Genus: Diplocynodon POMEL, 1847
Etymology: Greek, diplos, "double," Greek, cyno, "dog" and Greek, odon, "tooth."
= Baryphracta FREY, LAEMMERT & RIESS, 1987
= Diplyocynodus POMEL, 1847 (sic)
= Orthosaurus GREFFROY, 1833
Etymology:
= Plerodon MEYER, 1839
Etymology:
= Pleurodon MEYER, 1839 (sic)
= Saurocainus GERVIAS 1952
Etymology:

Species: plenidens (MEYER, 1838)
Etymology:
= Crocodilus plenidens MEYER, 1838
= Plerodon crocodiloides MEYER, 1839
Etymology:

Holotype:

Locality: Bohnerz of Mosskirch, Baden-Wurttemberg State, Germany.

Material:

Biostratigraphy:

Age: Miocene Epoch, Lower Neogene Period, Upper Tertiary Subera, Middle Cenozoic Era.

Material: Teeth.

Referred material:

LYDEKKER, 1888

Locality: Allstadt, near Mosskirch, Baden-Wurttemberg State, Germany.

Horizon:

Biostratigraphy:

Age: Middle (?) Miocene Epoch, Lower Neogene Period, Upper Tertiary Subera, Middle Cenozoic Era.

Material:

BMNH 43634: Imperfect crowns of 3 teeth.

_____________________________________________________________________________________

Species: hantoniensis (WOOD, 1844) POMEL, 1853
Etymology:
= Alligator hantoniensis WOOD, 1844

Holotype: NHMUK OR (old BMNH) 25166

Locality: Hordwell Cliff, near Lymington, , Hamphshire County, England, Southern United Kingdom.

Horizon: Totland Bay Member, Headon Hill Formation.

Biostratigraphy:

Age: Priabonian, Late Eocene Epoch, Paleogene Period, Lower Tertiary Subera, Lower Cenozoic Era-Oligocene Epoch, Upper Paleogene Period, Upper Tertiary Subera, Middle Cenozoic.

Material: Anterior portion of skull.

Referred material:

NHMUK OR (old BMNH) 5009: Teeth.

NHMUK R (old BMNH) 5010: Partial maxilla.

NHMUK R (old BMNH) 5121, 5212, 5216, 5217: Dentaries.

NHMUK R (old BMNH) 5213: Vertebra.

NHMUK R (old BMNH) 5215: Femur.

NHMUK R (old BMNH) 5217: partial angular.

NHMUK R (old BMNH) 5267: Partial basioccipital, partial vertebra patial osteoderm, partial femur, partial metapodial..

NHMUK OR (old BMNH) R1041: Fragmentary skull vertebrae.

NHMUK OR (old BMNH) R1042: Fragmentary skull of a juvenile.

NHMUK OR (old BMNH) R1043: Mandible, teeth, ribs, and scutes of an immature individual.

NHMUK OR (old BMNH) R1044: Right mandible of a juvenile.

NHMUK OR (old BMNH) R1045: Posterior portion of the left ramus of a juvenile, partial osteoderms. .

NHMUK OR (old BMNH) R1046: Numerous cervical vertebrae.

NHMUK OR (old BMNH) R1047, R1048: Dorsal vertebrae.

NHMUK OR (old BMNH) R1049: 3 lumbar vertebrae.

NHMUK OR (old BMNH) R1050: 6 sacral vertebrae.

NHMUK OR (old BMNH) R1051: 1st caudal vertebra.

NHMUK OR (old BMNH) R1052: Numerous caudal vertebrae.

NHMUK OR (old BMNH) R1053: Partial metapodials, partial phalanges.

NHMUK OR (old BMNH) R1054: Radialepartial phalanges.

NHMUK OR (old BMNH) R1068: 5 dorsal vertebrae, ribs and dermal scutes.

NHMUK R (old BMNH) 6323, 6852: Teeth.

NHMUK OR (old BMNH) R2053: Several phalangeals.

NHMUK OR (old BMNH) 20281: Partial lower jaw.

NHMUK OR (old BMNH) 25165: Teeth.

NHMUK OR (old BMNH) 25167: Anterior portion of a juvenile skull.

NHMUK OR (old BMNH) 25168: Left maxillo-palatine region of a juvenile.

NHMUK OR (old BMNH) 25169: Left maxillo-palatine region of a juvenile.

NHMUK OR (old BMNH) 25170: 2 fragments of a palate of a juvenile.

NHMUK OR (old BMNH) 25170a: Fragmentary skull of a juvenile.

NHMUK OR (old BMNH) 25174: Osteoderm.

NHMUK OR (old BMNH) 25175: 1st caudal vertebra.

NHMUK OR (old BMNH) 25176, 25177: 3 cervical vertebrae.

NHMUK OR (old BMNH) 25178: Lower jaw.

NHMUK OR (old BMNH) 25179: A sacral vertebra.

NHMUK OR (old BMNH) 25182: 3 dorsal vertebrae.

NHMUK OR (old BMNH) 25188: Quadrato-jugal arch.

NHMUK OR (old BMNH) 25186: Basisphenoid.

NHMUK OR (old BMNH) 25187: Quadrate.

NHMUK OR (old BMNH) 25188: jutals with partial quadratojugal.

NHMUK OR (old BMNH) 25191: Ribs.

NHMUK OR (old BMNH) 25195, 25199: Vertebre.

NHMUK OR (old BMNH) 25291: Articular.

NHMUK OR (old BMNH) 25193: Quadrate.

NHMUK OR (old BMNH) 25201: A cervical vertebra.

NHMUK OR (old BMNH) 25207: Lumbar vertebra.

NHMUK OR (old BMNH) 25208, 25209, 25210: Vertebrae.

NHMUK OR (old BMNH) 25215: Vertebra.

NHMUK OR (old BMNH) 25213-4: 2 dorsal vertebrae.

NHMUK OR (old BMNH) 25216-7: 3 pterygoids.

NHMUK OR (old BMNH) 25217: Pterygoid.

NHMUK OR (old BMNH) 25219: Jugal.

NHMUK OR (old BMNH) 25222: Dorsal vertebrae.

NHMUK OR (old BMNH) 25231, 25232: Femora.

NHMUK OR (old BMNH) 25236: Tibia.

NHMUK OR (old BMNH) 25237: Humerus.

NHMUK OR (old BMNH) 25238: FEmur.

NHMUK OR (old BMNH) 25239: Ulna.

NHMUK OR (old BMNH) 25242-52: 2 metapodials.

NHMUK OR (old BMNH) 25244: Partial radius, partial femur, partial metapodial.

NHMUK OR (old BMNH) 25245: Left coarcoid.

NHMUK OR (old BMNH) 25248: 5 ribs.

NHMUK OR (old BMNH) 25252-3: 2 ilia.

NHMUK OR (old BMNH) 25260: Fragmentary maxilla of a juvenile.

NHMUK OR (old BMNH) 25598: Teeth.

NHMUK OR (old BMNH) 29694: Subadult skull.

NHMUK OR (old BMNH) 30206, 30206a: Right humerii.

NHMUK OR (old BMNH) 30210-11-12-13: 4 femora.

NHMUK OR (old BMNH) 30215-17: 3 tibiae.

NHMUK OR (old BMNH) 30219: Humerus.

NHMUK OR (old BMNH) 30236, 30237: Ulnae.

NHMUK OR (old BMNH) 30239, 30242: 2 metapodials.

NHMUK OR (old BMNH) 30215, 30216, 30217: Tibiae.

NHMUK OR (old BMNH) 30247: Fragmentary left scapula, partial humerus, partial ulna.

NHMUK OR (old BMNH) 30222, 30223, 30228: Femora.

NHMUK OR (old BMNH) 30233, 30234, 30235, 30241: 4 fibulae.

NHMUK OR (old BMNH) 30236-7: 2 ulnae.

NHMUK OR (old BMNH) 30240: Metapodal.

NHMUK OR (old BMNH) 30242, 30389: Ulnae.

NHMUK OR (old BMNH) 30243: Metapodal.

NHMUK OR (old BMNH) 30246: Metapodal.

NHMUK OR (old BMNH) 30247: Partial femora, metapodal.

NHMUK OR (old BMNH) 30251: Pterygoid wity ectopterygoids and palatine.

NHMUK OR (old BMNH) 30287: Partial skull.

NHMUK OR (old BMNH) 30289: Partial proatlas, partial vertebra.

NHMUK OR (old BMNH) 30307, 30308, 30318: Angulars, osteodemrs.

NHMUK OR (old BMNH) 30317: Partial tooth.

NHMUK OR (old BMNH) 30336: Partial metaodal.

NHMUK OR (old BMNH) 30359: Coracoid.

NHMUK OR (old BMNH) 30372: Quadrate.

NHMUK OR (old BMNH) 30274: The homologous bone of the right side, lower jaw.

NHMUK OR (old BMNH) 30382: Quadrate, articular.

NHMUK OR (old BMNH) 30283-4: 2 smaller examples of the homologous bone.

NHMUK OR (old BMNH) 30384: Articular.

NHMUK OR (old BMNH) 30385: Angular.

NHMUK OR (old BMNH) 30388: Partial osteodemr.

NHMUK OR (old BMNH) 30389: Articular,ulna.

NHMUK OR (old BMNH) 30289: Partial premaxilla, partal nasal, patial jugal.

NHMUK OR (old BMNH) 30307-8: 2 right angular.

NHMUK OR (old BMNH) 30318: 2 dermal scutes.

NHMUK OR (old BMNH) 30338: Dentary.

NHMUK OR (old BMNH) 30354: 2 ischia.

NHMUK OR (old BMNH) 30359: Left coarcoid.

NHMUK OR (old BMNH) 30362: Ilium.

NHMUK OR (old BMNH) 30363: Left articular.

NHMUK OR (old BMNH) 30363a: 2 smaller examples of the homologous bone from the right side.

NHMUK OR (old BMNH) 30365: Dentary.

NHMUK OR (old BMNH) 30366-67: Dermal scutes.

NHMUK OR (old BMNH) 30369-70: 2 premaxilla.

NHMUK OR (old BMNH) 30372: 2 quadrates.

NHMUK OR (old BMNH) 30378: Dentary.

NHMUK OR (old BMNH) 30381: Partial rostrum fragment.

NHMUK OR (old BMNH) 30383: Dentary.

NHMUK OR (old BMNH) 30388: Articular joined with 30289>

NHMUK OR (old BMNH) 30392: Partial skull.

NHMUK OR (old BMNH) 30394, 30396: Mandibles.

NHMUK OR (old BMNH) 30397: Left ramus.

NHMUK OR (old BMNH) 30399: Femur.

NHMUK OR (old BMNH) 30392: Skull.

NHMUK OR (old BMNH) 30401: Radius.

NHMUK OR (old BMNH) 30402: Cervical and dorsal vertebrae.

NHMUK OR (old BMNH) 30414: Partial skull, partial premaxilla, quadrate, 2 fragments of a scapula.

NHMUK OR (old BMNH) 30414: Partial articular, partial surangular, several cervical ribs, partial ilium, partial femur, partial tibia.

NHMUK OR (old BMNH) 36812: Basioccipital.

NHMUK OR (old BMNH) 46434: Fragmentary dermal scutes.

RIO, MANNION, TSCHOPP, MARTIN & DELFINO, 2019

CAMSM C31041, C31042, C31043: Dentaries.

CAMSM C31044, 31045: Angualrs.

CAMSM C31-26-40: Cervical vertebrae.

CAMSM C31012-25: Dorsla vertebrae.

CAMSM C30999-31011: Caudal vertebrae.

CAMSM C31046-95, C32352-87: Osteoderms.

CAMSN unnumberd: 8 dentaries.

CAMSM TN 904: Lower jaw.

CAMSM TN 907: Partial skull, partial lower jaw.

CAMSM TN 918: Partial skull.

CAMSM TN917: Skull.

 

 

BENTON & SPENCER, 1995

Locality: Hordle Cliff, (SZ 253935-SZ 287915), Hampshire County, England, Southern United Kingdom.

Horizon: Totland Bay Member, Headon Hill Formation.

Biostratigraphy:

Age: Priabonian, Late Eocene Epoch, Paleogene Period, Lower Tertiary Subera, Lower Cenozoic Era.

Material:

CAMSM? Unnumbered

 

Locality: Bouldnor and Hamstead Cliffs, (SZ 391913), Isle of Wight County, England, Southern United Kingdom.

Horizon: Bouldnor Formation, Bembridge Marls Member, Hamstead Member, Rupelian.

Biostratigraphy:

Age: Lower Oligocene Epoch, Upper Paleogene Period, Upper Tertiary Subera, Middle Cenozoic.

Material:

Number: Not given:

 

= Diplocynodon cf. hantoniensis VIGNAUD, BRUNET, GUEVEL & JEHENNE, 1997

Locality: Razac d’Eymet, Dordogne, France.

Horizon:

Biostratigraphy:

Age: Lower Oligocene Epoch, Upper Paleogene Period, Upper Tertiary Subera, Middle Cenozoic.

Material:

Collection of Guevel: Skull.

 

WOLFE, FOWLER & BONDE, 2007

Locality: Bouldnor locality, Isle of Wight County, England, Southern United Kingdom.

Horizon: Bouldnor Formation, Bembridge Marls Member, Hamstead Member, Rupelian.

Biostratigraphy:

Age: Lower Oligocene Epoch, Upper Paleogene Period, Upper Tertiary Subera, Middle Cenozoic.

Material:

MSU-VP-Path-10-01: Osteoderm.

Note: Pathologic, necrotizing dermatitis.

 

= Crocodilus hastingsiae OWEN, 1847
Etymology: In reference to Hastings at Hordwell, Hampshire County, England, Southern United Kingdom.

Holotype: 30393

Locality: Hastings at Hordwell, Hampshire County, England, Southern United Kingdom.

Horizon: Headon Beds, Marchioness.

Biostratigraphy:

Age: Late Eocene Epoch, Paleogene Period, Lower Tertiary Subera, Lower Cenozoic Era.

Material: Skull, vertebrae, dorsal and ventral scutes.

Referred material:

OWEN, 1854

Locality: Hordwell, Hampshire County, England, Southern United Kingdom.

Horizon:

Biostratigraphy:

Age: Late Eocene Epoch, Paleogene Period, Lower Tertiary Subera, Lower Cenozoic Era.

Material:

ROYAL COLLAGE OF SURGEONS 9: Hind half of a left ramus.

ROYAL COLLAGE OF SURGEONS 10: Anterior part of a right ramus.

ROYAL COLLAGE OF SURGEONS 11: Right femur.

ROYAL COLLAGE OF SURGEONS 12: Right tibia.

ROYAL COLLAGE OF SURGEONS 13: 2 dermal scutes and a tooth.

 

= Alligator hantoniensis (WOOD, 1844) OWEN, 1847

Locality: Hastings at Hordwell, Hampshire County, England, Southern United Kingdom.

Horizon: Headon Beds, Marchionenss.

Biostratigraphy:

Age: Late Eocene Epoch, Paleogene Period, Lower Tertiary Subera, Lower Cenozoic Era.

Material:

Number: Not given: Skull and jaw specimens and vertebrae.

 

HUXLEY, 1859

Locality: Headon Beds, England, Southern United Kingdom.

Horizon:

Biostratigraphy:

Age:

Material:

Number: Not given: Dorsal and ventral scutes.

 

GRAMANN, 1958

Locality: Borken, Lower Hessen, Westfalen, North Rhine-Westphalia State, Germany.

Horizon:

Biostratigraphy:

Age: Lower Oligocene Epoch, Upper Paleogene Period, Upper Tertiary Subera, Middle Cenozoic.

Material:

Number: Not given: Anterior portion of a left mandibular ramus.

 

KUHN, 1938

Locality: Geisel Valley, Sachsen-Anhalt State, Germany.

Horizon:

Biostratigraphy:

Age: Middle Eocene Epoch, Paleogene Period, Lower Tertiary Subera, Lower Cenozoic Era.

Material:

Number: Not given: Imperfectly preserved skull and post-cranial material.

_____________________________________________________________________________________
Species: rateli (POMEL, 1846)
Etymology:
= Crocodilus ratelii POMEL, 1846

Holotype:

Locality: Allier, France.

Horizon:

Biostratigraphy:

Age: Lower Miocene Epoch, Lower Neogene Period, Upper Tertiary Subera, Middle Cenozoic Era.

Material: Right mandibular ramus.

Referred material:

BERG, 1966

Number: Not given: Skull and jaw material.

 

BUFFETAUT & CORNEE, 1982

Locality: Aquitanian of Venelles, Bouches-du-Rhone Department, France.

Horizon:

Biostratigraphy:

Age: Upper Oligocene Epoch, Upper Paleogene Period- Lower Miocene Epoch, Lower Neogene Period, Upper Tertiary Subera, Middle Cenozoic Era.

Material:

Number: Not given: Skull and most of skeleton mutilated by scavengers before burial.

 

= Diplocynodon cf. ratelii CHROUST, MAZUCH, IVANOV, EKRT & LUJAN, 2021

Locality: Ahnikov I, Most Basin, Czech Republic.

Horizon:

Biostratigraphy:

Age: Eggenburgian, MN3a, Early Miocene Epoch, Lower Neogene Period, Upper Tertiary Subera, Middle Cenozoic Era.

Material:

NMP PV 11603: Premaxilla.

NMP Pv 11604, 11605: Maxillae.

NMP Pv 11607-11609: Jugals.

NMP Pv 11611: Postorbital.

NMP Pv 11612: Squamosal.

NMP Pv 11614, 11615, 11609: Dentaries.

NMP Pv 11613: Basioccipital.

NMP Pv 11650- 11660: Teeth.

NMP Pv 11663-11665, 11667, 11668: Osteoderms.

NMP Pv 11627, 11621: Cervical vertebrae.

NMP Pv 11622, 11624, 11625, 11626: Dorsal trunk vertebrae.

NMP Pv 11623: Caudal vertebra.

NMP PV 11629: Partial humerus.

NMP Pv 11630: Partial femur.

NMP Pv 11631: Partial metatarsal.

 

Locality: Ahnikov II, Most Basin, Czech Republic.

Horizon:

Biostratigraphy:

Age: Eggenburgian, MN3a, Early Miocene Epoch, Lower Neogene Period, Upper Tertiary Subera, Middle Cenozoic Era.

Material:

NMP Pv 11632: Partial maxilla and dentary.

NMP Pv 11635-11637, 11649: Frontals.

NMP Pv 11639: Squamosal.

NMP Pv 11661, 11662: Teeth.

NMP Pv 11644-11666: Osteoderms.

NMP Pv 11628: Dorsal trunk vertebra.

NP Pv 11634: Partial femur.

 

Locality: VASSEUR, 1912

Locality: Agenais, South-west France.

Horizon:

Biostratigraphy:

Age: Upper Aquitanian, Lowermost Miocene Epoch, Lower Neogene Period, Upper Tertiary Subera, Middle Cenozoic Era.

Material:

Number: Not given: Teeth and dermal scutes.

 

= Diplocynodon gracile VALLANT, 1872
Etymology: Latin, gracilis, "slender."

Holotype:

Locality: NNE Clermont-Ferrand bzw. ca. 15 km NNE Vichy/Allier, France.

Horizon:

Biostratigraphy:

Age:

Material: Skull.

Referred material:

LYDEKKER, 1888

Locality: St. Gerand-le-Puy (Allier), France.

Horizon:

Horizon:

Biostratigraphy:

Age: Early Miocene or Late Oligocene Epoch, Upper Paleogene Period, Upper Tertiary Subera, Middle Cenozoic.

Material:

BMNH R789: Skull and numerous scutes and bones of a small individual.

BMNH 28641: Posterior portion of a skull.

The following specimens earthier belong to D. gracilis or D. darwini.

BMNH 26748: Posterior portion of a skull.

BMNH 28641: Left maxilla of a juvenile.

BMNH 28645: Right maxilla of a juvenile.

BMNH 26846: Tooth.

BMNH 30959: 2 teeth.

BMNH 30949, 30945-6: 3 fragments of skull.

BMNH 26841a-46: 2 fragments of skull.

BMNH 26840: Left ramus.

BMNH 26842: Fragmentary left mandible of a juvenile.

BMNH 26842: Left articular.

BMNH 26846: Dorsal vertebra.

BMNH 48471: Right humerus.

BMNH 30963: Left humerus of a juvenile.

BMNH 26846: 2 fragments of a humerus.

BMNH 26846a: Left ilium and pubis.

BMNH 30965: 8 dermal scutes.

 

Locality: Antiogin, Puy-de-Dome Department, France.

Horizon:

Biostratigraphy:

Age: Lower Miocene Epoch, Lower Neogene Period, Upper Tertiary Subera, Middle Cenozoic Era.

Material:

BMNH 27751: Fragmentary skull.

BMNH 27719: Posterior portion of a skull.

 


Locality: Cournon, Puy-de-Dome Department, France.

Horizon:

Biostratigraphy:

Age: Lower Miocene Epoch, Lower Neogene Period, Upper Tertiary Subera, Middle Cenozoic Era.

Material:

BMNH 27676: Fragmentary humerus, rib, vertebra and other bones.

 


Locality: Rott, near Bonn, Westfalen, North Rhine-Westphalia State, Germany.

Horizon:

Biostratigraphy:

Age: Lower Miocene Epoch, Lower Neogene Period, Upper Tertiary Subera, Middle Cenozoic Era.

Material:

BMNH 41089: Impression of a skull.

 

Locality: Caylux (Tarn-et-Garonne), France.

Horizon: Phosphorites.

Biostratigraphy:

Age: Lower Miocene Epoch, Lower Neogene Period, Upper Tertiary Subera, Middle Cenozoic Era.

Material:

BMNH R381: Anterior portion of a right ramus.

BMNH R429: Tooth.

BMNH R488: Fragmentary dorsal vertebra.

BMNH R382: Right humerus.

 

= Crocodilus bruchi MEYER, 1843
Etymology:
= Alligator bruchi (MEYER, 1843)
= Crocodilus medius MEYER, 1843
Etymology: Laitn, medius, "middle."
= Crocodilus brauniorum MEYER, 1843
Etymology:

Holotype:

Locality:

Horizon:

Biostratigraphy:

Age:

Material: Skull and jaw fragments.

 

= Crocodilus elaverensis BRAVARD, 1844
Etymology:

Holotype:

Locality: Bournoncle-Saint-Pierre, France.

Horizon: Limagne Basin.

Biostratigraphy:

Age:

Material: Scute.

Referred material:

Locality: Antoin, France.

Horizon:

Biostratigraphy:

Age:

Material:

BMNH 41089: Scute.

 

= Orthosaurus elaverensis (BRAVARD, 1844) GERVAIS, 1859
Etymology:

Holotype:

Locality: Saint-Gerand le Puy, France.

Horizon:

Biostratigraphy:

Age:

Material: Imperfect skull.

 

= Crocodilus aeduicus VALLANT, 1872
Etymology:

Holotype:

Locality: Saint-Gerand le Puy, France.

Horizon:

Biostratigraphy:

Age: Aquitanian, Miocene Epoch, Lower Neogene Period, Upper Tertiary Subera, Middle Cenozoic Era.

Material: Skull.

 

= Crocodilus rahtii LUDWING, 1877
Etymology:

Holotype:

Locality:

Horizon:

Biostratigraphy:

Age:

Material:

 

= Diplocynodon gracilis aeduicus BERG, 1966
Etymology:

Holotype:

Locality: Mainz Basin, Germany.

Horizon:

Biostratigraphy:

Age:

Material: Skull and jaw fragments.

 

= Diplocynodon rateli gracilis BERG, 1966
Etymology:

Holotype:

Locality: Orleans, Loiret, France.

Horizon:

Biostratigraphy:

Age:

Material: Mandible fragment.

_____________________________________________________________________________________
Species: gervaisi (GERVAIS, 1852)
Etymology:
= Crocodilus gervaisi GERVAIS, 1852

Holotype:

Locality: Le Puy, Haute Loire, France.

Horizon:

Biostratigraphy:

Age:

Material: Skull.

Referred material:

= Saurocainus gervaisi (GERVAIS, 1852)
Etymology:

Holotype:

Locality: Haute-Loire Department, France.

Horizon:

Biostratigraphy:

Age:

Material: No material was described.

 

= Crocodilus (Saurocainus) gervaisi (GERVAIS, 1852)
Etymology:

Holotype:

Locality: Ronzon, Haute-Loire Department, France.

Horizon:

Biostratigraphy:

Age:

Material: Lower jaw fragment.

Referred material:


BERG, 1966

Locality: Ronzon, Haute-Loire Department, France.

Horizon:

Biostratigraphy:

Age:

Material:

Number: Not given: Lower Jaw.

 

ZITTEL, 1890

Locality: Ronzon, Haute-Loire Department, France.

Horizon:

Biostratigraphy:

Age:

Material:

Number: Not given: Imperfect Skull.

 

= Saurocainus latus AYMARD, 1856
Etymology: Latin, latus, "wide."

Holotype:

Locality: Haute-Loire Department, France.

Horizon:

Biostratigraphy:

Age:

Material: No material was described.

_____________________________________________________________________________________

Species: buetikoensis (MEYER, 1854)
Etymology:
= Crocodilus buetikoensis MEYER, 1854

Holotype:

Locality: Buttikon, near Zurich, Zurich Canton, Switzerland.

Horizon:

Biostratigraphy:

Age: Tertiary Subera, Middle Cenozoic Era.

Material: Skull and lower jaw with associated dermal scutes.

Referred material:

SCHERER, 1978

Locality: The fissure filling of Appertshofen in the Southern Franconian Alb northerly of Ingolstadt, Bayern (Bavaria) State, Germany.

Horizon:

Biostratigraphy:

Age: Late Miocene Epoch, Lower Neogene Period, Upper Tertiary Subera, Middle Cenozoic Era.

Material:

BSP 1953 II7-309: Exoccipital, left premaxilla, fragment of left nasal, fragment of right prefrontal, right lachrymal, fragment of right quadrate, fragment of the anterior end right surangular, fragment of right symphysal, fragment of right dentary and teeth.

 

SCHERER, 1979

Locality: Viehhausen near Regensburg, Bayern (Bavaria) State, Germany.

Horizon:

Biostratigraphy:

Age: Miocene Epoch, Lower Neogene Period, Upper Tertiary Subera, Middle Cenozoic Era.

Material:

Palaontologisches Institut der Universitat Wurzburg Nr. 101/369: Fragmentary skull.

_____________________________________________________________________________________

Species: gracilis VAILLANT, 1872
Etymology: Latin, gracilis, "slender."

Holotype:

Locality:

Horizon:

Biostratigraphy:

Age:

Material:

Referred material:

Locality: Elgg, near Winterthur, Switzerland.

Horizon: Brown Coal.

Biostratigraphy:

Age: Eocene Epoch, Paleogene Period, Lower Tertiary Subera, Lower Cenozoic Era.

Material:

Number: Not given: Dermal scute.

_____________________________________________________________________________________

Species: darwini (LUDWING, 1877)
Etymology: In honor of Naturalist Charles R. Darwin (1809-1882).
= Alligator darwini LUDWING, 1877

Holotype: Me 5360

Locality: Messel near Darmstadt, Odenwald, Hesse State, Germany.

Horizon:

Biostratigraphy:

Age: Middle Eocene Epoch, Paleogene Period, Lower Tertiary Subera, Lower Cenozoic Era.

Material: Skull.

Referred material:

Me 4281: Fragmentary lower jaw.

Me 4319: Fragmentary skull.

Me 5347: Premaxilla, maxilla, nasals, and palatine fragment.

Me 5361: Fragment of lower jaw.

Me 5364: Skull.

Me 5374: Fragment of right lower jaw.

Me 5639: Fragment of lower jaw.

Me 5643: Fragmentary skull.

Me 5648: Deformed skull.

Me 5660: Lower jaw.

 


Locality and Horizon:?

Biostratigraphy:

Age:

Material:

SMF, Me 896: Sacral and caudal vertebrae, femur, pelvis and scutes.

 

= cf. Diplocynodon darwini HELLMUND, 2001

Locality: From former open pit of Mucheln-Westfield, Geiseltal, Sachsen-Anhalt State, Germany.

Horizon:

Biostratigraphy:

Age: Middle Eocene.

Material:

Number: Not given: Fragmentary specimen.
Note: With gastroliths.

 

LIEBIG, FREY, THEOBALD & WILLIS, 1996

Locality: Messel near Darmstadt, Odenwald, Hesse State, Germany.

Horizon: Lutetium.

Biostratigraphy:

Age: Middle Eocene Epoch, Paleogene Period, Lower Tertiary Subera, Lower Cenozoic Era.

Material:

HLMD Me 5478: Fragmentary skull with skin impressions from the mouth.

 

REDLICH, 1902

Locality: Bohemia.

Horizon: Brown Coal.

Biostratigraphy:

Age: Eocene Epoch, Paleogene Period, Lower Tertiary Subera, Lower Cenozoic Era.

Material:

Number: Not given: Skull and jaw fragments, teeth, dermal scutes, etc.

= Diplocynodon ebertsi (LUDWING, 1877)
Etymology:
= Crocodilus ebertsi LUDWING, 1877
= Caiman ebertsi (LUDWING, 1877)

Lectotype: Me 5366

Locality: Darmstadt, Hesse State, Germany.

Horizon:

Biostratigraphy:

Age: Middle Eocene Epoch, Paleogene Period, Lower Tertiary Subera, Lower Cenozoic Era.

Material: Dentary and right mandibula.

Referred material:

Me 5341: Fragment of a skull of a juvenile.

Me 5362: Fragmentary skull.

Me 5365: Fragment of a premaxilla, maxilla, and right nasal.

Me 5638: Fragment of rostrum.

 

REDLICH, 1902

Locality: Bohemia, Czech Republic.

Horizon: Brown Coal.

Biostratigraphy:

Age: Eocene Epoch, Paleogene Period, Lower Tertiary Subera, Lower Cenozoic Era.

Material:

Number: Not given: 3 dorsal vertebrae, 1st sacral, a left metatarsal I, a tooth and a rib fragment.

= Diplocynodon hallensis KUHN, 1938
Etymology:

Holotype:

Locality: Geisel Valley, near Halle, Sachsen-Anhalt State, Germany.

Horizon:

Biostratigraphy:

Age: Middle Eocene Epoch, Paleogene Period, Lower Tertiary Subera, Lower Cenozoic Era.

Material: Skull and imperfect skeleton.

Referred material:

BERG, 1967

Locality: Geissel Valley, Sachsen-Anhalt State, Germany.

Horizon:

Biostratigraphy:

Age: Middle Eocene Epoch, Paleogene Period, Lower Tertiary Subera, Lower Cenozoic Era.

Material:

Number: Not given:

_____________________________________________________________________________________

 

Species: muelleri (KALIN, 1936) PIRAS & BUSCALIONI, 2006
= Hispanochampsa mülleri KALIN, 1936
Etymology:

Holotype: NMB Spa 4 (= "T" 1)

Locality: El Talladell, Tarrega, Catalunya, Lleida Province, Spain.

Horizon: Calcaries de Tarrega Formation.

Biostratigraphy: Theridomyx major zone.

Age: Lower Oligocene Epoch, Upper Paleogene Period, Upper Tertiary Subera, Middle Cenozoic.

Material: Skull.

Referred material:

"T" 2: Skull.

PIRAS & BUSCALIONI, 2006

NMB, Spa 4, Spa 73, IGME 1565N, 1567N, IEI 74, MMB 28128, MCT 1531, 1528:

 

MACALUSO, MARTIN, del FAVERO & DELFINO, 2019

Locality: Monteviale, Vicenza Province, Italy.

Horizon:

Biostratigraphy:

Age: Late Rupelina age, Lower Oligocene Epoch, Upper Paleogene Period, Upper Tertiary Subera, Middle Cenozoic.

Material:

MGP-PG 10170, 11407, 28164, 26816, 26817 (skull and lower jaw of same individual), 26833, 26845, 26846, 26858: Skulls.

MGP-PD 26845, 26850, 26862: 3 rostri.

MGP-PB 8648, 26819, 26835, 26837, 26838, 26842, 26957, 26859, 31998: 9 isolated skull fragments.

MGP-PD 26839: Several disarticulated teeth.

MGP-PD 27999-26854 (a single fractures slab), 26818, 26820, 26831, 26822, 26823, 26824, 26825-26830 (a single fractured slab), 26826, 26827, 26828, 26829, 26831-26834 (derived from the same individual, 26836, 26840, 26841, 26847, 26848, 26849, 26851, 26852-26853 (a single fractured slab), 26855, 26856, 31997: Various postcranial elements.

MGP-BD 27403: Almost complete specimen.

Note: Lable states 'Monte Bolca,' but more likely than not comes from Monteviale.

NMB-Bc.6: Anterior half of skeleton with a complete skull with bones belonging to one or two smaller individuals.

NHMUK 2789: A slab particularly altered by pyrite oxidation containing the thoracic portion of a small individual and part of tis limbs.

MGP-PD unnumbered: 7 fragments of mandibles and maxillae, more than 40 disartticulated teeth, 5 slabs with poorly preserved portions of the vertebral column, 5 isolated vertebrae, 7 slabs with fragmentary adn disarticulated limbs adn several ribs and osteoderms.

 

 

= Diplocynodon monsvialensis (FABIANI, 1914)
Etymology: In reference to Monteviale, Vicenza Province, Italy.
= Crocodilus monsvialensis FABIANI, 1914
= Asiatosuchus monsvialensis (FABIANI, 1914) FRANCO & PICCOLI, 1993

Lectotype: MGP-PD 26843

Locality: Monteviale, Vicenza Province, Italy.

Horizon:

Biostratigraphy:

Age: Late Rupelina age, Lower Oligocene Epoch, Upper Paleogene Period, Upper Tertiary Subera, Middle Cenozoic.

Material:

 

= Crocodilus dalpiazi FABIANI, 1915
Etymology:
= Diplocynodon dalpiazi (FABIANI, 1915) FRANCO, PICCOLI & TCHAPRASSIAN, 1992

Lectotype: MGP-PD 26814

Locality: Monteviale, Vicenza Province, Italy.

Horizon:

Biostratigraphy:

Age: Late Rupelina age, Lower Oligocene Epoch, Upper Paleogene Period, Upper Tertiary Subera, Middle Cenozoic.

Material:


= Diplocynodon marini BATALLER, 1941
Etymology:

Holotype:

Locality: Tarrega, Catalunya, Spain.

Horizon:

Biostratigraphy:

Age: Oligocene Epoch, Upper Paleogene Period, Upper Tertiary Subera, Middle Cenozoic.

Material: Incomplete skull and skeleton.

= Diplocynodon guerini BATALLER, 1941
Etymology:

Holotype:

Locality: Spain.

Horizon:

Biostratigraphy:

Age: Oligocene Epoch, Upper Paleogene Period, Upper Tertiary Subera, Middle Cenozoic.

Material:
_____________________________________________________________________________________

Species: levantinicum HUENE & NIKOLOFF, 1963
Etymology:

Holotype: No holotype listed.

Lectotype: NMNHS 31-1 (as per Massonne & Böhme, 2022).

Locality: Kipra coal-seam of teh underground mines Nadeshda, West-Mariza lignite field, Open Graben, north of the city of Dimitrovgrad, 140 km SE of Sofia, Central Bulgaria.

Horizon:

Biostratigraphy:

Age: Late Oligocene Epoch, Upper Paleogene Period, Upper Tertiary Subera, Middle Cenozoic.

Material: Cast of the right dentary fragment from teh first to the 14th alveolous.

Referred material:

NMNHS FR 31-2: Cast of right anterior dentary fragment.

 

Locality: Radievo 1, West-Mariza lignite field, Open Graben, north of the city of Dimitrovgrad, 140 km SE of Sofia, Central Bulgaria.

Horizon:

Biostratigraphy:

Age: Late Oligocene Epoch, Upper Paleogene Period, Upper Tertiary Subera, Middle Cenozoic.

Material:

NMNHS FR 30-1: Left premaxilla fragment.

NMNHS FR 30-2: Left dentary fragment.

NMNHS FR 30-3: Dentary fragment.

NMNHS FR 30-4: Right dentary fragment.

NMNHS FR 30-5: Left surangualr fragment.

NMNHS FR 30-6: Left ventral angular fragment.

NMNHS FR 30-7: Left posterior antular fragment.

NMNHS FR 30-8: Left articualr fragment.

NMNHS FR 30-9: Right articular fragment.

NMNHS FR 30-10: Cervical vertebra fragment.

NMNHS FR 30-11: Caudal vertebral fragment.

NHMHS FR 30-12: Right coracoid fragment.

NMNHS FR 30-13: Right ulan fragment.

NMNHS FR 30-14: Metatarsl fragment.

NMNHS FR 30-15: Phalane.

NMNHS FR 30-16 to 18: Dorsal osteoderms.

NMNHS FR 30-19, 20: Ventral osteodersm.

NHMHS FR 30-21: Left posterior maxilla fragment.

NMNHS FR 30-22: Left jugal frament, matching the maxilla fragment.

NMNHS FR 30-23: Right anterior dentary fragment.

NMNHS FR 30-24: Tooth.

NMNHS FR 30-25: Tooth.

NMNHS FR 30-16: Tooth fragment.
_____________________________________________________________________________________

Species: deponiae (FREY, LAEMNERTI & RIESS, 1987) DELFINO & SMITH, 2012
Etymology:
= Baryphracta deponiae FREY, LAEMNERTI & RIESS, 1987

Holotype: SMF, Me 899

Locality: Messel near Darmstadt, Odenwald, Hesse State, Germany.

Horizon: Lutetium.

Biostratigraphy:

Age: Eocene Epoch, Paleogene Period, Lower Tertiary Subera, Lower Cenozoic Era.

Material: Skull and skeleton.

Referred material:

IRSNB R 161: Nearly complete skull and skeleton.

_____________________________________________________________________________________

Species: tormis BUSCALIONI, SANZ, & CASANOVAS, 1992
Etymology:

Holotype: IPS-9001

Locality: Buero basin, province of Salamanca, Spain.

Horizon: Upper Rhenanian of Salamaca sector.

Biostratigraphy:

Age: Eocene Epoch, Paleogene Period, Lower Tertiary Subera, Lower Cenozoic Era.

Material: Skull with mandible with lacks the occiput.

Referred material:

ST-7066: Partially disarticulated skull lacking the premaxillae.

ST-344: Fragment of cranial table with occiput (previously figured and identified as Diplocynodon aff. gracilis by Jimenex-Fuentes 1983)

ST-7320 and ST-7505: Isolated mandibular elements.

_____________________________________________________________________________________

Species: elavericus MARTIN, 2010
Etymology: ‘elavericus’ referes to the Latin name of Allier River: Elaver. Note that Bravard (1843) used ‘Crocodilius elaveris’ to designate various crocodilian remains from Allier but did not provide any description, nor did he figure any specimens. As Vaillant (1872) pointed out, this early mention cannot be considered as having any right of priority over Diplocynoond ratelii, given the absence of a description or figure. ‘Crocodilus elaveris’ should be considered as a nomen nudum.

Holotype: Rhinopolis B3

Locality: Domerat, Allier Department, France.

Horizon:

Biostratigraphy:

Age: Middle Priabonian, Late Eocene Epoch, Paleogene Period, Lower Tertiary Subera, Lower Cenozoic Era.

Material: Complete skull and mandibular fragments and some osteoderms.

_____________________________________________________________________________________

Species: kochi VENCZEL & CODREA, 2022
Etymology: A tribute to Koch Antal, eminent geologist from Transylvania, Cluj-Napoca (Kolozsvar, Klausenburg), who brought an important contribution to the field of Transylvanian stratigraphy:

Holotype: UBB V.1453

Locality: Cluj-Manastur former limestone quarry, Cluj-Nopoca, Transylvania, Romania.

Horizon: Cluj Limestone Formation.

Biostratigraphy:

Age: Priabonian Stage, Late Eocene Epoch, Late Middle Paleogene Period, Lower Tertiary Subera, Lower Cenozoic Era.

Material: Nearly complete skull lacking from its right side the anterolateal margin of premaxilla, posteriormost part of the maxilla, anterior part of the jugal and from the left side the postorbital, squamosal, jugal and quadratojugal.

Referred material:

VENCZEL, 2023

Locality: Limstone Quarry, Southwestern vicinity of Leghia Village, Gilau sedemntary area, Cluj County, Transylvanian Basin, Romania.

Horizon: Vistea Limestone Formation (or upper coarse limestone Koch, 1894).

Biostratigraphy:

Age: Priabonian Stage, Late Eocene Epoch, Late Middle Paleogene Period, Lower Tertiary Subera, Lower Cenozoic Era.

Material:

UBB V.1004/1: Paired dentaries.

UBB V. 1004/2: Posterior part of right mandible.

UBB V. 1008: Isolated tooth.

UBB V.1007: Trunk vertebra.

UBB V.1005, 1006: Femora.

 

Locality: Santului Valley near the locality of Trezna, Meses sedimentray area, Salaj County, Transylvanian Basin, Romania.

Horizon: Turbuta Formation (may corresopne to the Valea Nadasului Formation, Farcas, 2011).

Biostratigraphy:

Age: Priabonian Stage, Late Eocene Epoch, Late Middle Paleogene Period, Lower Tertiary Subera, Lower Cenozoic Era.

Material:

MTC 26015: Left jugal.

MTC 26016: Isolated dentary.

UBB B. uncatalogued: 3 isolated teeth.

 

Locality: Within the village of Radaia, Gilau sedimnetary area, Cluj County, Transylvanian Basin, Romania.

Horizon: Valea Nadasului Formation.

Biostratigraphy:

Age: Priabonian Stage, Late Eocene Epoch, Late Middle Paleogene Period, Lower Tertiary Subera, Lower Cenozoic Era.

Material:

UBB V. 1449: Partial left jugal.