J13562: Distal end of a left tibia.

 

HUENE, 1906

Locality: Stonesfield, (SP 387171), Oxfordshire County, England, Southern United Kingdom.

Horizon: Stonesfield Member, Sharps Hill Formation.

Biostratigraphy:

Age: Middle Bathonian Stage, Middle Dogger Epoch, Middle Jurassic.

Material:

Number: Not given: Fragment of posterior portion of a skull.

 

HUENE, 1966

Locality: Stonesfield, (SP 387171), Oxfordshire County, England, Southern United Kingdom.

Horizon: Stonesfield Member, Sharps Hill Formation.

Biostratigraphy:

Age: Middle Bathonian Stage, Middle Dogger Epoch, Middle Jurassic.

Material:

Number: Not given: Dorsal vertebra.

 

LYDEKKER, 1888

Locality: Daston, Northamptonshire County, England, Southern United Kingdom.

Horizon: Inferior Oolite.

Biostratigraphy:

Age: Bajocian Stage, Dogger Epoch, Early Jurassic.

Material:

BMNH 47152: A small tooth.

 

 

Locality: Selsly Hill, Gloucestershire County, England, Southern United Kingdom.

Horizon: Inferior Oolite.

Biostratigraphy:

Age: Bajocian Stage, Dogger Epoch, Early Jurassic.

Material:

BMNH R497: Middle part of a crown.

 

Locality: Unknown, Cornbrash, England, Southern United Kingdom.

Horizon: Cornbrash Formation.

Biostratigraphy:

Age: Late Bathonian Stage, Dogger Epoch, Early Jurassic.

Material:

BMNH 47169: A centrum of a trunk vertebra.

 

Locality: Neocomian Bone-bed of Potton, Bedfordshire County, England, Southern United Kingdom.

Horizon:

Biostratigraphy:

Age: Early Cretaceous.

Material:

BMNH 42028: Fragmentary lumbar? vertebra.

 

Locality: Coral Rag of Dry Sandford, Oxfordshire County, England, Southern United Kingdom.

Horizon:

Biostratigraphy:

Age: Early Jurassic.

Material:

BMNH R1027: Cast of the sacrum.

 

Locality: Sarsgrove, Sarsden, chipping Norton, Wiltshire County, England, Southern United Kingdom.

Horizon: Great Oolite.

Biostratigraphy:

Age: Early Jurassic.

Material:

BMNH R413: Median metatarsal.

 

OWEN, 1854

Locality: Stammerham, West Sussex County, England, Southern United Kingdom.

Horizon: Wealden Formation.

Biostratigraphy:

Age: Early Cretaceous.

Material:

Royal Collage of Surgeons: A colored plaster cast of a posterior dorsal vertebra.

 

Locality: Near Battle, Sussex County, England, Southern United Kingdom.

Horizon: Wealden Formation.

Biostratigraphy:

Age: Early Cretaceous.

Material:

Royal Collage of Surgeons: A dorsal centrum, caudal, posterior caudal, of a juvenile.

 

Locality: In the Neighborhood of Battle, Sussex County, England, Southern United Kingdom.

Horizon: Wealden Formation.

Biostratigraphy:

Age: Early Cretaceous.

Material:

Royal Collage of Surgeons: A portion of a neuropophysis with one of the zygapophyses.

 

Locality: Stonesfield, (SP 387171), Oxfordshire County, England, Southern United Kingdom.

Horizon: Stonesfield Member, Sharps Hill Formation.

Biostratigraphy:

Age: Middle Bathonian Stage, Middle Dogger Epoch, Middle Jurassic.

Material:

Royal Collage of Surgeons: Right femur, shaft of a femur, left tibia, cast of a middle phalanx, a block with a portion of a long slightly compressed bone, a block containing a crown of a tooth, fragments of a tooth crown.

 

 

Locality: Sussex County, England, Southern United Kingdom.

Horizon: Wealden Formation.

Biostratigraphy:

Age: Early Cretaceous.

Material:

Royal Collage of Surgeons: Cast of a ungual phalanx.

 

Locality: Unknown.

Horizon:

Biostratigraphy:

Age: Unknown.

Material:

Royal Collage of Surgeons: Fragment of ribs.

 

ROYO Y GOMEZ, 1927

Locality: Benageber, Valence Province, Spain.

Horizon: Wealden Formation.

Biostratigraphy:

Age: Early Cretaceous.

Material:

Number: Not given:

 

SAUVAGE, 1900

Locality: Vesullien, Saint-Gaultier, Indre Department, France.

Horizon:

Biostratigraphy:

Age: Bathonian Stage, Dogger Epoch, Middle Jurassic.

Material:

Number: Not given: Tooth.

 

UBAGHS, 1892

Locality: Limbourg, Belgium.

Horizon:

Biostratigraphy:

Age: Late Cretaceous.

Material:

Number: Not given: Fragmentary tooth.

 

WEISHAMPEL, 1990, 1992 (Benton pers. comm.)

Locality: Northamptonshire County, England, Southern United Kingdom.

Horizon: Inferior Oolite.

Biostratigraphy:

Age: Aalenian-Bajocian Stage, Lower Dogger Epcoh, Early Jurassic.

Material:

Number: Not given:

 

Locality: Gloucestershire County, England, Southern United Kingdom.

Horizon: Inferior Oolite.

Biostratigraphy:

Age: Aalenian-Bajocian Stage, Lower Dogger Epoch, Early Jurassic.

Material:

Number: Not given:

 

Locality: Gloucestershire County, England, Southern United Kingdom.

Horizon: Forest Marble.

Biostratigraphy:

Age: Late Bathonian Stage, Dogger Epoch, Middle Jurassic.

Material:

Number: Not given:

= Metriacanthosaurus brevis WELLES, POWELL & PICKERING, 1995 (nomen nudum)
Etymology: Latin, brevis, "short."

Holotype:

Locality:

Horizon:

Biostratigraphy:

Age:

Material:

OUM J29888: Right pectoral girdle.

BMNH 31806: Right femur.

_____________________________________________________________________________________

Species: phillipsi WELLS, POWELL & PICKERING, 1995
Etymology:

Holotype:

Locality: Swindon, Wiltshire County, England

Horizon: Kimmeridge Clay.

Biostratigraphy:

Age: Kimmeridgian Stage, Middle Malm Epoch, Late Jurassic.

Material:

OUM J29886: Right tibia.

BMNH 31809: Left tibia.

OUM J13586: Left metatarsals II-IV

_____________________________________________________________________________________