Genus: Paronychodon COPE, 1876 (nomen dubium)
Etymology: Greek, para, “near,” Greek, onych- (onyx) “claw”, and Greek, odon, "tooth": Near claw tooth.

Species: lacustris COPE, 1876 (nomen dubium)
Etymology: Latin, lacustris, "lacustrine."

Holotype: AMNH 3018

Locality: Judith River, Montana.

Horizon: Hell Creek Formation.

Biostratigraphy:

Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.

Material: Tooth.

Referred material:

ARMSTRONG-ZEIGLER, 1978

Locality: MNA Locality A, 30 miles southwest of Farmington, New Mexico.

Horizon: Fruitland Formation.

Biostratigraphy:

Age: Judithian Age, Late Campanian Stage, Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.

Material:

MNA Pl. 1627: Several isolated teeth.
Note: Dromaeosauridae incertae according to LUCAS, MATEER, HUNT & O’NEILL, 1987.


BASZIO, 1997

Locality: Alberta, Canada.

Horizon: Milk River Formation.

Biostratigraphy:

Age: Aquilian age, Late Santonian Stage, Senonian subepoch, Upper Gulf epoch, Late Cretaceous

Material:

UA MR-4: 46-54: Teeth.

 

Locality: Alberta, Canada.

Horizon: Horseshoe Canyon Formation (Lower Edmonton Formation), Edmonton Group.

Biostratigraphy:

Age: Edmontonian age, lower Maastrichtian Stage, Upper Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.

Material:

RTMP (preliminary number) 1941: Tooth.

 

Locality: Alberta, Canada.

Horizon: Scollard Formation, Uppermost Edmonton Group.

Biostratigraphy:

Age: Lancian Age, uppermost Maastrichtian Stage, uppermost Senonian subepoch, Upper Upper Gulf Epoch, Late Cretaceous.

Material:

Alberta 1: 97, Alberta 2: 121: Teeth.

 

Locality:

Horizon: Lance Formation.

Biostratigraphy:

Age: Lancian Age, uppermost Maastrichtian Stage, uppermost Senonian subepoch, Upper Upper Gulf Epoch, Late Cretaceous.

Material:

UA BTB: 134, 135, 137, 146: Teeth.


BREITHAUPT, 1982

Locality: East Flank of Rock Springs Uplift, Sweetwater County, Wyoming.

Horizon: Lance Formation.

Biostratigraphy:

Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.

Material:

UW 14091, 14092: Isolated teeth.

 


BREITHAUPT, 1985, DEMAR & BREITHAUPT, 2006

Locality: UW V-81006, False Rocks Quarry (part of BQFR, Barwin Quarry False Rocks), southeastern Wind River Basin, Wyoming.

Horizon: Mesaverde Formation.

Biostratigraphy:

Age: Judithian Age, Campanian Stage, Senonian subepoch, Upper Gulf epoch, Late Cretaceous.

Material:

UW 34819, AMNH 12881, 12882: Teeth.

 

CARPENTER, 1982

Locality: UCMP V-5711, Bushy Tailed Blowout, Quarry area includes the floor of a small blowout approximately 10 yards northwest of the highest point on the ridge forming the northeastern side of the blowouts of loc. V-5003, S 1/2, Sec. 20, T37N, R64W, Lance Creek, Powder River Basin, Niobrara County, Wyoming.

Horizon: Lance Formation.

Biostratigraphy:

Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.

Material:

UCM 38288, 38459: Teeth of a baby.

 


Locality: UCMP V73087, Flat Creek 5, Garfield County, Montana.

Horizon: Hell Creek Formation.

Biostratigraphy:

Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.

Material:

UCMP 124990, 124991, 124992: Teeth of babies.

SANKEY, 2008

UCMP 123342, 187156, 187159, 187162, 187164: Teeth.

 


ESTES, 1964

Locality: UCMP V-5003, S 1/2, Sec. 20, T37N, R64W, Lance Creek, Powder River Basin, Niobrara County, Wyoming.

Localities: UC V-5616, Altman Blowout, this large blowout, site of the UCMP field-party camps form 1956 to 1960, is just east of the crest of the divide separating Lance Creek and Hancock Creek and 100 yards east of a dirt road. Fossils were found in anthills, talus, and sandstone exposed in the blowout, which is in the NE 1/4, Sec. 9, T37N, R64W, Eastern Wyoming.

Locality: UCMP V-5620, Lull Quarry, near the head of a tributary of Buck Creek, Powder River Basin, Niobrara County (Formerly Converse County), Wyoming.

Locality: UCMP V-5711, Bushy Tailed Blowout, Quarry area includes the floor of a small blowout approximately 10 yards northwest of the highest point on the ridge forming the northeastern side of the blowouts of loc. V-5003, S 1/2, Sec. 20, T37N, R64W, Lance Creek, Powder River Basin, Niobrara County, Wyoming.

Locality: UCMP V-5815, Hatcher Number 2, western side of a cross-stratified sandstone erosional remnant near the center of a large blowout on the crest of a ridge, the distance form the nearest point on the road, approximately 150 yards south of the quarry, to the Johnson ranch is 0.4 miles, Eastern Wyoming.

Locality: UCMP V-5918, Fish tooth Hill, Anthills and talus found in and around a badland area on the eastern side of a north-trending ridge extending from the crest of the divide between Cow and Dogie Creeks, Eastern Wyoming.

Horizon: Lance Formation.

Biostratigraphy:

Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.

Material:

Number: Not given: Teeth.

 


ESTES, BERBERIAN, & MESOZLEY, 1969

Locality: Bug Creek Anthills, Montana.

Horizon: Hell Creek Formation.

Biostratigraphy:

Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.

Material:

MCZ 3645: Two teeth.

 


= cf. Paronychodon lacustris LARSON, 2009

Locality: Alberta Province, Canada.

Horizon: Deadhorse Coulee Member, Milk River Formation.

Biostratigraphy:

Age: Aquilian age, Latest Santonian Stage, Senonian subepoch, Upper Gulf epoch, Late Cretaceous.

Material:

UALVP 48365-48413, 49596-49684, ROM 56623: Teeth.


SAHNI, 1972

Locality: Clambank Hollow, Montana.

Horizon: Judith River Formation.

Biostratigraphy:

Age: Judithian Age, Late Campanian Stage, Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.

Material:

AMNH 8522: Large tooth.

AMNH 8523: Small tooth.

AMNH 8522: Tooth.

3 other teeth.

 

SANKEY, 2008

Locality: UCMP V-5616, Altman Blowout, this large blowout, site of the UCMP field-party camps from 1956 to 1960, is just east of the crest of the divide separating Lance Creek and Hancock Creek and 100 yards east of a dirt road. Fossils were found in anthills, talus, and sandstone exposed in the blowout, which is in the NE 1/4, Sec. 9, T37N, R64W, Eastern Wyoming.

Horizon: Lance Formation.

Biostratigraphy:

Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.

Material:

UCMP 46585: Tooth.

 

Locality: UCMP V-5620, Lull Quarry, near the head of a tributary of Buck Creek, Powder River Basin, Niobrara County (Formerly Converse County), Wyoming.

Horizon: Lance Formation.

Biostratigraphy:

Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.

Material:

UCMP 53286, 124400, 187120, 187121, 187123, 187126, 187128, 187112, 187114, 187102, 187108, 187085, 187086, 187087, 187089, 187091, 187092, 187093, 187096, 187098, 187099, 187100, 187130, 187133, 187134: Teeth.

 

Locality: UCMP V-5815, Hatcher Number 2, western side of a cross-stratified sandstone erosional remnant near the center of a large blowout on the crest of a ridge, the distance form the nearest point on the road, approximately 150 yards south of the quarry, to the Johnson ranch is 0.4 miles, Eastern Wyoming.

Horizon: Lance Formation.

Biostratigraphy:

Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.

Material:

UCMP 73076: Tooth.

 

Locality: UCMP V65238, Niobrara County, Wyoming.

Horizon: Lance Formation.

Biostratigraphy:

Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.

Material:

UCMP 214007-214016: Teeth.

 

Locality: LACM 3075, = UCMP V72206, Meniscoessus skull, Garfield County, Montana.

Horizon: Hell Creek Formation.

Biostratigraphy:

Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.

Material:

UCMP 119787: Tooth.

 

Locality: UCMP V75173, Garfield County, Montana.

Horizon: Hell Creek Formation.

Biostratigraphy:

Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.

Material:

UCMP 214051-214055: Teeth.

 

Locality: UCMP V73076, Engdahl Anthills 2, Garfield County, Montana.

Horizon: Hell Creek Formation.

Biostratigraphy:

Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.

Material:

UCMP 120192: Tooth.

 

Locality: UCMP V73077, Engdahl Anthills 3, Garfield County, Montana.

Horizon: Hell Creek Formation.

Biostratigraphy:

Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.

Material:

UCMP 120254: Tooth.

 

The following UCMP localities are either in Montana or Wyoming, and either Lance or Hell Creek Formations.

UCMP V5821: UCMP 53065: Tooth.

UCMP V73092: UCMP 123525: Tooth.

UCMP V77131: UCMP 187150, 186931: Teeth.

UCMP V74107: UCMP 124405: Tooth.

UCMP V80091: UCMP 187151: Tooth.

UCMP V80119: UCM 186911: Tooth.

UCMP V82013: UCMP 187145, 187146: Teeth.

UCMP V82022: UCMP 187082: Tooth.

UCMP V83118: UCMP 186900: Tooth.

UCMP V84130: UCMP 214047-214050: Teeth.

UCMP V87034: UCMP 187214: Tooth.

UCMP V87035: UCMP 186913: Tooth.

UCMP V87038: UCMP 187141: Tooth.

UCMP V87151: UCMP 186908: Tooth.

UCMP V87084: UCMP 187144: Tooth.

 

= Paronychodon cf. P. lacustris SANKEY, STANDHART & SCHIEBOUT, 2005

Locality: LSUM 113, Running Lizard Site, Big Bend National Park, Brewster County, Texas.

Horizon: Aguja Formation.

Biostratigraphy:

Age: Early Maastrichtian Stage, Senonian subepoch, Upper Gulf epoch, Late Cretaceous.

Material:

LSUMG 1310, 1311, 5107, 5993, 5996: Teeth.

 


STENERSON & O’CONNER, 1994

Locality: Northwestern South Dakota.

Horizon: Hell Creek Formation.

Biostratigraphy:

Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.

Material:

Number: Not given: Teeth.

 

= Paronychodon sp. cf. P. lacustris STOKOSA, 2005

Locality: Eastern Wyoming.

Horizon: Hell Creek Formation.

Biostratigraphy:

Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.

Material:

SDSM 15101b: Tooth.

 

SULLIVAN & LUCAS, 2006

Locality: New Mexico.

Horizon: De-na-Zin Member, Kirtland Formation.

Biostratigraphy:

Age: Kirtlandian age, Upper Campanian Stage, Senonian Subepoch, Gulf Epoch, Late Cretaceous.

Material:

SMP VP-1354: Non-serrated tooth with wrinkled enamal.


TRIEBOLD, 1997

Locality: Sandy Site, Harding County, South Dakota.

Horizon: Hell Creek Formation.

Biostratigraphy:

Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.

Material:

Number: Not given: Teeth.

 

WHITMORE, 1988

Locality: SDSM Locality V851, Zerbst Locality, east of the Cheyenne River, Niobrara County, Wyoming.

Horizon: Lance Formation.

Biostratigraphy:

Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.

Material:

SDSM 12459: 2 teeth.

SDSM 15101: 1 tooth fragment.

 


Locality: SDSM Locality V852, Hanson Locality, Niobrara County, Wyoming.

Horizon: Lance Formation.

Biostratigraphy:

Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.

Material:

SDSM 12460: 1 tooth.

 

 

Locality: SDSM Locality V853, Leonard’s Lizard Site, Niobrara County, Wyoming.

Horizon: Lance Formation.

Biostratigraphy:

Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.

Material:

SDSM 12461: 1 tooth.

 

Locality: SDSM Locality V685, Greasewood Creek Locality, northern side of the south fork of the Greasewood Creek, Niobrara County, Wyoming.

Horizon: Lance Formation.

Biostratigraphy:

Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.

Material:

SDSM 12707: 1 tooth.

 


WROBLEWSKI, 1995, 1998

Locality: Western Hanna Basin, Carbon County, Wyoming.

Horizon: Ferris Formation, near the K/T Boundary.

Biostratigraphy:

Age: Lancian age, uppermost Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.

Material:

Number: Not given: Teeth.

_____________________________________________________________________________________

Species: caperatus (MARSH, 1889) OLSHEVSKY, 1991 (nomen dubium)
Etymology: Latin, caperatus, "wrinkled."
= Tripriodon caperatus MARSH, 1889 (nomen dubium)
= Menisocoessus caperatus (MARSH, 1889) HAY, 1902 (nomen dubium)
= Triprotodon caperatus (MARSH, 1889) CHURE & McINTOSH, 1989 (sic)
Dipriodon caperatus (MARSH, 1889) CURRIE, RIGBY & SLOAN, 1990

Holotype: YPM 11853

Locality: Wyoming.

Horizon: Laramie Formation.

Biostratigraphy:

Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.

Material: Isolated lower incisor.

Referred material:

Number: Not given: Teeth

_____________________________________________________________________________________

Species: asiaticus (NESOV, 1995) SUES & AVERIANOV, 2013
= Euronychodon asiaticus NESOV, 1995
Etymology: In reference to Asia, where the specimen was found.
= Paronychodon cf. lacustris NESOV, 1985
= Paronychodon -like NESOV, 1992

Holotype: TsNIGR museum #9/12454

Locality: Lower part of site CBI-14, Dzhyrakuuk, Kizylkum Desert, Uzbekistan.

Horizon: Middle of Bissekt Formation, (Bissektiyskaya Svita).

Biostratigraphy:

Age: Coniacian Stage, Lower Senonian Subepoch, Gulf Epoch, Early Late Cretaceous.

Material: 7 teeth.

Referred material:

SUES & AVERIANOV, 2013

ZIN PH 3/1/16, 1052/16, 1053/16, 1055/16, 1056/16, 1057/16, 1058/16, 1060/16, 1062/16, 1063/16, 1064/16, 1065/16, 1066/16, 1969/16, 1070/16, 1072/16, 1149/16, 1150/16, 1151/16, 1152/16, 1833/16: Teeth.

 

Locality: SKH-20, Khodzhakul, Kyzylkum Desert, Uzbekisan.

Horizon: Khodzhakul Formation.

Biostratigraphy:

Age: Cenomanian Stage, Upper Gallic Subepoch, Lowermost Gulf Epoch, Middle Cretaceous¬-Turonian Stage, upper Gallic subepoch, lower Gulf Epoch, Early Late Cretaceous.

Material:

ZIN PH 1071/16, 1221/16: Teeth.

 

Locality: SSHD-8, Sheikhdzheili, Kyzylkum Desert, Uzbekisan.

Horizon: Khodzhakul Formation.

Biostratigraphy:

Age: Cenomanian Stage, Upper Gallic Subepoch, Lowermost Gulf Epoch, Middle Cretaceous¬-Turonian Stage, upper Gallic subepoch, lower Gulf Epoch, Early Late Cretaceous.

Material:

ZIN PH 1220/16, 1228/16, 1229/16, 1230/16: Teeth.

 

Locality: CBI-17a, Dzhyrakuuk, Kizylkum Desert, Uzbekistan.

Horizon: Middle of Bissekt Formation, (Bissektiyskaya Svita).

Biostratigraphy:

Age: Coniacian Stage, Lower Senonian Subepoch, Gulf Epoch, Early Late Cretaceous.

Material:

ZIN PH 1054/16, 1059/16, 1068/16: Teeth.

 

Locality: CBI-5a, Dzhyrakuuk, Kizylkum Desert, Uzbekistan.

Horizon: Middle of Bissekt Formation, (Bissektiyskaya Svita).

Biostratigraphy:

Age: Coniacian Stage, Lower Senonian Subepoch, Gulf Epoch, Early Late Cretaceous.

Material:

ZIN PH 1061/16, 1067/16, 1148/16: Teeth.

 

Locality: CBI-4, Dzhyrakuuk, Kizylkum Desert, Uzbekistan.

Horizon: Middle of Bissekt Formation, (Bissektiyskaya Svita).

Biostratigraphy:

Age: Coniacian Stage, Lower Senonian Subepoch, Gulf Epoch, Early Late Cretaceous.

Material:

ZIN PH 1147/16: Tooth.

 

Locality: UCH-4, Uchkuduk, Kizylkum Desert, Uzbekistan.

Horizon: Middle of Bissekt Formation, (Bissektiyskaya Svita).

Biostratigraphy:

Age: Coniacian Stage, Lower Senonian Subepoch, Gulf Epoch, Early Late Cretaceous.

Material:

ZIN PH 1345/16: Tooth.