Ichnogenus: Rhynchosauroides BEASLEY in MAIDWELL, 1911
Etymology: Rhynchosaurus and Greek, oides, "similar to, likeness."
= Akropus RUHLE von LILIENSTERN, 1939
Etymology:
= Dactylotherium WALDTHIER, 1917
Etymology:
= Eurichnus LULL, 1942
Etymology:
= Ganasauripus VALENTINI, CONTI & MARIOTTI, 2007
Etymology: From "Gana," Fairy figure of the "Ladini" legends, a brook nymph.
= Hamatopus RUHLE von LILIENSTERN, 1939
Etymology:
= Pontopous NOPCSA, 1923
Etymology:
= Rhynchocephalichnus HUENE, 1941
Etymology:

Ichnospecies: sublacertoides (PHOLIG, 1893)
Etymology:
= Protiritonichnites sublacrtoides PHOLIG, 1893

Holotype:

Locality: Hildburghausen, Thuringina State, Germany.

Horizon:

Biostratigraphy:

Age: Buntsandstein, Scythian Epoch, Early Triassic.

Material:
_____________________________________________________________________________________

Ichnospecies: rectipes BEASELY in MAIDWELL, 1911
Etymology:

Holotype:

Locality:

Horizon:

Biostratigraphy:

Age:

Material:

Referred material:

= Rhynchosauroides cf. rectipes KOTANSKI, GIERLINSKI & PTASZYNSKI, 2004

Locality: Belvedere (Bkherdous), Djurdura Mountains, Algeria.

Horizon: Haizer-Akouker Unit.

Biostratigraphy:

Age: Middle Triassic.

Material:

Muz PIG 1672.II.1: Track on slab 1.


_____________________________________________________________________________________

Ichnospecies: membranipes MAIDWELL, 1911
Etymology:
= Pontopus membranipes (MAIDWELL, 1911), O. KUHN, 1958

Holotype:

Locality:

Horizon:

Biostratigraphy:

Age:

Material:

Referred material:

KING & THOMPSON, 2000

Locality: Hilbre (gird reference SJ 186 876) Wirral, Northwest England, Southern United Kingdom.

Horizon: Sherwood Sandstone Group.

Biostratigraphy:

Age: Early to Middle Triassic.

Material:

LIVCM 1995.163. A-AQ (slab 1): 5 print casts.

_____________________________________________________________________________________

Ichnospecies: minutipes MADIWELL, 1911
Etymology:

Holotype:

Locality: Weston near Runcorn, Cheshire County, England, Southern United Kingdom.

Horizon: Keuper.

Biostratigraphy:

Age: Late Triassic.

Material:
_____________________________________________________________________________________

Ichnospecies: pisanus (FUCINI, 1915)
Etymology:
= Ichnites pisanus FUCINI 1915
= Rhynchocephalichnus pisanus (FUCININ, 1915) HUENE, 1942

Holotype:

Locality: La Costia, Italy.

Horizon:

Biostratigraphy:

Age: Late Triassic.

Material: Manus and pes prints.
_____________________________________________________________________________________

Ichnospecies: beasleyi NOPCSA, 1923
Etymology:

Holotype:

Locality: Runcorn, Cheshire County, England, Southern United Kingdom.

Horizon: Keuper.

Biostratigraphy:

Age: Late Triassic.

Material:
_____________________________________________________________________________________

Ichnospecies: maidwelli NOPCSA, 1923
Etymology:

Holotype:

Locality: Storeton near Birkenhead, Cheshire County, England, Southern United Kingdom.

Horizon: Keuper.

Biostratigraphy:

Age: Late Triassic.

Material:
_____________________________________________________________________________________

Ichnospecies: tirolicus ABEL, 1926
Etymology: In referece to Subtirol, Braies Dolomites, Bolzano, Northern Italy, the geographical region of origin.

Holotype: University of Wien.

Locality: Lapadures, Flatschkofel-Col Valacia, Braies Dolomites, Bolzano, Northern Italy.

Horizon: Uper Voltago Congomerate, Richthofen Conglomearte and Morbiac Dark Limestone.

Biostratigraphy:

Age: Lower Ladinian Stage, Upper Mid Triassic Epoch, Late Triassic.

Material: 3 imcomplete manus-pes tracks.

Referred material:

AVANZINI, 1999, AVANZINI & RENESTO, 2002

Locality: Upper Val di Non, Val d’Adige, Bolzano, Northern Italy.

Horizon:

Biostratigraphy:

Age: Anisian Stage, Lower Mid Triassic Epoch, Middle Triassic.

Material:

Number: Not given: Trackways.

 


DIEDRICH, 2002

Locality: Hagen, Silbersee, North Rhine-Westphalia State, Germany.

Horizon: Bed 1, Upper Wellenkalk Member, Lower Muschelkalk.

Biostratigraphy: Horizon II.

Age: Anisian Stage, Lower Mid Triassic Epoch, Middle Triassic.

Material:

NMO No. pal. 121/1: Pes imprint, negative.

NMO No. pal. 121/2: Claw imprent negative.

 


Locality: Detmold, Buchenberg, North Rhine-Westphalia State, Germany.

Horizon: Upper Wellenkalk Member, Lower Muschelkalk.

Biostratigraphy:

Age: Anisian Stage, Lower Mid Triassic Epoch, Middle Triassic.

Material:

Bed 1, Horizon 1

LLM No. mm12: Pes imprint negative.

LLM No. mm26: Manus/pes imprint negative set.

LLM No. mm27: Manus imprint negative.

Bed II, Horizon VIII

LLM No mm1: Manus imprint negative.

LLM no. mm2: Pes imprint negative.

LLM No. mm3: Pes imprint negative.

LLM No. mm4: Pes imprinet negative.

LLM No. mm7: Manus/pes set imprint negatives.

LLM No. mm19: Pes imprint negative.

LLM No. mm29: Manus imprint negative.

Bed II, Horizon IV

LLM No. mm13: Pes imprint negative.

Bed II, Horizon III

LLM No. mm14: Pes imprint negative.

 


Locality: Ostheim, Bayern (Bavaria) State, Germany.

Horizon: orbicularis Member, Middle Muschelkalk.

Biostratigraphy:

Age: Mid Triassic Epoch, Middle Triassic.

Material:

Bed I, Horizon I

GPIM No. B4B.14-1: Pes imprint.

Bed I, Horizon II

GPIM No. B4B. 14-2: Manus/pes imprint negative.

Bed I, Horizon III

GPIM No. B4B. 14-3: Manus imprint negative.

Bed I, Horizon V

GPIM No. B4B. 14-6: Pes imprint negative.

Bed I, Horizon VI

GPIM No. B4B 14-5: Pes imprint negative.

 

 

Locality: Niederlistingen, Thuringina State, Germany.

Horizon: orbicularis Member, Middle Muschelkalk.

Biostratigraphy:

Age: Mid Triassic Epoch, Middle Triassic.

Material:

RW No. mm1-1: Pes imprint negative.

VALDISERII & AVANZINI, 2007

Locality: Tisens (Bozen), Adige Valley, Trentino, Italy.

Horizon:

Biostratigraphy:

Age: Pelsonian, Anisian Stage, Lower Mid Triassic Epoch, Middle Triassic.

Material:

BzBG 2d, 6, 9d, 10, 12, 14, 15a/b, 20, 24-28, 30-33, 35, 37, 40a/b, 41a/b, 42, 43, 43d, 40a/b, 51, 52a/b, 53, 56, 58, 64, 66, 68a/b, 69, 71, 73, 75d: 88 footprints on 43 slabs.

Note: Some show skin and phalangeal pad impressions.

_____________________________________________________________________________________

Ichnospecies: schlauersbachensis (WEISS, 1934)
Etymology: In reference to Schlauersbach, France.
= Aetosauripus schlauersbachensis WEISS, 1934

Holotype:

Locality: Schlauersbach, France.

Horizon: Keuper, Plattensandstein.

Biostratigraphy:

Age: Late Triassic.

Material:
_____________________________________________________________________________________

Ichnospecies: schochardti (RHULE von LILIENSTERN, 1939)
Etymology:
= Akropus schochardti RHULE von LILIENSTERN, 1939

Holotype:

Locality: Thurgina State, Germany.

Horizon:

Biostratigraphy:

Age: Buntsandstein, Scythian Epoch, Middle Triassic.

Material:

Referred material:

Locality: Arizona.

Horizon: Holbrook Member, Moenkopi Formation.

Biostratigraphy:

Age: Early Anisian Stage, Lower Mid Triassic Epoch, Middle Triassic.

Material:

Number: Not given:

 

= Rhynchosauroides cf. schochardti MUJAL, FORTUNY, BOLET, OMS & LOPEZ, 2017

Locality: Port del Canto, Pyrenean Basin, Catalan Pyrenees, northeast Iberian Peninsula, Spain.

Horizon:

Biostratigraphy:

Age: Late Olenekian Stage, Late Scythian Epoch, Late Early Triassic.

Material:

Ichnospecies I

IPS-82612, 82619: Isolated tracks.

In situ: Isoalted tracks.


= Akropus langi RUHLE von LILIENSTERN, 1939
Etymology:

Holotype:

Locality:

Horizon:

Biostratigraphy:

Age:

Material:

 


= Hamatopus wildfeueri RUHLE von LILIENSTERN, 1939
Etymology:

Holotype:

Locality:

Horizon:

Biostratigraphy:

Age:

Material:

 

= Rhynchosauroides moenkopiensis HAUBOLD, 1970
Etymology: In reference to the Moenkopi Formation.

Holotype:

Locality: Shadow Mountian, V 4208, Cameron, Coconino County, Arizona.

Horizon: Holbrook Member, Moenkopi Formation.

Biostratigraphy:

Age: Perovkan, Anisian Stage, Early Mid Triassic Epoch, Middle Triassic.

Material:

UCMP 37343, 37344, 37795-97, 37800: Manus and pes tracks.
Note: Group 2 of PEABODY, 1948

 


_____________________________________________________________________________________

Ichnospecies: palmatus (LULL, 1942)
Etymology:
= Collettosaurus palmatus LULL, 1942
= Akropus palmatus (LULL, 1942) PEABODY, 1948

Holotype: YPM 14100-14103

Locality: Old Thermopolis road about 18 miles northwest of Lysite, Fremont County, Wyoming.

Horizon: Chugwater Formation about 40 feet below the Alcova Limestone.

Biostratigraphy:

Age: Spathian Stage, Uppermost Scythian Epoch, Late Early Triassic.

Material: 4 slabs.

Referred material:

= Rhynchosauroides cf. palmatus MARCHETTI, COLLARETA, BELVEDERE & LEONARDI, 2021

Locality: Monti Pisani Massif, Tuscany, Central Italy.

Horizon: Member S4, Quarziti del Monte Serra Formation.

Biostratigraphy:

Age: Middle Triassic.

Material:

MGPUF-IGF 5209: 2 logn trackways aligned in teh same direction or slightly divergin. Both show straight tail impression and partial overste of the pes on teh manus, concave epirelief and convex hyporelief.

MGPUF-IGF 5195: Trackway with sinous tail impression, convex hyporelief.

MGPUF-IGF 5162: 2 consecutive right pes-manus couples and isoalted tracks, convex hyporelief.

MGPUF-IGF 5208: Right manus imprint, convex hyporelief.

MGPUF-IGF 5182: Right pes-manus couple, convex hyporelief.

MGPUF-IGF 54T: Left manus imprint, concave epirelief.

MGPUF-IGF 467T: Left pes imprint, convex hyporelief.

MGPUF-IGF 508T: Isoalted tracks, concave epirelif.

MGPUF-IGF 512T: Incomplete step cycle with straight tail impression.

MGPUF-IGF 515: Isolated tracks, concave epirelief.

MGPUF-IGF 566T: Right manus imprint, convex hyporelief.

MGPUF-IGF 670T: Left pes-manus couple showing complete overstep and isolated tracks, convex hyporelif.

MGPUF-IGF 711T: Incomplete step cycle, convex hyporelief.

MGPUF-IGF 778T: Left manus imprint, convex hyporelief.

MGPUF-IGF 820T: Left pes-manus couple, concave eporelief.

MGPUF-IGF 927T: Right manus imprint, convex hyporelif.

MGPUF-IGf 992T: Isoalted tracks, convex hyporelief.

MGPUF-IGF 1005T: 2 right pes-manus couples, convex hyporelief.

MGPUF-IGF 1023T: Isoalted tracks, convex hyporelief.

MGPUF-IGF 1101T: Left manus imprint, convex hyporelief.

MGPUF-IGF 1105T: Left manus imprint, onvex hyporelief.

MGPUF-IGF 5154: Isoalted tracks, concave epirelief.

MGPUF-IGF 5160: Isolated tracks, concave eprielif and convex hyporelif.

MGPUF-IGF 5163: Incomplete step cycle, convex hyporelief.

MGPUF-IGF 5164: Isoalted tracks, conex hyporelif.

MGPUF-IGF 5168: Left pes-manus couple, convex hyporelif.

MGPUF-IGF 5178: Left pes-manus couple and right manus imprint convex hyporelif.

MGPUF-IGF 5185: Left manus, concave epirelief.

MGPUF-IGF 5193: Isolated tracks, convex hyporelief.

MGPUF-IGF 5201: Isolated tracks, convex hyporelif.

MGPUF-IGF 5204: Isoalted tracks, convex hyporelif.

MGPUF-IGF 5208B: Trackway and incomplete stepcyclek with stratigh tail impression, convex hyporelief.

MGPUF-IGF 8638: Left pes-manus couple, concave epirelief.

MGPUF-IGF 8639: Right manus imprint, concave epirelief.

MGPUF-IGF 8641: Right pes imprint, convex hyporelif.

MGPUF-IGF 8643: Isoalted tracks, concave epirelief.

MGPUF-IGF 8648: Incomplte step cycle, convex hyporelif.

MGPUF-IGF 8652: Right pes-manus couple, complete onverstep, concave epirelief.

MGPUF-IGF 8654: Right manus improint and isolated tracks, convex hyporelif.

MGPUF-IGF 8655: Isoalted tracks, convex hyporelif.

MGPUF-IGF 12566: 2 isolated pes-manus couples and isoalted tacks, concave epirelief.

MSNUP-I 12994: Left pes-manus couple showing complete overstep adn isoalted tracks, concave eprielief.

MSNUP-I 12997: Left pes-manus coupole, concave epirelief.

MSNUP-I 13477: Isolated tracks, convex hyporelif.

MSNUP-I 13480: Several isloated tracks, convex hyporelif.

MSNUP-I 13481: left pes-manus couple and isoalted tracks, concave epirelief.

MSNUP-I 13484: Left pes-manus couiple showing complete overstep and 2 isolated tracks, concave epirelief.

MSNUP-I 13487: Isolated tracks, convex hyporelif.

MSNUP-I 13488: Isoalted tracks, convex hyporelif.

MSNUP-I 13494: Incomplete step cycle, right pes-manus couple showing complete overstep and isolated tracks, convex hyporelif.

MSNUP-I 13496/13521: left manus imprint, concave epirelief adn convex hyporelief.

MSNUP-I 13514: Right pes-manus couple, complete overstep, convex hyporelif.

MSNUP-I 13519: Right pes-manus couple, concave epirelief and convex hyporelif.

MSNUP-I 13520: Right pes-manus couple, concave epirelif and convex hyporelif.

MSNUP-I 13532/13535: Left manus imprint, concave epirelief and concave hyporelif.

MSNUP-I 13538: Right pes-manus coulpe, partial overstep and isolated tracks, convex hyporelif.

MSNUP-I 13542: Left pes-manus couple, complete overstep and isolated tracks, convex hyporelif.

MSNUP-I 13549: Right pes-manus couple, complete overstep, digit bifurcation, concave eprielif.

MSNUP-I 13551: Several isolated tracks, concve epirelief and convex hyporelif.

MSNUP-I 16945: Isolated pes and manus imprints, convex hyporelif.

GAMPS-Malesci_243: Pes-manus couple and isolated tracks, concave epirelief and convex hyporelif.

 

 

= Eurichnus jenseni LULL, 1942
Etymology:
= Eurichrinus jeneseni BRANSON, 1947 (sic)

Holotype: YPM VP-14100

Locality: Old Thermopolis road about 18 miles northwest of Lysite, Fremont County, Wyoming.

Horizon: Chugwater Formation.

Biostratigraphy:

Age: Spathian Stage, Uppermost Scythian Epoch, Late Early Triassic.

Material: A slab of red sandstone containing a sequence of 2 impressions each of the right manus and pes, a partial impression of the left manus, and the tail trace.

Referred material:

YPM VP-14102: A slab with manus impressions.

_____________________________________________________________________________________

Ichnospecies: etruscus (HUENE, 1941)
Etymology:
= Rhynchocephalichnus etruscus HUENE, 1941

Holotype:

Locality: Verrucano, Italy.

Horizon:

Biostratigraphy:

Age: Late Triassic.

Material: Manus and pes prints.
_____________________________________________________________________________________

Ichnospecies: brunswickii (RYAN & WILLARD, 1947))
Etymology:
= Kintneria brunswickii RYAN & WILlARD, 1947

Holotype:

Locality: Bucks County, Pennsylvania.

Horizon: Lockatong Formation.

Biostratigraphy:

Age: Late Triassic.

Material:

Referred material:

BAIRD, 1964

Locality: Phoneixville Tunnel, Chester County, Pennsylvania.

Horizon: Lockatong Formation.

Biostratigraphy:

Age: Late Triassic.

Material:


AMNH 2972: Manus imprint.
Note: “Microsauiran amphibian footprint” from Moodie, 1909

 


SMITH, 1995

Locality: Wayne Covington of the State Correctional Institution at Graterford, Eastern Pennsylvania.

Horizon: Passiac Formation.

Biostratigraphy:

Age: Norian Stage, Middle Late Triassic Epoch, Late Triassic.

Material:

Number: Not given: Footprints.

 

= Rhynchosauroides cf. R. brunswickii SUES, & OLSEN, 2015

Locality: Locality B and C, Cliffs and beach just south-southwest of Paddy Island, Kings County, Nova Scotia Province, Canada.

Horizon: Red Head Member, lower Blomidon Formation, Newark Supergroup.

Biostratigraphy:

Age: Early Norian Stage, Lower-Middle Late Triassic Epoch, Late Triassic.

Material:

In situ: Manus and pes prints.

 


= Orthodactylus howelli BOCK, 1952
Etymology:
= Rhynchosauriodes howelli (BOCK, 1947) BAIRD, 1957

Holotype: ANS 15219

Locality: Near Schwenkville, Montgomery County, Pennsylvania.

Horizon: Brunswick shale.

Biostratigraphy:

Age: Late Triassic.

Material: Left manus imprint which is deceptively distorted by digital drag marks.

_____________________________________________________________________________________

Ichnospecies: franconicus (HELLER, 1956)
Etymology:
= Rhynchocephalichnus franconicus HELLER, 1956

Holotype:

Locality: Ansbach, and Hassfurt, France.

Horizon: Benkerstein, and Hangednes des Semionontensandstein, Middle Keuper.

Biostratigraphy:

Age: Late Triassic.

Material:

Referred material:

HELLER, 1958

Locality: Near Hasfurt a. m. Germany.

Horizon: Keuper.

Biostratigraphy:

Age: Late Triassic.

Material:

Number: Not given: Pes and manus prints.

_____________________________________________________________________________________

Ichnospecies: peabodyi (FABER, 1958) DEMATHIEU & OOSTERNIK, 1983
Etymology:
= Chirotherium peabodyi FABER, 1958

Holotype:

Locality: Winterswikj, Netherlands.

Horizon: Lower Muschelkalk.

Biostratigraphy:

Age: Anisian Stage, Lower Mid Triassic Epoch, Early Middle Triassic.

Material:

Referred material:

ALBAT, 1999

Locality: Borgholzhausen, Teutoburger Wald, Germany.

Horizon: Lower Muschelkalk.

Biostratigraphy:

Age: Anisian Stage, Lower Mid Triassic Epoch, Early Middle Triassic.

Material:

Trackways. Some swimming some walking.

 


DEMATHIEU & OOSTERNIK, 1983

Locality: Winterswijk, eastern part of Holland.

Horizon: Lower Muschelkalk.

Biostratigraphy:

Age: Anisian Stage, Lower Mid Triassic Epoch, Early Middle Triassic.

Material:

Jagt, Poppe, Platte D5, D50, D11, D27, D17, D16, D78a, D78b, D70, D98, D168a, D168b: Trackways.

 


DIEDRICH, 1998

Locality: Track Horizon 1, Wichlinghausen, Central Teutoburg Forest, Lower Saschen (Saxony) State, North West Germany.

Horizon: Oolith Zone, Lower Muschelkalk, Bithyn.

Biostratigraphy:

Age: Anisian Stage, Lower Mid Triassic Epoch, Early Middle Triassic.

Material:

Palaontologischesa Museum der Westfalischen Wilhelms-Universitat Munster, Nr. B4 B.6-3916,1 Wi Fa 1: Left pes.

 


DIEDRICH, 2000

Locality: Osnabrueck, Osnabrueck Hills, Lower Saschen (Saxony) State, North West Germany.

Horizon: Lower Wellenkalk, Oolith Member (Graue Zwischenittel), Lower Muschelkalk.

Biostratigraphy:

Age: Anisian Stage, Lower Mid Triassic Epoch, Early Middle Triassic.

Material:

GPI Nr.-B4B 10-Os 1:

GPI Nr.-B4B 10-Os 2: Pes (negative), (track horizon V, muW1C).

GPI Nr.-B4B 10-Os 3:

GPI Nr.-B4B 10-Os 4: Left manus (negative), (track horizon V, muW1C).

GPI Nr.-B4B 10-Os 5-14:

GPI Nr.-B4B 10-Os 5-18: Digits 2, 3, 4 of pes? (negative), (track horizon WI, mu W1C).

GPI Nr.-B4B 10-Os 5-30: Left manus (negative), (track horizon VII, MuW1C).

 

Locality: Near Dissen, Teutoburger Wald, Osnabrueck Hills, Lower Saschen (Saxony) State, North West Germany.

Horizon: Lower Wellenkalk, Oolith Member (Graue Zwischenittel), Lower Muschelkalk.

Biostratigraphy:

Age: Anisian Stage, Lower Mid Triassic Epoch, Early Middle Triassic.

Material:

GPI No. B4B.6-Di 17-9, 11, 13, 16, 19-20:

GPI No. B4B.6-Di 12: Left manus (negative) (track horizon X, Graues Zwischenmittel (muOo)).

GPI No. B4B. 6-Di 14: Manus? (negative) (track horizon XV, Graues Zwischenmittel (MuOo)).

 


Locality: Bissendorf, Osnabrueck Hills, Lower Saschen (Saxony) State, North West Germany.

Horizon: Lower Wellenkalk, Oolith Member (Graue Zwischenittel), Lower Muschelkalk.

Biostratigraphy:

Age: Anisian Stage, Lower Mid Triassic Epoch, Early Middle Triassic.

Material:

GPI No. B4B.-10 Bis 1, 6, 7:

 


Locality: Eistrup, Osnabrueck Hills, Lower Saschen (Saxony) State, North West Germany.

Horizon: Lower Wellenkalk, Oolith Member (Graue Zwischenittel), Lower Muschelkalk.

Biostratigraphy:

Age: Anisian Stage, Lower Mid Triassic Epoch, Early Middle Triassic.

Material:

GPI No. B4B.10-E 1: Left manus (negative) (track horizon 1, muW1C)).

GPI No. B4B.10-E 2: Right manus, (negative) (track horizon II, muW1C)).

GPI No. B4B.10-E 3-6, 14:

 


Locality: Handarpe, Osnabrueck Hills, Lower Saschen (Saxony) State, North West Germany.

Horizon: Lower Wellenkalk, Oolith Member (Graue Zwischenittel), Lower Muschelkalk.

Biostratigraphy:

Age: Anisian Stage, Lower Mid Triassic Epoch, Early Middle Triassic.

Material:

GPI No.-B4B. 10-Ha: Left manus (negative, (track horizon 1, Graues Zwischenmittel (muOo)).

GPI No.-B4B. 10 – Ha 1: Right manus, (negative) (track horizon I, Graues Zwischenmittel (muOo)).

GPI No.-B4B. 10 – Ha 2: 3 and 4 pes prints (negative) (track horizon I, Graues Zwischenmittel (muOo)).

GPI No. –B4B. 10-Ha 3: Right manus (negative) (track horizon V, Graues Zwischenmittel (muOo)).

GPI No.-B4B. 10 – Ha 5: Manus? (negative) (track horizon I, Graues Zwischenmittel (muOo)).

GPI No.-B4B. 10-Ha 6-14, 18, 30:

 


DIEDRICH, 2002

Locality: Geilsdorf, Thuringia State, Southeastern Germany.

Horizon: Lower Muschelkalk.

Biostratigraphy:

Age: Anisian Stage, Lower Mid Triassic Epoch, Early Middle Triassic.

Material:

muOo, Geilsdorf. Nr. –B4B, 11-1: Left manus.

muOo, Geilsdorf. Nr. –B4B, 11-2: Scratch marks.

muOo, Geilsdorf. Nr. –B4B, 11-3: Tail drag mark.

_____________________________________________________________________________________

Ichnospecies: hyperbates BAIRD, 1947
Etymology:

Holotype: ANS 15210

Locality: Smith Clark quarry near Milford, 1.25 km north of Delaware River Bridge, Hunterdon County, New Jersey.

Horizon: Lower Passic Formation (Old Upper Brunswick Formation), about 5100 meters above the Triassic Base, Level B, Newark Series.

Biostratigraphy:

Age: Late Triassic.

Material: A trackway of one pes and 3 manus imprints associated with the type of Chirotherium copei.

Hypodigm:

Number: Not given: An isolated left pes on same slab.

MCZ 210: A partial left pes associated with C. parvum.

_____________________________________________________________________________________

Ichnospecies: petri DEAMTHIEU, 1966
Etymology:

Holotype: Plaque 2. I..FU-I

Locality: Northeast of Massif, France.

Horizon:

Biostratigraphy:

Age: Late Triassic.

Material: An isolated 5 toed footprint.

Hypodigm: 436 footprints.

 

DEAMTHIEU, 1970

Locality: Saint-Germain-au-Mont-d’Or, Rhone, France.

Horizon:

Biostratigraphy:

Age: Late Triassic.

Material:

Number: Not given: 4 footprints.

 

 

Locality: Chasselas, Sanone-et-Loire Department, France.

Horizon:

Biostratigraphy:

Age: Late Triassic.

Material:

Number: Not given: 6 footprints.

 

 

DEMATHIEU & SAMANA, 1970

Locality: Region of Largentiere, Ardeche Department, France.

Horizon:

Biostratigraphy:

Age: Late Triassic.

Material:

Number: Not given: Footprints.

_____________________________________________________________________________________

Ichnospecies: pusillus HAUBOLD, 1966
Etymology:

Holotype:

Locality: Harras near Hildburg, Thuringia State, Germany.

Horizon:

Biostratigraphy:

Age: Buntsandstein, Scythian Epoch, Late Triassic.

Material:
_____________________________________________________________________________________

Ichnospecies: sphaerodactylus DEMATHIEU, 1967
Etymology:

Holotype:

Locality: Mot d’Or lyonnais, Rhone Department, France.

Horizon:

Biostratigraphy:

Age: Middle Triassic.

Material:
_____________________________________________________________________________________

Ichnospecies: majus DEMATHIEU, 1967
Etymology:

Holotype:

Locality: Chasselay, Mont d’Or lyonnais, Rhone Department, France.

Horizon:

Biostratigraphy:

Age: Middle Triassic.

Material: Fragmentary footprint.

Hypodigm: 5 manus and 2 pes prints.
_____________________________________________________________________________________

Ichnospecies: bornemanni HAUBOLD, 1966
Etymology:
= Dactylotherium bornemanni WALDTHIER 1917

Holotype:

Locality: Harras near Hildburghausen, Thuringia State, Germany.

Horizon:

Biostratigraphy:

Age: Buntsandstein, Scythian Epoch, Middle Triassic.

Material:

Referred material:

Locality: Gad Berka, Thuringia State, Germany.

Horizon:

Biostratigraphy:

Age: Buntsandstein, Scythian Epoch, Middle Triassic.

Material:

Number: Not given:

_____________________________________________________________________________________

Ichnospecies: pallinii CONTI, LEONARDI, MARIOTTI,& NICOSIA, 1977
Etymology:

Holotype: N. S. 75/5/a-f

Locality: Bletterbach site, Redagno-Radein, Northern Italy.

Horizon: Arenaria di Val Gardena Formation.

Biostratigraphy:

Age: Late Permian.

Material: 3 consecutive manus-pes sets.

Paratype:

IGPF (old N. S.) 75/5/g-n: Trackway 2, trackway with 3 consecutive manus-pes sets, one partial trackway wiht one pes-manus set adn one pes.

N. S. 34/74/58: Left manus-pes set.

Referred material:

VALENTINI, CONTI & MARIOTTI, 2007

57 footprints preserved as natural casts and natural molds.

Published material:

N. S. 34/73/53/a: Left manus.

N. S. 34/73/53/b-c: Right manus-pes set.

Unpublished material:

N. S. 75/5/o-q: Left manus-pes set and left pes of the next set.

N. S. 34/79/a-d: 2 consecutive manus-pes sets.

N. S. 34/79/e-g: Left manus-pes set and the right manus of the previous set.

N. S. 34/70/h: Left pes.

N. S. 34/79/i: Right manus.

N. S. 34/80: Left manus-pes set.

N. S. 34/82: Left manus.

N. S. 34/201: Right manus-pes set.

N. S. 34/202: Right manus-pes set.

N. S. 34/203: Left manus.

N. S. 34/204: Right manus-pes set and isolated left manus.

N. S. 34/205/a-b: Right manus-pes set.

N. S. 34/206: Right manus.

N. S. 34/207: Right manus.

N. S. 34/208: Right manus-pes set.

N. S. 34/209/a-d, N. S. 34/209/e-g, N. S. 34/209/h, N. S. 34/209/1: Natural molds of natural casts.

N. S. 34/79/a-d, N. S. 34/79/e-g, N. S. 34/79/h, N. S. 34/70/i, N. S. 34/210: Right manus-pes set.

 

MARCHETTI, VOIGT & KLEIN, 2017

Locality: Trentino-Alto Adige region, Dolomites Region, NE Italy.

Horizon: Val Gardena Sandstone Formation.

Biostratigraphy:

Age: Late Permian.

Material:

IGPF 58/7: 9 pes and manus imprints.

MGD 73-90: Several tracks associated with fish casts.

MGP-PD 27712: Fragmetnary left pes-manus couple.

MGR 0001:Right pes-manus couples.

MGR 0028: 4 pes manus couples and isoalted tracks of differnt trackways.

MGR 0029: 6 tracks of a trackway and tail trace.

NMS 99/11: Trackway with 5 manus-pes sets.

NMS-BL 007: Left pes-manus coule and further tracks.

NMS-BL 021: Right manus-pes couple.

NSM-NS 34/25: 2 pes-manus couples of an incomplete trackway.

NMS-NS 34/26: 2 pes-manus couples of an incomplete trckway.

NMS-NS 35/51: Several tracks of differnt trackway, together with cf. Dromopus.

NMS-NS 34/79: At least 8 tracks of several trackways.

NMS-PZO 967: Left manus-pes couple.

NMS-NO NUM 4: Several tracks of different trackways.

NMS-NO NUM 5: Left pes-manus coupole.

UR 73/101: Right pes-manus couple.

?UR 73/113: Left pes-manus couple and ohter tracks.

UR 74/7: 3 pes-manus couples and other tracks.

UR 74/14: Right pes-manus couple.

UR 74/48: Left pes-manous couple.

UR 74/58: 3 treacks arraned in an incomlpete step cycle.

UR 75/5 (partial counterpart of IGPF 75/5):

UR 75/9: Isoalted pes and manus tracks.

?UR 75/10: Partial trackway wiht tail impressions.

UR 82/2: 3 trackways.

UR-NS 34/26b: 2 consecutie pes-manus couples with straight tail track and isolated imprints.

UR-NS 34/26c: Several pedal and manual tracks.

UR-NS 34/28: 2 long trackways and several shorter trackway segments with distinct tail/body-tracy.

UR-NS 34/28c: Left manus-pes couple.

UR-NS 34/29: 3 trackways, one with straight tail impression.

US-NS 34/33: Incomlpete ste cycle and isoalted tracks.

US-NS 34/216: Partial trackway.

UR-NS 34/217: Right manus.

UR-NS 34/224: 2 isolated pes-manus couples.

UR-NS 34/227: Right ?pes/?maus imprint.

UR-NS 34/229: Left pes-manus couple and other tracks.

UR-NO NUM 1: Pes-manus couple and other tracks.

UR-NO NUM 5: Right pes-manus couple.

UR-NO NUM 24: Indistinct trackway with tail trace.

UR-NO NUM 25: Indistinct trackway.

UR-NO NUM 26: 2 consecutive pes-manus couples adn straight tail impression, further isoalted tracks.

UR-NO NUM 27: Incomlete step cycle, wiht straight tail trace.

UR-NO NUM 28: Right pes-manus couple.

UR-NO NUM 29: Several tracks.

UR-NO NUM 30: Isolated imprints.

UR-NO NUM 31: Left pes-manus couple and tail impression.

UR-NO NUM 32: Right pes-manus couple and subsquent right pes, associated tail impression.

UR-NO NUM 33: left pes-manus couple.

UR-NO NUM 34: Several isoalted imprints.

UR-NO NUM 35: Right pes-manus couple.

UR-NO NUM 36: Right pes-manus couple and subsequent right pes, associated tail impression.

UR-NO NUM 37: 3 isoalted pres-manus couples.

UR-NO NUM 38: Left pes-manus coule.

 

= Eumekichnium gampsodactylum LEONARDI, 1951 non NOPCSA, 1923

Locality:

Horizon:

Biostratigraphy:

Age:

Material:

 

= Dromopus lacrtoides LEONARDI, 1974

Locality:

Horizon:

Biostratigraphy:

Age:

Material:

= Ganasauripus ladinus VALENTINI, CONTI & MARIOTTI, 2007

Etymology: From the "Ladini," the autocthounous inhabitants of the Dolomitic area.

Holotype: N.S. 34/28/D

Locality: Bletterbach site, Redagno-Radein, Northern Italy.

Horizon: Arenaria di Val Gardena Formation.

Biostratigraphy:

Age: Late Permian.

Material: Trackway.

Paratype:

UR-NS (old N.S). 34/80/1: 2 isolated manus-pes sets.

Referred material:

More than 100 footprints preserved as natural molds and natural casts.

Material referred by Conti et al. (1977) to lacertoid footprints:

N.S. 34/75/1: Slab showing dozens of footprints differently oriented.

Material referred by Conti et al. (2000) to Rhynchosauroides isp. (= Lepidosauria indet. f1 Conti et al., 1977):

N. S. 34/73/96/a: Right manus.

N. S. 34/73/96/b: Right pes.

N. S. 34/73/106/a: Right manus.

N. S. 34/73/106/b: Left pes.

N. S. 34/74/40/d: Left manus.

N. S. 34/74/40/e: Left pes.

N. S. 34/75/40: Left pes.

Material: referred by Conti et al. (2000) to Rhynchosauroides isp. (= Laoporus isp. in Conti et al., 1977)

N. S. 34/73/6/c-d: Unpublished material:

N. S. 34/28/A1-7, N. S. 34/28/B1-11, N. S. 34/28/C1-10, N. S. 34/28/D1-23, N. S. 34/28/E1-27: 5 trackways.

N. S. 34/28-1/a-d: 2 consecutive manus-pes sets.

N. S. 34/28-2/a-b: Left manus-pes set.

N. S. 34/80/1/a-b: Left manus-pes set.

N. S. 34/80/1/c-d: Left manus-pes set.

N. S. 34/224/a: Left pes.

N. S. 34/224/b-c: Right manus-pes set.

N. S. 34/225/a: Left pes.

N. S. 34/225/b: Right manus.

N. S. 34/226: Left pes.

N. S. 34/227: Right manus.

N. S. 34/228: Right pes.

N. S. 34/229: Several footprints differently oriented.

_____________________________________________________________________________________

Ichnospecies: virgiliae DEMATHIEU, RAMOS & SOPENA, 1978
Etymology: In honor of Doctor Carmina Virgili.

Holotype: Muestra DT-24

Locality: Rillo de Gallo, Iberica, Guadalajara Province, Spain.

Horizon:

Biostratigraphy:

Age: “Buntsandstein”, Scythian Stage, Middle Triassic.

Material: Footprints.

Hypodgim:

Numbers: Not given: Several footprints.

_____________________________________________________________________________________

Ichnospecies: brevidigitatus PTASZYNSKI, 1990
Etymology:

Holotype: MWGUW 01144: 5.2, and ZPAL R.7/41

Locality: Right bank of Swislina River, Wiory, near Ostrowiec Swietokrzyski, Swietokrzyskie (Holy Cross) Mountains, Poland.

Horizon: Wiory Formation, Labyrinthodontid beds.

Biostratigraphy:

Age: Middle Buntsandstein, Scythian Stage, Early Triassic.

Material: Plaster cast of imprint of left pes.

Paratypes:

KR 11 4: 16: Imprint of left pes.

TPW 27: 60.2: Imprint of right pes.

TPW 28: 111.1-2: 2 successive imprints of pes.

ZPAL R.7/42: 2: Rigth pes.

Referred material:

KR 11:4.5-4: Set of imprints of right pes and manus.

KR 11: 4.22-24-21-13: With paratype KR 11: 4.16, group of poorly preserved imprints of manus and pes, representing probably and incomplete trackway.

KR 11 4: 10: Isolated imprint of manus.

KR 2: 6.3-4: set of left manus-pes imprints.

MWGUW 01141: 3.37-36: Set of imprints of left pes and manus.

TPW 29: 5.3: Plaster cast isolated imprint of pes.

TPW 27, TPW 30-35: Numerous prints.

PTASZYNSKI, 2000

TP 100: 1-4: 2 consecutive sets of pes and manus.

TP 100: 5-7: Fragmentary track.

ZPAL R.7/29: 1-10: Long trackway.

ZPAL R.7/29: 11-16: Fragmentary track.

ZPAL R.7/29: 17, 18: Isolated set of prints.

ZPAL R.7/28: 1-4: Fragmentary track.

ZPAL R.7/28: 5-8: Fragmentary track.

ZPAL R.7/44: 1, 2: 2 sucessive pes prints.

_____________________________________________________________________________________

Ichnospecies: rdzaneki PTASZYNSKI, 2000
Etymology: In honor of Kazimierz Rdzanek, co-author of the first description of vertebrate footprints from Wiory.

Holotype: ZPAL R.7/20a: 3, 4

Locality: Right bank of Swislina River, Wiory, near Ostrowiec Swietokrzyski, Swietokrzyskie (Holy Cross) Mountains, Poland.

Horizon: Wiory Formation, Labyrinthodontid beds.

Biostratigraphy:

Age: Middle Buntsandstein, Scythian Epoch, Early Triassic.

Material: Set of pes and manus prints.

Referred material:

ZPAL R.7/30a: 1-8: Track composed of 4 sets of consecutive pes and poorly preserved manus prints.

ZPAL R.7/30b: 1-5: Track composed of 4 consecutive pes prints and 1 manus.

ZPAL R.7/30b: 6-10: Fragmentary track.

ZPAl R.7/30A: 9, 10: Isolated set of pes and manus prints.

ZPAL R.7/21: 84-89: A short track.

TP 442ab: 1-7, ZPAL R.7/11: 15-21: Part and counterpart of same track.
Note: ZPAL R.7/11: 18-21 has web-like structure between the right pes and claw traces.

ZPAL R.7/11: 5-14: Shallow, complete track consisting of 5 consecutie sets of pes and manus prints.

_____________________________________________________________________________________
Ichnospecies: kuletae PTASZYNSKI & NIEDZWIEDZKI, 2004
Etymology: In honor of Maria Kuleta, researcher and expert on the Holy Cross Mountains Buntsandstein lithostratigraphy and sedimentary environment.

Holotype: Muz. PIG OS-220/27:11, 12

Locality: Tumlin Grod Quarry, 11 km NW of Kielce, Holy Cross Mountains, Poland.

Horizon: Tumlin Sandstone.

Biostratigraphy:

Age: Lower Buntsandstein, Upper Dorashamian Stage (Latest Tatarnian Stage), Latest Lopingian Substage, Latest Zechstein Epoch, Latest Permian.

Material: Set of left and manual imprints, 1-14 seven sets of imprints, among them the 5th left set of pedal manual imprints,

15-31, 9 consecutive sets of prints.

119-136, 9 consecutive sets of imprints.

Referred material:

Muz. PIG OS-220/26: 2-5 trackway consist of two sets and the next consecutive manual imprints.

Muz. PIG OS-220/28: 1-11 6 consecutive sets of imprints, the second represented only by manual imprint.

Muz. PIG OS-220/29: 1 isolated left pedal imprint.

Muz. PIG OS-220/73: 1-26 eleven consecutive sets of pedal and manual imprints.

Muz. PIG OS-220/73: 27-31 three consecutive sets of pedal and manual imprints.

Muz. PIG OS-220/73: 88, 89: set of right pedal and manual imprints, the largest specimen accounted to this ichnospecies.

Muz. PIG OS-220/73: 33-38: 3 consecutive sets of imprints.

Muz. PIG OS-220/73: 39-41: Right pedal imprint and the next (left) set o fimprints.

Muz. PIG OS-220/27: 76, 77: Isolated set of right pedal and manual imprints.

Muz. PIG OS-220/27: 84, 85 isolated set of imprints.

Muz. PIG OS-220/27: 78, 79: Isolated set of imprints.


Locality: Sosnowica Hill Quarry, 8 km N of Kielce, Holy Cross Mountains, Poland.

Horizon: Tumlin Sandstone.

Biostratigraphy:

Age: Lower Buntsandstein, Upper Dorashamian Stage (Latest Tatarnian Stage), Latest Lopingian Substage, Latest Zechstein Epoch, Latest Permian.

Material:

Muz. PIG OS-220/1: Set of right pedal and manual imprints.

Muz. PIG OS-220/6: 1, 2 set of left pedal and manual imprints.

_____________________________________________________________________________________

Ichnospecies: retroversipes SILVA, FERIGOLO, CARVALHO, & FERNADES, 2008
Etymology: Specific name retroversipes, from Latin, and pes turned back.

Holotype: MCN-PIC.002/1-10

Locality: Predebron outcrop, municipality of Sao Joao do Polesine, Rio Grande do sul State, Brazil.

Horizon: Alemoa Member, Santa Maria Formation.

Biostratigraphy:

Age: Late Triassic.

Material: Trackway in concave epirelief.

Paratypes:

MCN-PIC.01/1-11, MCN-PIC.003/1-15, MCN-PIC.004/1-14, MCN-PIC.005/1-5: Trackways in concave epirelief.

Referred material:

MCN-PIC.006/1-35, MCN-PIC.025/1-7: Tracks and trackways preserved in concave epirelief.