Genus: Torvosaurus GALTON & JENSEN, 1979
Etymology: Latin, torvus, "savage, cruel, wild" and Greek, sauros, "lizard."
= Torovosaurus KURZANOV, 1989 (sic)

Species: ingens (JANENSCH, 1920) emend SOTO, TORINO & PEREA, 2020 (nomen dubium)
Etymology: Latin, ingens, "enormous, massive."
= Megalosaurus? ingens JANENSCH, 1920 (nomen dubium)
= Ceratosaurus ingens (JANENSCH, 1920) ROWE & GAUTHIER, 1990

Holotype: MB R 1050

Locality: Tendaguru site B, Tendaguru, Tanzania, East Africa.

Horizon: Upper Dinosaur (Saurian bed) Member, Tendaguru Formation.

Biostratigraphy:

Age: Late Tithonian Stage, Upper Malm Epoch, Late Jurassic.

Material: Type a: Tooth.
Note: Tyrannosaurus size.

 

Locality: Tendaguru, Tanzania, East Africa.

Horizon: Lower Dinosaur (Saurian bed) Member, Middle Tendaguru Formation.

Biostratigraphy:

Age: Kimmeridigian Stage, Middle Malm Epoch, Late Jurassic.

Material:

MB R 1953, 1064: 2 tooth crowns.

 

Locality: Tendaguru, Tanzania, East Africa.

Horizon: Middle Dinosaur (Saurian bed) Member, Middle Tendaguru Formation.

Biostratigraphy:

Age: Kimmeridigian Stage, Middle Malm Epoch, Late Jurassic.

Material:

MB R 1054, 1057, 1058, 1061, 1067: 5 tooth crowns.

 

Locality: Tendaguru, Tanzania, East Africa.

Horizon: Upper Dinosaur (Saurian bed) Member, Tendaguru Formation.

Biostratigraphy:

Age: Tithonian Stage, Upper Malm Epoch, Late Jurassic.

Material:

MB R 1060, 1069: 2 teeth.

 

Locality: Tendaguru, Tanzania, East Africa.

Horizon: Unknown, Tendaguru Formation.

Biostratigraphy:

Age: Kimmeridigian Stage, Lower Malm Epoch, Late Jurassic.

Material:

MB R 1082: Tooth.

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Species: tanneri GALTON & JENSEN, 1979
Etymology: Named in honor of N. Eldon Tanner, first counselor in the First Presidency of The Church of Jesus Christ of Latter-day Saints.
= Edmarka BAKKER, KRALIS, SIEGWARTH & FILLA, 1992
Etymology: In honor of Dr. Bill Edmark, University of Colorado alumnus, who has made notable contributions to science and technology.
= Megalosaurus tanneri (GALTON & JENSEN, 1979) PAUL, 1988

Holotype: BYUVP 2002

Locality: Dry Mesa Quarry 1, sec. 23, T.50N, R.14, Mesa County, on the Uncompahgre Plateau, in Western Colorado.

Horizon: Lower Upper Brushy Basin Member, Morrison Formation.

Biostratigraphy: Zone 2.

Age: Lower Part, Comobluffian age, Kimmeridgian Stage, uppermost Malm Epoch, Late Jurassic.

Material: Left humerus.

Referred material:

BYUVP 2001: Right humerus, right radius, left and right ulna.

BYUVP 2003: Left dentary.

BYUVP: 2004a: Cervical 5.

BYUVP 2004b: Cervical 6.

BYUVP 2004c: Fragmentary cervical 7.

BYUVP 2004d: Fragmentary cervical 8.

BYUVP 2005: Dorsal vertebra 01.

BYUVP 2006: Fragmentary dorsal vertebra 05.

BYUVP 2007: Fragmentary dorsal vertebra 06.

BYUVP 2008: Fragmentary dorsal vertebra 12.

BYU 2006-2010: Dorsal vertebrae 5, 7, 10 and 12.

BYU 2010-2012: Left metacarpals I, II, and III.

BYU 2013: Right ilium.

BYU 2014: Left and right pubes.

BYU 2015: Right ischium.

BYU 2016: Right tibia, fibula, astragalus and calcaneum.

BYU 2017: Right tibia.

BYUVP 4882: Right premaxilla.

BYUVP 4883: Left and right jugal.

BYUVP 4884: Fragmentary atlas cervical 1.

BYUVP 4860: Fragmentary cervical 3.

BYUVP 4890: Fragmentary dorsal vertebra 14.

BYUVP 4976a: Fragmentary caudal 08 and fragmentary chevron.

BYUVP 4976b: Fragmentary caudal 09 and fragmentary chevron.

BYUVP 4976c: Fragmentary caudal 10 and fragmentary chevron.

BYUVP 4976d: Fragmentary caudal 11 and fragmentary chevron.

BYUVP 4976e: Fragmentary caudal 12 and fragmentary chevron.

BYUVP 4976f: Fragmentary caudal 12 and fragmentary chevron.

BYUVP 4976g: Fragmentary caudal 14 and fragmentary chevron.

BYUVP 4976h: Fragmentary caudal 15 and fragmentary chevron.

BYUVP 4976i: Fragmentary caudal 16.

BYUVP 4998: Dorsal vertebra 04.

BYUVP 5004: Fragmentary caudal 04 or 05.

BYUVP 5005: Left metatarsal III.

BYUVP 5009: Right metatarsal III.

BYUVP 5020: Left calcaneum.

BYUVP 5029: Right astragalus

BYUVP 5077: Right metatarsal II.

BYUVP 5086: Fragmentary caudal 02 or 03.

BYUVP 5110: Right quadrate.

BYUVP 5129: Left fibula.

BYUVP 5136: Fragmentary right fibula.

BYUVP 5147: Left metatarsal II.

BYUVP 5241: Left metatarsal III.

BYUVP 5277: Right metatarsal III.

BYUVP 5278: Right metatarsal IV.

BYUVP 5279: Left metatarsal IV.

BYUVP 5280: Left metatarsal III.

BYUVP 5281: Left metatarsal II.

BYUVP 5286: Left lachrymal.

BYUVP 5276: Right metatarsal II.

BYUVP 8966: Left metatarsal IV.

BYUVP 9013: Fragmentary caudal 12.

BYUVP 9090: Fragmentary dorsal vertebra 10.

BYUVP 9120: Fragmentary dorsal vertebra 07.

BYUVP 9121: Fragmentary dorsal vertebra 09.

BYUVP 9135: Fragmentary caudal 22.

BYUVP 9249: Fragmentary right postorbital.

BYUVP 9620: Right fibula.

BYUVP 9621: Right astragalus.

BYUVP 9622: Right calcaneum.

McINTOSH, 1981

CM 14995: Cast of BYU 2020: Claw.

\

Composite skull.

 

Localitiy: Calico Gulch Quarry (195 km from the Dry Mesa Quarry), Moffat County, Colorado,

Horizon: Lower Upper Brushy Basin Member, Morrison Formation.

Biostratigraphy: Zone 2.

Age: Lower Part, Comobluffian age, Kimmeridgian Stage, uppermost Malm Epoch, Late Jurassic.

Material:

BYU 2018: Left phalanx of digit I of manus.

 


Locality: Quarry face as exposed inside the Dinosaur National Monument Visitor Center, Carnegie Quarry, Vernal, Uintah County, Utah.

Horizon: Brushy Basin Member, Morrison Formation.

Biostratigraphy: Zone 3.

Age: Upper Part, Comobluffian age, Kimmeridgian Stage, uppermost Malm Epoch, Late Jurassic.

Material:

DMNH 2243: Posterior dorsal vertebra.

 

DALMAN, 2014

Locality: Quarry 12, Como Bluff, Albany County, Wyoming.

Horizon: Member? Morrison Formation.

Biostratigraphy: Zone?

Age: Kimmeridgian/Tithonian Stage, Malm Epoch, Late Jurassic.

Material:

YPM VP 058269: A nearly complete left tibia that is missing the distal end.


HANSON & MAKOVICKY, 2014

Locality: Freezeout Hills Quarry 6, Carbon County, Wyoming.

Horizon: Brushy Basin, Member, Morrison Formation.

Biostratigraphy:

Age: Early Tithonian Stage, Malm Epoch, Late Jurassic.

Material:

FMNH PR 3060: 3 midline fragments of gastralia, right metacarpal III, right manual phalanx III-2, Left metatarsals II-IV, and left pedla phalanx I-1.

 

LULL, 1927

Locality: (Marsh’s Quarry, same?) Quarry 9 (Mammal Quarry), Como Bluff, (Sec. 12, T22N, R77W), Carbon County, Wyoming.

Horizon: Morrison Formation.

Biostratigraphy: Zone ?.

Age: Age?, Stage?, Epoch?, Late Jurassic.

Material:

Unnumbered: A large theropod tooth.

 


= cf. T. tanneri TURNER & PETERSON, 1999

Locality: Meyer Sites 1 and 2, Garden Park area, Canon City, Fremont county, Colorado.

Horizon: Salt Wash Member, Morrison Formation.

Biostratigraphy: Zone 2.

Age: Comobluffian age, Kimmeridgian Stage, uppermost Malm Epoch, Late Jurassic.

Material:

Number: Not given:


= Edmarka rex BAKKER, KRAILS, SIEGWARTH & FILLA, 1992
Etymology: Latin, rex, "king."

Holotype: CPS 1005

Locality: Nail Quarry, Como Bluff, Como Bluff East (East of Marshall road), Albany County, Wyoming.

Horizon: Lower Upper Brushy Basin Member, Morrison Formation.

Biostratigraphy: Zone 2.

Age: Comobluffian age, Kimmeridgian Stage, uppermost Malm Epoch, Late Jurassic.

Material: Left jugal, missing only the extreme dorsal tip of the postorbital process and the posterior tip of the lower quadratojugal process.

Referred material:

CPS 1004: Dorsal ribs.

CPS 1002: Left scapulocoracoid.

CPS 1005: Proximal caudal.

CPS 1003: Left coracoid.

 


Locality: Louise Quarry, Como Bluff, West (west of Marshall Road), Albany County, Wyoming.

Horizon: Brushy Basin Member, Morrison Formation.

Biostratigraphy: Zone 2.

Age: Comobluffian age, Kimmeridgian Stage, uppermost Malm Epoch, Late Jurassic.

Material:

CPS 401: 6 right dorsal ribs.

Note: Torvosauridae?


_____________________________________________________________________________________

Species: gurneyi HENDRICKX & MATEUS, 2014
Etymology: In honor of the paleoartist James Gurney, creator of the utopic world of Dinotopia.
= Torvosaurus tanneri MATEUS, WALEN & TELLES ANTUNES, 2006

Holotype: ML 1100

Locality: Cliffs of Praia da Vermelha, Lourinha, Estremadura Province, West coast of Portugal.

Horizon: Porto Novo Member, Lourinha Formation.

Biostratigraphy:

Age: Upper Kimmeridgian Stage, Middle Malm Epoch, Late Jurassic.

Material:Left maxilla with tooth.

 

Referred material:

FUB PG Ther 1: Tooth.

ML 430: An incomplete tibia.

ML 632: A parital femur.

 

MATEUS, MATEUS, TELLES ANTUNES, MATEUS, TAQUET, RIBEIRO & MANUPPELLA, 1997

Number: Not given: Several well preserved clutches, one egg has an embryo of a theropod? Centra, proximal part of a right femur, right tibia, partial left scapula?, distal part of a femur, part of a skull floor with occipital condyle and spheno-occipital tubercles.

ML 1186: cranial and post cranial material of embryos.

 

= cf. Torvosaurus sp MALAFAIA, ORTEGA, SILVA & ESCASO, 2008

Locality: Torres Vedras, Praia da Corva, Lourinha, Estremadura Province, West coast of Portugal.

Horizon: Porto Novo Member, Alcobaca Formation, Lourinha Group.

Biostratigraphy:

Age: Kimmeridgian Stage, Middle Malm Epoch, Late Jurassic.

Material:

ALT-SHN.116: Fragment of a maxilla.

 

= cf. Torvosaurus gurneyi MALAFAIA, MOCHO, ESCASO, NARVAEZ & ORTEGA, 2024

= Megalosaurus pombali, Megalosaurus insignis LAPPARENT & ZBSZEWSKI, 1957

Locality: Montoito, Lourinha, Estremadura Province, Portugal.

Horizon: Bonbarral Formation.

Biostratigraphy:

Age: Tithonian Stage, Middle Malm Epoch, Late Jurassic.

Material:

MG 4813: Fragment of tooth crown.

 

Locality: Paraia de S. Bernardino, Peniche Municipality, Portugal.

Horizon: Alcobaca Formation.

Biostratigraphy:

Age: Luistanien, Kimmeridgian Stage, Middle Malm Epoch, Early Jurassic.

Material:

MG 4818: Fragment of tooth fragment.

 

Locality: Porto Dinheiro, Lourinha, Estremadura Province, Portugal.

Horizon: Praia da Amoreira-Porto Novo Formation.

Biostratigraphy:

Age: Late Kimmeridgian-Tithonian Stage, Middle Malm Epoch, Late Jurassic.

Material:

MNHN/UL.EPt.8628: Large mesial tooth.

 

MALAFAIA, MOCHO, ESCASO & ORTEGA, 2017

Locality: Praia da Vermelha, Peniche, Portugal.

Horizon: Upper levels of the Praia da Amoreira-Porto Novo Formation.

Biostratigraphy:

Age: Late Kimmeridgian-lowermost Tithonian Stage, Malm Epoch, Late Jurassic.

Material:

SHN.400: An almost complet right maxilla.