Species: neocesariensis (DE KAY, 1842) LEIDY, 1852
Etymology:
= Gavialis neocesariensis DE KAY, 1842

Holotype: ANSP (?) 9437

Locality: Vincentown, Big Timber Creek, Burlington County, New Jersey.

Horizon: Vincentown Limestone, Rancocas Group.

Biostratigraphy:

Age: Late Maastrichtian Stage, uppermost Senonian subepoch, Gulf Epoch, Late Cretaceous.

Material: Portion of left dentary, portion of right dentary.

Referred material:

BAIRD, 1986

Locality: Loc. B, Landover, southeast of intersection of Interstate Route I-495 (Capital Beltway) and MD Route 202 (Landover road), McCormic Construction site, Developed as Inglewood Business Center
(southeast of Brouwers & Hazel'z loc.2), Prince Georges County, Maryland.

Horizon: Severn Formation, Middle Maastrichtian, Late Cretaceous.

Material:

USNM 336478: Teeth and osteoscute fragments.

USNM 387368: 2 teeth.

Fonger Collection: Fragmentary vertebrae.

 

Locality: Loc. C. Excavations for and exposures behind Hampton Mall Shopping Center, southwest of intersection of Route I-495 and Central Avenue (= Loc. 3a of Brouwers & Hazel, Prince Georges County, Maryland.

Horizon: Severn Formation.

Biostratigraphy:

Age: Early Late Maastrichtian Stage, uppermost Senonian subepoch, Gulf Epoch, Late Cretaceous.

Material:

Number: Not given: Teeth and scutes.

 

Locality: Loc. D. North side of US route 50 about 2 to 3 miles east of intersection with Route I-495, Prince Georges County, Maryland.

Horizon: Severn Formation.

Biostratigraphy:

Age: Early Late Maastrichtian Stage, uppermost Senonian subepoch, Gulf Epoch, Late Cretaceous.

Material:

Number: Not given: Teeth and scutes.

 

ESTES & BERBERIN, 1970

Locality: Bug Creek Anthills, SW 1/4, Sec. 28, T24N, R44E, McCone County, Montana.

Horizon: Hell Creek Formation.

Biostratigraphy:

Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.

Material:

Number: Not given: Teeth.

 

= Thoracosaurus cf. neocesariensis GALLEGHER, 1993

Locality: Blackwood Terrace, Gloucester County, New Jersey.

Horizon: MFL, Basal Hornerstown Formation, Greensand, Rancocas Group.

Biostratigraphy:

Age: Late Maastrichtian Stage, uppermost Senonian subepoch, Gulf Epoch, Late Cretaceous.

Material:

ANSP unnumbered: Lower jaw.

 

GALLEGHER, 1993

Locality: MFL, Inversand Pit, Sewell, Gloucster County, New Jersey.

Horizon: Basal Hornerstown Formation, Greensand, Rancocas Group.

Biostratigraphy:

Age: Late Maastrichtian Stage, uppermost Senonian subepoch, Gulf Epoch, Late Cretaceous.

Material:

NJSM 11885:

Locality: Landover, Prince Georges County, Maryland.

Horizon: Severn Formation.

Biostratigraphy:

Age: Early Late Maastrichtian Stage, uppermost Senonian subepoch, Gulf Epoch, Late Cretaceous.

Material:

Private collections: Teeth, scutes, and fragmentary vertebrae.

 

Locality: Blackwood, Camdon County, New Jersey.

Horizon: Vincentown Limestone, Rancocas Group.

Biostratigraphy:

Age: Late Maastrichtian Stage, uppermost Senonian subepoch, Gulf Epoch, Late Cretaceous.

Material:

ANSP 9364:

 

Locality: Mt. Holly, Burlington County, New Jersey.

Horizon: Navesink Formation, Monmouth Group.

Biostratigraphy:

Age: Middle Maastrichtian Stage, uppermost Senonian subepoch, Gulf Epoch, Late Cretaceous.

Material:

ANSP 9435, 9358:

 

Locality: Vincentown, Burlington County, New Jersey.

Horizon: Hornerstown Formation (?), Rancocas Group.

Biostratigraphy:

Age: Late Maastrichtian Stage, uppermost Senonian subepoch, Gulf Epoch, Late Cretaceous.

Material:

ANSP 10079:

 

HARTSTEIN, DECINA & KEIL, 1999

Locality: Near the town of Bowie, 76°42’30”, 38°57’50:, Prince Georges County, Maryland.

Horizon: Severn Formation.

Biostratigraphy:

Age: Early Late Maastrichtian Stage, uppermost Senonian subepoch, Gulf Epoch, Late Cretaceous.

Material:

Decina collection: Tooth.

McCloskey collection: Scute.

 

LAURENT, BUFFETAUT & LE LOEUFF, 2000

Locality: Cassagnau 2, foothills of the Pyrenees near the village of Marignac-Laspeyres, Haute-Garonne Department, France.

Horizon: Marnes d’Auzas Formation.

Biostratigraphy:

Age: Maastrichtian Stage, uppermost Senonian subepoch, Gulf Epoch, Late Cretaceous.

Material:

MDE-Cas1-199: Nearly complete skull.

 

LEIDY, 1864

Locality: Vincentown, Burlington County, New Jersey.

Horizon: Hornerstown Formation, Rancocas Group.

Biostratigraphy:

Age: Late Maastrichtian Stage, uppermost Senonian subepoch, Gulf Epoch, Late Cretaceous.

Material:

Number: Not given: Fragmentary skull, teeth.

LYDEKKER, 1888

BMNH 41842: Cast of a skull figured by LEIDY 1864.

 

Locality: Blackwood town, Camden County, New Jersey.

Horizon:

Biostratigraphy:

Age: Late Cretaceous.

Material:

Number: Not given: Teeth, fragmentary upper jaw.

 

Locality: Big Timber Creek, Gloucester County, New Jersey.

Horizon:

Biostratigraphy:

Age: Late Cretaceous.

Material:

Number: Not given: Tooth.

 

Locality: Mount Holly, Burlington County, New Jersey.

Horizon: Greensand.

Biostratigraphy:

Age: Late Cretaceous.

Material:

Number: Not given: A dermal plate.

 

Locality: Burlington County, New Jersey.

Horizon:

Biostratigraphy:

Age: Late Cretaceous.

Material:

Number: Not given: Vertebrae.

 

MANNING & DOCKERY, 1992

Locality: Frankstown site, Prentiss County, Mississippi.

Horizon: Basal Demopolis Formation.

Biostratigraphy:

Age: Late Campanian Stage, Senonian subepoch, Upper Gulf epoch, Late Cretaceous.

Material:

MPPM 1990.47.2: Partial osteoscute.

MPPM 1991.27.2: Tooth crown.

 

MILLER, 1955

Locality: Big Timber Creek, Gloucester County, New Jersey.

Horizon: Vincentown Formation.

Biostratigraphy:

Age: Eocene Epoch, Paleogene Period, Lower Tertiary Subera, Lower Cenozoic Era.

Material:

ANSP 9374, 9381:

 

= Thoracosaurus sp. cf. T. neocesariensis MULDER, 1997, 1998

Locality: Ankerpoort-Nekami quarry near Bemelen, Limburg Province, the Netherlands.

Horizon: Probably Kekum Member of the Meerssen Member, Maastricht Formation.

Biostratigraphy: Belemnitella junior Zone or Belemnella casimirovensis Zone.

Age: Late Maastrichtian Stage, Upper Senonian Subepoch, Upper Gulf Epoch, Late Cretaceous.

Material:

Museum Natura Docet Denekamp Collection, No. K 20.01.882: Cervical vertebra.

K. 20.01.883, K. 20.01.885: Dorsal vertebrae.

 

SCHEIN, LACOVARA, GALLAGHER & POOLE, 2006

Locality: New Jersey.

Horizon: Hornerstone Formation.

Biostratigraphy:

Age: Late Maastrichtian Stage, uppermost Senonian subepoch, Gulf Epoch, Late Cretaceous-Danian Stage, Paleocene Epoch, Lower Paleogene Period- Early Eocene Epoch, Paleogene Period, Lower Tertiary Subera, Lower Cenozoic Era.

Material:

Number: Not given: 30 isolated teeth, a partial premaxilla, the caudal portion of left dentary, a partial surangular, the dorsocaudal region of the skull including both frontals and portions of the postorbitals, parietals, suamosals, brain case including the basioccipital, basipterygoid process, exoccipital, partial foramen magnum, and a partial parocciptial process, atlas, axis, partial left ilium, partial ischium, 2 partial humeri, a scapula and coraocoid, 9 cervical vertebrae, 1 complete and numerous partial cervical ribs, 12 dorsal vertebrae with numerous partial ribs, 1 sacral vertebra, a partial pubis, 9 caudal vertebrae, femora, astragalus and partial calcaneum, 2 complete phalnges, a partial metacarpal and approximately 30 osteoderms.

 

= Crocodilus dekayi LEIDY, 1852
Etymology:
= Thoracosaurus (Crocodiles) dekayi (LEIDY, 1852) MOOK, 1925

Holotype: Number not given.

Locality: Southern New Jersey.

Horizon: Greensands.

Biostratigraphy:

Age: Late Cretaceous.

Material: Parts of a jaw.

 

= Crocodilus (Gavialis?) clavirostris MORTON, 1844
Etymology:

Holotype: ANSP 10079

Locality: Farm of William Irick, near Vincentown, Burlington County, New Jersey.

Horizon:

Biostratigraphy:

Age: Late Cretaceous.

Material: Skull lacking anterior tip.

 

= Sphenosaurus clavirostris AGASSIZ, 1849
Etymology:

Holotype: Number not given.

Locality: Near Vincentown, Burlington County, New Jersey.

Horizon: Ferruginous marl.

Biostratigraphy:

Age: Late Cretaceous.

Material: Skull missing anterior tip.

 

= Crocodilus basitruncatus OWEN, 1849
Etymology:
= Holops basitruncatus (OWEN, 1849)
= Holopsisuchus basitruncatus (OWEN, 1849)

Holotype: Number not given.

Locality: Big Timber Creek, Gloucester County, New Jersey.

Horizon: Vincentown Limestone, Rancocas Group.

Biostratigraphy:

Age: Late Maastrichtian Stage, uppermost Senonian subepoch, Gulf Epoch, Late Cretaceous.

Material: Cervical vertebra.

Referred material:

OWEN, 1954

Locality: New Jersey.

Horizon:

Biostratigraphy:

Age: Late Cretaceous.

Material:

ROYAL COLLEGE OF SURGEONS 17: 3rd cervical vertebra.

ROYAL COLLEGE OF SURGEONS 18: 4th cervical.

ROYAL COLLEGE OF SURGEONS 19: 5th cervical.

ROYAL COLLEGE OF SURGEONS 20: Fragmentary dorsal vertebra.

 

MILLER, 1955

Locality: Big Timber Creek, Gloucester County, New Jersey.

Horizon: Vincentown Formation.

Biostratigraphy:

Age: Eocene Epoch, Paleogene Period, Lower Tertiary Subera, Lower Cenozoic Era.

Material:

ANSP 9436, 9169, 9383, 9355-56:

 

= Crocodilus basifissus OWEN, 1849 (nomen dubium)
Etymology:
= Thoracosaurus basifissus (OWEN, 1849) (nomen dubium)

Holotype: Number not given.

Locality: Vincentown, Burlington County, New Jersey.

Horizon: Vincentown Limestone, Rancocas Group.

Biostratigraphy:

Age: Late Maastrichtian Stage, uppermost Senonian subepoch, Gulf Epoch, Late Cretaceous.

Material: Cervical vertebrae, fragmentary vertebrae.

Referred material:

OWEN, 1854

Locality: New Jersey.

Horizon:

Biostratigraphy:

Age: Late Cretaceous.

Material:

ROYAL COLLEGE OF SURGEONS 21: 3rd or 4th cervical.

ROYAL COLLEGE OF SURGEONS 22: 6th cervical.

ROYAL COLLEGE OF SURGEONS 23: Posterior cervical vertebrae, 7th?

 

= Thoracosaurus grandis LEIDY, 1865
Etymology: Latin, grandis, "large, full grown."

Syntypes: ANSP 9358, 9435

Locality: Mount Holly, Burlington County, New Jersey.

Horizon: Navesink Formation, Monmouth Group.

Biostratigraphy:

Age: Middle Maastrichtian Stage, uppermost Senonian subepoch, Gulf Epoch, Late Cretaceous.

Material: A dermal scute and half of another.

 

= Crocodilus tenebrosus LEIDY, 1865
Etymology:
= Holopsisuchus tenebrosus (LEIDY, 1865)
= Holops tenebrosus (LEIDY, 1865)

Holotype: ANSP 9161, 9162, 9169, 9355, 9356, 9383, 9436

Locality: Big Timber Creek, Gloucester County, New Jersey.

Horizon: Vincentown Limestone, Rancocas Group.

Biostratigraphy:

Age: Late Maastrichtian Stage, uppermost Senonian subepoch, Gulf Epoch, Late Cretaceous.

Material: Portions of two humeri and vertebrae, two cervical, a dorsal, the sacral, and 2 caudal vertebrae and portions of both humeri.

Note: See SPAMER & DASCHLER, 1995 for figures.

 

= Holops obscurus (LEIDY, 1865)
Etymology:
= Holopsisuchus obscurus LEIDY, 1865

Holotype: RU 1-K-V.-12 (Now at the AMNH)

Locality: Barnesboro, Gloucester County, New Jersey.

Horizon: Hornerstown Formation, Rancocas Group.

Biostratigraphy:

Age: Late Maastrichtian Stage, uppermost Senonian subepoch, Gulf Epoch, Late Cretaceous.

Material: 4 vertebrae, the shaft of a femur and 4 dermal bones.

Referred material:

Locality: Arneytown, Burlington County, New Jersey.

Horizon: Greensands.

Biostratigraphy:

Age: Late Cretaceous.

Material:

Number: Not given:

 

GALLEGHER, 1993

Locality: Pemberton, New Jersey.

Horizon: Hornerstown Formation, Rancocas Group.

Biostratigraphy:

Age: Late Maastrichtian Stage, uppermost Senonian subepoch, Gulf Epoch, Late Cretaceous.

Material:

ANSP 9185-9:

 

= Thoracosaurus brevispinus COPE, 1867
Etymology:
= Holops brevispinus (COPE, 1867)
= Holopsisuchus brevispinus (COPE, 1867)

Holotype: Geol. Mus. Rutgers College Coll. (Now AMNH 2200)

Locality: Burlington County, New Jersey.

Horizon:

Biostratigraphy:

Age: Late Cretaceous.

Material: Posterior part of a skull, vertebrae.

Paratype: Number not given.

Locality: St. George's, New Castle County, Delaware.

Horizon: Late Cretaceous.

Material: Right humerus and a dorsal vertebra.

 

Paratype: Number not given.

Locality: Burlington County, New Jersey.

Horizon:

Biostratigraphy:

Age: Late Cretaceous.

Material:

 

Hypotype: Number not given.

Locality: Monmouth County, New Jersey.

Horizon:

Biostratigraphy:

Age: Late Cretaceous.

Material:

 

Paralectotype (Lectotype) 9168

Locality: St. George’s , New Castle County, Delaware.

Horizon:

Biostratigraphy:

Age: Cretaceous.

Material: Cervical vertebra.

 

= Holops cordatus COPE, 1869
Etymology:
= Holopsisuchus cordatus (COPE, 1869)

Holotype: AMNH 2206

Locality: "From Barnesboro", Gloucester County, New Jersey.

Horizon: Hornerstown Formation, Rancocas Group.

Biostratigraphy:

Age: Late Maastrichtian Stage, uppermost Senonian subepoch, Gulf Epoch, Late Cretaceous.

Material: Dorsal vertebra.

 

= Holops glyptodon COPE, 1869
Etymology:
= Holopsisuchus glyptodon (COPE, 1869)

Syntypes:

Locality: Barnesboro, Gloucester County, New Jersey.

Horizon: Hornerstown Formation, Rancocas Group.

Biostratigraphy:

Age: Late Maastrichtian Stage, uppermost Senonian subepoch, Gulf Epoch, Late Cretaceous.

Material:

9388, 9389, 9390, 9391, 9392, 9393: Teeth fragments.

 

= Holops pneumaticus COPE, 1872
Etymology:
= Holopsisuchus pneumaticus (COPE, 1872)

Holotype: American Museum Cope Collections 2217

Locality: Harrisonville, Gloucester County, New Jersey.

Horizon: Greensands.

Biostratigraphy:

Age: Late Cretaceous.

Material: Teeth, muzzle, cervical vertebrae and long bones.

Referred material:

MOOK, 1931

Locality: Permutiut Company at Birmingham, New Jersey.

Horizon: Marl pits, Hornerstown Marls.

Biostratigraphy:

Age: Late Paleocene Epoch, Lower Paleogene Period- Early Eocene Epoch, Paleogene Period, Lower Tertiary Subera, Lower Cenozoic Era.

Material:

New Jersey State Museum: Posterior top of skull, posterior portion of dentary, 18 teeth, 3 cervical, 1 dorsal 1 lumbar, 2 sacral, and 9 caudal vertebrae, right scapula, left coracoid, left humerus, right and left ulnae, both ilia, both ischia, left pubis, both femora, both tibiae, 1 metapodial, 1 tarsal, a partial sternal, 10 ribs and many dermal scutes.

 

= Thoracosaurus mullicensis TROXELL, 1925
Etymology:

Holotype: YPM 283

Locality: Mullica Hill, Gloucester County, New Jersey.

Horizon: Marl beds.

Biostratigraphy:

Age: Late Cretaceous.

Material: Part of a right maxillary with teeth, posterior end of left maxillary, ends of both quadrates, 2 foot bones, 1 of the atlantal neural halves, fragments of ribs, scutes and dentary.

 

= Thoracosaurus meirsanus TROXELL, 1925
Etymology:

Holotype: YPM 404

Locality: Hornerstown, Monmouth County, New Jersey.

Horizon: Marl Beds, Hornerstown Formation, Rancocas Group.

Biostratigraphy:

Age: Late Maastrichtian Stage, uppermost Senonian subepoch, Gulf Epoch, Late Cretaceous.

Material: Table of the skull, parietal, frontal, post-frontal, lachrymal, and squamosal, the occipital condyle, the articular ends of the quadrates, a portion of the quadratojugal and jugal, parts of the mandible, angular, dentary, splenial, etc., and many unidentified fragments.

_____________________________________________________________________________________

Species: macrorhynchus (BLAINVILLE, 1839-64)
Etymology: Greek, macro, “big, large” and Greek, rhynchus, “beak, snout.”
= Crocodilus macrorhynchus BLAINVILLE, 1839-64
= Tomistoma macrorhynchus (BLAINVILLE, 1839-64)

Holotype: MNHN 1902-22

Locality: Bourgogne, Marne Department, France.

Horizon:

Biostratigraphy:

Age: Paleocene Epoch, Lower Paleogene Period, Lower Tertiary Subera, Cenozoic Era.

Material: Incomplete skull and jaw material.

Referred material:

GERVAIS, 1859

Number: Not given: Skull, and jaw material, vertebrae, a derma scute, etc.

 

KOKEN, 1888

Locality: Maastricht, Limburg Province, The Netherlands.

Horizon:

Biostratigraphy:

Age: Late Cretaceous.

Material:

Number: Not given: Imperfect skull.

 

LYDEKKER, 1888

Locality: Pisolite of Mont Aime, Marne Department, France.

Horizon:

Biostratigraphy:

Age: Late Cretaceous.

Material:

BMNH 28296: Teeth.

BMNH 28296a: Centra.

BMNH 28296b: 3 distal femora.

 

= Gavialis isorhynhus POMEL, 1847
Etymology:

Holotype:

Locality: Pisolite of the Paris Basin, France.

Horizon:

Biostratigraphy:

Age: Late Cretaceous.

Material:

_____________________________________________________________________________________

Species: scanicus TROEDSSON, 1924
Etymology:

Holotype: LO 3076

Locality: Sweden.

Horizon:

Biostratigraphy:

Age: Late Cretaceous.

Material: Skull, and lower jaw, cervical and dorsal vertebrae, ribs, humeri, dermal scutes and gastraliths.

LO 3077T, LO 3083T, LO 3086T, LO 3098T, LO 3105T, LO 3103T:

Referred material:

BORISAK, 1913

Locality: Crimea, Sweden.

Horizon:

Biostratigraphy:

Age: Late Cretaceous.

Material:

Number: Not given: Skull.

 

= Gavialis cf. macrorhynchus LUNDGREN, 1890

Locality: Malmo, Sweden.

Horizon:

Biostratigraphy:

Age: Senonian Subepoch, Gulf Epoch, Late Cretaceous.

Material:

Number: Not given: Rostral fragment, skull, teeth and scutes.

_____________________________________________________________________________________

Species: cherifiensis LAVOCOT, 1955
Etymology:

Holotype:

Locality: South Morocco.

Horizon:

Biostratigraphy:

Age: Cenomanian Stage, Uppermiddle Gallic Subepoch, Lowermost Gulf Epoch-Turonian Stage, Lowermost Senonian Subepoch, Lower Gulf Epoch, Early Cretaceous.

Material: Skull fragments.
_____________________________________________________________________________________

Species: borissiaki EFIMOV, 1988
Etymology:
= Thoracosaurus macrorhynchus BORISYAK, 1913

Holotype: TsNIGRI c. 3373

Locality: Inkerman, Crimea Region, European Russia.

Horizon:

Biostratigraphy:

Age: Maastrichtian Stage, Upper Senonian Subepoch, Gulf Epoch, Late Cretaceous.

Material: Skull missing anterior tip.