Genus: Ricardoestesia CURRIE, RIGBY & SLOAN, 1990
Etymology: In honor of Richard Estes, whose 1964 paper on Lance Formation microvertebrate
fossils demonstrated the use of theropod teeth in faunal studies.
= Asiamericana NESOV, 1995
Etymology: From the continent Asiamercia, considered to have been a single
land mass during the Coniacian.
= Richardoestesia CURRIE, RIGBY & SLOAN,
1990 (sic)
Note: Richardoestesia is a common misspelling of the name. The authors
wanted the name to be spelt Ricardoestesia (pers. comm..) as per George
Olshevsky’s
(1992) taxa list, Ricardo is the Latinization of Richard.
Species: gilmorei CURRIE, RIGBY & SLOAN, 1990
Etymology: In honor of G. W. Gilmore who first described this specimen.
= Richardoestesia gilmorei CURRIE, RIGBY & SLOAN, 1990 (sic)
Holotype: NMC 343
Locality: Dinosaur Provincial Park, Sec. 30, tp. 20, RIIW, 4th mer. 2 1/2 miles south of the mouth of Little Sandhill creek, Red Deer River, Newell County, Alberta Province, Canada.
Horizon: Dinosaur Park Formation, upper Judith River Group.
Biostratigraphy:
Age: Judithian age, Middle Campanian Stage, middle Senonian subepoch, upper Gulf Epoch, Late Cretaceous.
Material: Remains of a pair of dentaries with unerupted and germ teeth.
Referred material:
CURRIE, RIGBY & SLOAN, 1990
RTMP 80.8.298, 84.89.274, 83.129.11, 80.16.1230: Teeth.
Note: Troodontid?
BASZIO, 1997Locality: Alberta, Canada.
Horizon: Dinosaur Park Formation, upper Judith River Group.
Biostratigraphy:
Age: Judithian age, Middle Campanian Stage, middle Senonian subepoch, upper Gulf Epoch, Late Cretaceous.
Material:
RTMP 65.26.13, 85.50.204, 86.171.61, 86.23.105, 86.23.90, 87.116.60, 89.36.156, 83.36.242: Teeth.
Locality: Alberta, Canada.Horizon: Milk River Formation.
Biostratigraphy:Age: Aquilian age, Late Santonian Stage, Senonian subepoch, Upper Gulf epoch, Late Cretaceous
Material:
UA MR-4: 18-20, 4-17: Teeth.
Locality: Stephen’s site, Wheatland County, Alberta Province, Canada.
Horizon: Horseshoe Canyon Formation (Lower Edmonton Formation), Edmonton Group.
Biostratigraphy:
Age: Edmontonian age, lower Maastrichtian Stage, Upper Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.
Material:
RTMP 1016-1018: Teeth.
Locality: Janet’s site, Alberta, Canada.Horizon: Horseshoe Canyon Formation (Lower Edmonton Formation), Edmonton Group.
Biostratigraphy:
Age: Edmontonian age, lower Maastrichtian Stage, Upper Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.
Material:
RTMP 1026, 1027: Teeth.
Locality: Alberta, Canada.Horizon: Scollard Formation, Uppermost Edmonton Group.
Biostratigraphy:
Age: Lancian Age, uppermost Maastrichtian Stage, uppermost Senonian subepoch, Upper Upper Gulf Epoch, Late Cretaceous.
Material:
KUA-1: 85, 120, 108, LSF-140: 105, KUA-18: 114: Teeth.
Locality:
Horizon: Lance Formation.
Biostratigraphy:
Age: Lancian Age, uppermost Maastrichtian Stage, uppermost Senonian subepoch, Upper Upper Gulf Epoch, Late Cretaceous.
Material:
UA BTB: 135, 154, 155, 158-160: Teeth.
EATON, GARDNER, KIRKLAND, BRINKMAN, & NYDAM, 2014
Locality: UMNH VP Locality 119, Iron County, Utah.
Horizon: Iron Springs Formation.
Biostratigraphy:
Age: Upper Santonian Stage-Early Campanian Stage, Upper Senonian Sub-epoch, Upper Gulf Epoch, Late Cretaceous.
Material:
UMNH VP 19935: Tooth.
FIORILLO & CURRIE, 1994
Localities: Careless Creek Quarry, Emily’s Ankle Quarry, Hidden Valley Quarry, Blackbird Ridge Quarry, Wheatland and Golden Valley Counties, Central Montana.
Horizon: Judith River Formation.
Biostratigraphy:
Age: Judithian age, Campanian Stage, Senonian subepoch, Upper Gulf epoch, Late Cretaceous.
Material: ANSP 15824, 15942, 15952, 17647, 17782, 17784, 17983, 18101, 18104, 18114, 18115, 18119: Teeth.
Note: Exact distribution not given.
LEWIS, HECKERT, LUCAS & WILLIAMSON, 2007
Locality: NMMNH L. 5636, New Mexico.
Horizon: Allison Member, Menefee Formation.
Biostratigraphy:
Age: Lower Campanian Stage, Senonian Subepoch, Gulf Epoch, Late Cretaceous.
Material:
Number: Not given: Teeth.
= cf. Richardoestesai gilmorei LARSON, 2009
Locality: Alberta Province, Canada.
Horizon: Deadhorse Coulee Member, Milk River Formation.
Biostratigraphy:
Age: Aquilian age, Latest Santonian Stage, Senonian subepoch, Upper Gulf epoch, Late Cretaceous.
Material:
UALVP 47982, 47985, 47986, 47990, 47997, 48011, 48014, 48038, 48055, 48057, 48060, 48072, 48079, 48080, 48112-48115, 48125, 48130, 48138-48146, 48148-48150, 48152-48162, 48164-48170, 48172-48183, 48188, 48189, 48104-48197, 48199, 48202-48205, 48208-48217, 48189, 48914-48917, 48199, 48202-48205, 48208-48217, 48219, 48220, 48224-48226, 48231-48238, 48242, 48245-48247, 48250-48253, 48255, 48256, 48258-48263, 48265-48267, 48270-48277, 48457, 48459, 48466, 48469, 48470, 48473-48475, 49685-50047: Teeth.
= Richardoestesia cf. gilmorei SANKEY, 2001 (sic)
Locality: LSUMG VL-489, Talley Mountain fossil site, Big Bend National Park, Brewster County, Texas.
Horizon: Aguja Formation.
Biostratigraphy:
Age: Campanian Stage, Senonian subepoch, Upper Gulf epoch, Late Cretaceous.
Material:
LSUMG V-6237: Fragmentary tooth.
STEIN, 2021
Locality: Tooth Draw Deposit, Butte County, South Dakota.
Horizon: Hell Creek Formation.
Biostratigraphy:
Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.
Material:
TD-15-237: Tooth.
Number: Not given: Teeth.
= Richardoestesia sp. cf. R. gilmorei STOKOSA, 2005
Locality: Northwestern South Dakota.
Horizon: Hell Creek Formation.
Biostratigraphy:
Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.
Material:
SDSM 64372b: Tooth.
_____________________________________________________________________________________
Species: isosceles emend SANKEY, 2001
Etymology: In reference to the shape of the tooth in lateral view, an isosceles
trangle.
= Richardoestesia isosceles SANKEY, 2001 (sic)
Holotype: LSUMG 6238
Locality: LSUMG VL-489, Talley Mountain fossil site, Big Bend National Park, Brewster County, Texas.
Horizon: Aguja Formation.
Biostratigraphy:
Age: Edmontonian age, Lower Upper Campanian Stage, Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.
Material: Tooth.
Referred material:
LSUMG V-6233, 6234, 6235, 6237, 6238, 6050: Teeth.
Locality: LSUMG VL-492, Talley Mountain fossil site, Big Bend National Park, Brewster County, Texas.Horizon: Aguja Formation.
Biostratigraphy:
Age: Edmontonian age, Lower Upper Campanian Stage, Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.
Material:
LSUMG V-6264: Tooth.
RTMP 91.1709:
A.M. 8113 (ESTES, 1969)
LSUMG 5483 and 6239
CARBOT-CHANONA & RIVERA-SYLVA, 2011
Locality: Ocozocoualta, Chiapas State, Mexico.
Horizon: Ocozocoautla Formation.
Biostratigraphy:
Age: Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.
Material:
IHNFG-0537: Isolated maxillary tooth.
DEMAR & BREITHAUPT, 2006
Locality: UW V-81006, False Rocks Quarry (part of BQFR, Barwin Quarry False Rocks), southeastern Wind River Basin, Wyoming.
Horizon: Mesaverde Formation.
Biostratigraphy:
Age: Judithian Age, Campanian Stage, Senonian subepoch, Upper Gulf epoch, Late Cretaceous.
Material:
UW 34822: Tooth.
Locality: UW V-81032, Wyoming.
Horizon: Mesaverde Formation.
Biostratigraphy:
Age: Judithian Age, Campanian Stage, Senonian subepoch, Upper Gulf epoch, Late Cretaceous.
Material:
UW 34821: Tooth.
DUMONT, TAFFOREAU, BERTIN, BHULLAR, FIELD, SCHULP, STRILISKY, THIVICHON-PRINCE, VIRIOT, & LOUCHART, 2016
Locality: Wendy's Site, Pinhorn Ranch, Alberta Province, Canada.
Horizon: Oldman Formation.
Biostratigraphy:
Age: Middle Campanian Stage, Senonian Subepoch, Gulf Epoch, Late Cretaceous.
Material:
RTMP 1989.103.0025: Tooth.
Note: Orginally refered to as a bird by SANKEY, & BRINKMAN, 2002.
EATON, GARDNER, KIRKLAND, BRINKMAN, & NYDAM, 2014
Locality: UMNH VP Locality 12, Iron County, Utah.
Horizon: Iron Springs Formation.
Biostratigraphy:
Age: Upper Santonian Stage-Early Campanian Stage, Upper Senonian Sub-epoch, Upper Gulf Epoch, Late Cretaceous.
Material:
UMNH VP 24115: Tooth.
= Rhicardoestesia sp. cf. R. isosceles GARRISON, BRINKMAN, NICHOLS, LAYER, BURGE & THAYN, 2007
Locality: Eolambia II (Eo 2) site (Cifelli #2 Quarry), Emery County, Utah.
Horizon: Mussentuchit Member, Uppermost Cedar Mountain Formation.
Biostratigraphy:
Age: Mussentuchitan age, Uppermost Albian-Lowermost Cenomanian Stage, Upper Gallic subepoch, Lowest Gulf Epoch, Late Early to Early Middle Cretaceous.
Material:
Number: Not given:
= cf. Richardoestesia isosceles LARSON, 2009
Locality: Alberta Province, Canada.
Horizon: Deadhorse Coulee Member, Milk River Formation.
Biostratigraphy:
Age: Aquilian age, Latest Santonian Stage, Senonian subepoch, Upper Gulf epoch, Late Cretaceous.
Material:
UALVP 48147, 48151, 48171, 48185-48487, 48190-48193, 48198, 48200, 48201, 48206, 48207, 48218, 48222, 48227-48230, 48236, 48239-48241, 48243, 48244, 48248, 48254, 48257, 48264, 48268, 48269, 48278-48264, 48451, 50048-50209, CMNFV 8727: Teeth.
LEWIS, HECKERT, LUCAS & WILLIAMSON, 2007
Locality: NMMNH L. 5636, New Mexico.
Horizon: Allison Member, Menefee Formation.
Biostratigraphy:
Age: Lower Campanian Stage, Senonian Subepoch, Gulf Epoch, Late Cretaceous.
Material:
Number: Not given: Teeth.
= Tyrannosauridae incerte sedis by RUSSELL, 1987 (moved to the Richardoestesia isosceles EVANS, BARRETT & SEYMOUR, 2012)Locality: Henry Farm locality, Scollard Canyon, Alberta Province, Canada.
Horizon: Scollard Formation, Uppermost Edmonton Group.
Biostratigraphy:
Age: Lancian Age, uppermost Maastrichtian Stage, uppermost Senonian subepoch, Upper Upper Gulf Epoch, Late Cretaceous.
Material:
ROM 26345: Incomplete tooth.
SANKY, 2008
Locality: UCMP V-5620, Lull Quarry, near the head of a tributary of Buck Creek, Powder River Basin, Niobrara County (Formerly Converse County), Wyoming.
Horizon: Lance Formation.
Biostratigraphy:
Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.
Material:
UCMP 213983-213986: Teeth.
Locality: V73077, Engdahl Anthills 3, Garfield County, Montana.
Horizon: Hell Creek Formation.
Biostratigraphy:
Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.
Material:
UCMP 213987-213991, 120257: Teeth.
Locality: UCMP V73087 (Flat Creek 5), McCone County, Montana.
Horizon: Hell Creek Formation.
Biostratigraphy:
Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.
Material:
UCMP 213922-213979, 213992-214006: Teeth.
Locality: V73103, Dry Fork Coluee, Garfield County, Montana.
Horizon: Hell Creek Formation.
Biostratigraphy:
Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.
Material:
UCMP 186842: Teeth.
The following UCMP localities are either in Montana or Wyoming, and either Lance or Hell Creek Formations.
UCMP 71118: UCMP 186837: Tooth.
UCMP V80088: UCMP 186827: Tooth.
UCMP V80091: UCMP 186835: Tooth.
UCMP V80092: UCMP 186824, 186825: Tooth.
UCMP V82023: UCMP 186833, 186833: Teeth.
UCMP V82060: UCMP 186834, 186389: Tooth.
UCMP V83044: UCMP 186829: Tooth.
UCMP V87030: UCMP 186836: Tooth.
UCMP V87072: UCMP 213980-213982: Teeth.
UCMP V87082: UCMP 186840: Tooth.
SANKY, STANDHART & SCHIEBOUT, 2005
Locality: LSUM 140, Talley Mountain, Big Bend National Park, Brewster County, Texas.
Horizon: Aguja Formation.
Biostratigraphy:
Age: Edmontonian age, Lower Upper Campanian Stage, Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.
Material:
LSUMG 6140, 6151: Teeth.
Locality: LSUM 492, Big Bend National Park, Brewster County, Texas.Horizon: Aguja Formation.
Biostratigraphy:
Age: Edmontonian age, Lower Upper Campanian Stage, Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.
Material:
LSUMG 6264: Tooth.
Locality: LSUM 741, Big Bend National Park, Brewster County, Texas.
Horizon: Aguja Formation.
Biostratigraphy:
Age: Late Campanian – Early Maastrichtian Stage, Senonian subepoch, Upper Gulf epoch, Late Cretaceous.
Material:
LSUMG 5933, 5934: Teeth.
Locality: LSUM 113, Running Lizard Site, Big Bend National Park, Brewster County, Texas.Horizon: Aguja Formation.
Biostratigraphy:
Age: Early Maastrichtian Stage, Senonian subepoch, Upper Gulf epoch, Late Cretaceous.
Material:
LSUMG 5939: Tooth.
Note: as per SANKEY, 2001…Based on large collections of dinosaur teeth in the RTMP, teeth from R. isosceles and R. gilmorei have been identified from the Milk River, Oldman, Dinosaur Park, Horseshoe Canyon, and Scollard Formations of Alberta; the Frenchman Formation of Saskatchewan; the Lance Formation of Wyoming (BASZIO, 1997); and the Hell Creek Formation of North and South Dakota (personal observation). However, the two taxa have different relative abundances at different sites, reflecting their different paleoecologies: R. isosceles is abundant in the Milk River, Scollard, Lance and Hell Creek Formations and are rare in the Oldman, Dinosaur Park, and Horseshoe Canyon Formations; the latter units representing wetter, more closed habitats…
_____________________________________________________________________________________
Species: asiatica (NESOV, 1995) SUES & AVERIANOV,
2013
= Asiamericana asiatica NESOV,1995
Etymology: From the continent Asiamerica, considered to have been a single land
mass during the Coniacian.
= Genus: Nova? NESOV, 1992
Holotype: St. Petersburg Central Geological Research Museum, CCMGE 460/12457
Locality: Locality CBI-14, Dzhyrakuuk, Kizylkum Desert, Uzbekistan.
Horizon: Bissekty Formation.
Biostratigraphy:
Age: Coniacian Stage, Lower Senonian Subepoch, Lower Gulf Epoch, Late Cretaceous.
Material: 3 teeth.
Referred material:
Number: Not given.
Similar teeth have been found from the Santonian to Lower Campanian of Kazakhstan.
SUES & AVERIANOV, 2013
ZIN PH 1073/16-1078/16, 1080/16-1108/16, 1110/16-1114/16, 1116/16-1118/16, 1120/16, 1122/16-1124/16, 1127/16,1128/16-1133/16, 1135/16-1137/16, 1139/16, 1141/16-1144/16: Teeth.
AVERIANOV & SUES, 2019
Morpohotype A
ZIN PH 1079/16, 1129/16, 2455/16, 2520/16: Teeth.
Morphotype B
ZIN PH 2593/16: Tooth.
Motphotype C
ZIN PH 1094/16, 1112/16, 1134/16, 1145/16, 2486/16, 2450/16, 2461/16, 2489/16, 2502/16, 2503/16, 2505/16, 2530/16, 2564/16, 2578-2580/16, 2600/16: Teeth.
Morphotype D
ZIN PH 2472/16, 2490/16, 2521/16, 2506/16, 2559/16, 2588/16, 2563/16: Teeth.
Morphotype E
ZIN PH 2553/16, 2565/16, 2597/16: Teeth.
Locality: SKH-20, Khodzhakul, Kyzylkum Desert, Uzbekisan.
Horizon: Khodzhakul Formation.
Biostratigraphy:
Age: Cenomanian Stage, Upper Gallic Subepoch, Lowermost Gulf Epoch, Middle Cretaceous¬-Turonian Stage, upper Gallic subepoch, lower Gulf Epoch, Early Late Cretaceous.
Material:
ZIN PH 1222/16: Tooth.
Locality: SSHD-8, Sheikhdzheili, Kyzylkum Desert, Uzbekisan.
Horizon: Khodzhakul Formation.
Biostratigraphy:
Age: Cenomanian Stage, Upper Gallic Subepoch, Lowermost Gulf Epoch, Middle Cretaceous¬-Turonian Stage, upper Gallic subepoch, lower Gulf Epoch, Early Late Cretaceous.
Material:
ZIN PH 1223/16, 1224/16, 1225/16, 1226/16, 1227/16: Teeth.
Locality: CBI, Dzhyrakuuk, Kizylkum Desert, Uzbekistan.
Horizon: Middle of Bissekt Formation, (Bissektiyskaya Svita).
Biostratigraphy:
Age: Coniacian Stage, Lower Senonian Subepoch, Gulf Epoch, Early Late Cretaceous.
Material:
ZIN PH 2315/16: Teeth.
Locality: CBI-17, Dzhyrakuuk, Kizylkum Desert, Uzbekistan.
Horizon: Middle of Bissekt Formation, (Bissektiyskaya Svita).
Biostratigraphy:
Age: Coniacian Stage, Lower Senonian Subepoch, Gulf Epoch, Early Late Cretaceous.
Material:
ZIN PH 1140/16: Teeth.
Locality: CBI-17a, Dzhyrakuuk, Kizylkum Desert, Uzbekistan.
Horizon: Middle of Bissekt Formation, (Bissektiyskaya Svita).
Biostratigraphy:
Age: Coniacian Stage, Lower Senonian Subepoch, Gulf Epoch, Early Late Cretaceous.
Material:
ZIN PH 1119/16, 1121/16: Teeth.
Locality: CBI-5, Dzhyrakuuk, Kizylkum Desert, Uzbekistan.
Horizon: Middle of Bissekt Formation, (Bissektiyskaya Svita).
Biostratigraphy:
Age: Coniacian Stage, Lower Senonian Subepoch, Gulf Epoch, Early Late Cretaceous.
Material:
ZIN PH 1109/16: Teeth.
Locality: CBI-5a, Dzhyrakuuk, Kizylkum Desert, Uzbekistan.
Horizon: Middle of Bissekt Formation, (Bissektiyskaya Svita).
Biostratigraphy:
Age: Coniacian Stage, Lower Senonian Subepoch, Gulf Epoch, Early Late Cretaceous.
Material:
ZIN PH 1115/16, 1125/16: Teeth.
Locality: CBI-4, Dzhyrakuuk, Kizylkum Desert, Uzbekistan.
Horizon: Middle of Bissekt Formation, (Bissektiyskaya Svita).
Biostratigraphy:
Age: Coniacian Stage, Lower Senonian Subepoch, Gulf Epoch, Early Late Cretaceous.
Material:
ZIN PH 1128/16: Tooth.
Locality: CBI-4v, Dzhyrakuuk, Kizylkum Desert, Uzbekistan.
Horizon: Middle of Bissekt Formation, (Bissektiyskaya Svita).
Biostratigraphy:
Age: Coniacian Stage, Lower Senonian Subepoch, Gulf Epoch, Early Late Cretaceous.
Material:
ZIN PH 1127/16: Tooth.
Locality: CDZH-17a, Dzhyrakuuk, Kizylkum Desert, Uzbekistan.
Horizon: Middle of Bissekt Formation, (Bissektiyskaya Svita).
Biostratigraphy:
Age: Coniacian Stage, Lower Senonian Subepoch, Gulf Epoch, Early Late Cretaceous.
Material:
ZIN PH 1138/16: Tooth.
Locality: ZK-3, Zengekurgon, Kizylkum Desert, Uzbekistan.
Horizon: Middle of Bissekt Formation, (Bissektiyskaya Svita).
Biostratigraphy:
Age: Coniacian Stage, Lower Senonian Subepoch, Gulf Epoch, Early Late Cretaceous.
Material:
Morphotype A
ZIN PH 1231/16: Tooth.
AVERIANOV & SUES, 2014
Locality: Dzhyrakuuk, Kizylkum Desert, Uzbekistan.
Horizon: Aitym Formation.
Biostratigraphy:
Age: Coniacian Stage, Lower Senonian Subepoch, Lower Gulf Epoch, Late Cretaceous.
Material:
Morphotype C
ZIN PH 2605/16: Tooth.