Genus: Leptonectes McGOWAN, 1996
Etymology: Greek, leptos, "slender" and Greek, nektes, "swimmer."
= Leptopterygius HUENE 1922 non TROSCHEL, 1860 (Leptopterygius coccoi, Actinopterygii, clingfish)
Etymology: Greek, leptos, “slender”, Greek, pterygion, “fin” + us: Slinder fin.
= Strongylostinus HAWKINS, 1840
Etymology:

Species: tenuirostris (CONYBEARE 1822) McGOWAN, 1996
Etymology: Latin, tenius, "narrow, thin" and Latin, rostris, "snout."
= Ichthyosaurus tenuirostris CONYBEARE 1822
= Leptopterygius tenuirostris (CONYBEARE, 1822), HUENE, 1992

Holotype: Lost.

Locality: Lyme Regis? Dorset County, England, Southern United Kingdom.

Horizon:

Biostratigraphy:

Age: Lower Lias Epoch, Early Jurassic?

Material: Tooth.

Neotype: BSGS GSM (old IGS) 51236

Locality: Street, Somerset County, England, Southern United Kingdom.

Horizon: Blue Lias Formation.

Biostratigraphy: P. tilmanni or P. planorbis biozone.

Age: Hettangian Stage, Early Lias Epoch, Early Jurassic (Rhaetian Stage, Uppermost Late Triassic, Latest Triassic).

Material: Skull and skeleton.

Note: Ichthyosaurus tenuirostris McGowan, 1974, Massare, 1988, Leptopterygius tenuirostris Appleby, 1979, McGowan, 1989, 1991, 1993, Leptonetes tenuirostris McGowan & Motani, 2003 (as per Weeden & Chapman, 2022).

Referred material:

NHMUK PV (old BMNH) R498: Skull and skeleton.

Note: Ichthyosaurus tenuirostris Lydekker, 1889, 1892, Woodward 1893, McGowan, 1974, Massare, 1988, Leptopterygous tenuirostris Huene, 1922, McGowan, 1989, McGowan, 1993, Leptonectes tenuirostris Motani, 1999, McGowan & Motani, 2003 (from Weeden & Chapman, 2022).

OUMNH PAL-J 10305 (old OUM J10305): Skull and fragmentary skeleton.

Note: Ichthyosaurus tenuirostris McGowan, 1974, 1986, Leptopterygius tenuirostris McGowan, 1989, 1993, Leptonectes tenuirostris McGowan, 1995, 2003, McGowan & Motani,2003, 'Leptopterygous tenuirostris' Massare et al, 2021 (from Weeden & Chapman, 2022).

TTNCM (old SCM) 8372: Skull and skeleton.

Note: Ichthyosaurus tenuirostris McGowan, 1973, Leptopterygius tenuirostris McGowan, 1989, 1993 (from Weeden & Chapman, 2022).

WEEDEN & CHAPMAN, 2022

BRSLI (BATGM) M3356: Poorly preserved skeleton.

Note: Leptopterygius tenuirostris McGowan, 1989, 1993, Leptonectes tenuirostris McGowan, 1996.

 

 

DELAIR, 1960

Locality: Lyme Regis coast, Pinhay Bay- Charmouth, (SY 3291-SY 3794), Dorset County, England, Southern United Kingdom.

Horizon:

Biostratigraphy:

Age: Hettangian-Pliensbachian Stage, Lower Lias Epoch, Early Early Jurassic.

Material:

NHMUK PV OR (old BMNH) R.38522: Nearly complete skeleton.

Note: Ichthyosaurus tenuirostris Lydekker, 1889, Leptonectes tenuirostris McGowan, 1996 (from Weeden & Chapman, 2022).

NHMUK PV (old BMNH) R1211: Nearly complete skeleton.

NHMUK PV (old BMNH) R1126:

NHMUK PV (old BMNH) 40348: Skull of a juvenile with crushed cranium.

Note: Ichthyosaurus tenuirostris Lydekker, 1889, Leptonectes tenuirostris McGowan, 1996 (from Weeden & Chapman, 2022).

Dorset County Museum G.4, G.5:

Specimens at the Osvord University Museum.

VINCENT, TAQUET, FISCHER, BARDET, FALCONNET, & GODEFROIT, 2014

MNHN AC 9937: A partial postcranial skeleton including 20 presacral vertebrae, ribs, pectoral girdle, both humeri, ulnae, radii and femora.

 

= Leptopterygius cf. tenuirostris

Locality: Lyme Regis coast, Pinhay Bay- Charmouth, (SY 3291-SY 3794), Dorset County, England, Southern United Kingdom.

Horizon:

Biostratigraphy:

Age: Hettangian-Pliensbachian Stage, Lower Lias Epoch, Early Early Jurassic.

Material:

NHMUK PV (old BMNH) 32396: A small left scapula.

NHMUK PV (old BMNH) 2036: A slab with coracoids and anterior portion of a vertebral column.

NHMUK PV (old BMNH) 1189: A slab with coracoids.

 

DELAIR, 1974

Locality: Barnstone Cement Company's pit, 1 mile west of Barnstone Church, Nottinghamshire County, England, Southern United Kingdom.

Horizon:

Biostratigraphy:

Age: Lower Lias Epoch, Early Jurassic.

Material:

VR 57: Right humerus, radius, ulna of a moderately sized right limb.

 


Locality: Found in an old quarry adjacent to Tengrove Lane, near Upton, Somersetshire County, England, Southern United Kingdom.

Horizon:

Biostratigraphy:

Age: Lower Lias Epoch, Early Jurassic.

Material:

Mr. L. Walrond private collection: Humerus, ulna and radius.

 

GODEFROIT, 1992

Locality: Differt, Lorraine, Belgium.

Horizon:

Biostratigraphy:

Age: Middle Lias Epoch, Middle Early Jurassic.

Material:

IRSNB R121: Humerus, radius and ulna.

 

LOMAX & MASSARE, 2012

Locality: Street, Somersetshire County, England, Southern United Kingdom.

Horizon:

Biostratigraphy:

Age: Lower Lias Epoch, Early Jurassic.

Material:

CAMSM J 35279: Fragmentary skull and fragmentary skeleton with two embryo's (CASM J 35718).

 

LYDEKKER, 1891

Locality: A quarry near Overleigh, Street, Somerset County, England, Southern United Kingdom.

Horizon:

Biostratigraphy:

Age: Lower Lias Epoch, Early Jurassic.

Material:

NHMUK PV (old BMNH) XI: Skull and skeleton.

 

= Leptonectes cf. tenuirostris MAISCH & MATZKE, 2003

Locality: Lyme Regis, Dorset County, England.

Horizon:

Biostratigraphy:

Age: Hettangian or Sinemurian Stage, Lower Lias Epoch, Early Jurassic.

Material:

MBI 007: 3-dimensional skull lacking most of the snout.

 

MARTIN, FREY & REISS, 1986

Locality: Barrow-Upon-Soar, Leicestershire County, England, Southern United Kingdom.

Horizon: Possibly bed 9, 'bottom floor' limestone, Blue Lias Formation.

Biostratigraphy: P. planorbis sub-biozone, P. planorbis Biozone.

Age: Lower Hettangian Stage, Lower Lias Epoch, Early Jurassic.

Material:

LEICT G124.1992 (Leicestershire Museum, OS 90.1953): Near-complete skeleton with partial body outline in neck, forelimb, rear dorsal and base of tail regions.

Note: Possible foetus in situ near the pelvic girdle (Martin et al., 1986).

Note: Ichthyosaurus tenuirostris Harrison, 1977, Martin et al, 1986, Leptopterygius tenuirostris McGowan, 1989, Leptonectes tenuirostris McGowan, 1996 (from Weeden & Chapman, 2022).

 

MASSARE, LOMAX, WILLIAMS & HOWELLS, 2024

Locality: ?Weston, possibly the northern suburb of Bath, Gloucestershire County, England, Southern United Kingdom.

Horizon:

Biostratigraphy:

Age: Lias Epoch, Early Jurassic.

Material:

MANCH L.9663: Fragmentary skull and fragmentary skeleton.

 

Locality:

Horizon:

Biostratigraphy:

Age:

Material:

MANCH L.9910: Forefin, both coracoids and rib fragments.

 

McGOWAN, 1989

Locality: Street, Somerset County, England, Southern United Kingdom.

Horizon: Blue Lias Formation.

Biostratigraphy: P. tilmanni or P. planorbis Biozone.

Age: Hettangian Stage, Lower Lias Epoch, Early Jurassic.

Material:

BGS 51236: Skull and skeleton, the authenticity of the material is in question.

GTS L/AG/ARCH/18: Near complete skeleton.

BATGM M3560: Fragmentary skeleton it may be Ichthyosaurus communis.

BATGM M3558: Fragmentary skeleton possibly a composite skeleton.

BATGM M3568: Fragmentary skeleton possibly with an Ichthyosaurus communis skull.

BATGM M3575: Fragmentary skeleton with an incomplete Ichthyosaurus communis skull.

BATGM M3564: Fragmentary skeleton that has been tampered with.

BATGM M3573, R1123, 38709, GTSL.AG.ARCH.7: Fragmentary skeletons.

OUMNH PAL-J.10319 (old OUM J10319): Fragmentary skeleton probably composite skeleton.

Note: Ichthyosaurus tenuirostris McGowan, 1989, Leptonectes tenuirostris McGowna, 1996 (from Weeden & Chapman, 2022).

WEEDEN & CHAPMAN, 2022

CAMSM J35718: Foetus or neonate skull and articulated vertebrae spanning 2 limestone blocks.

CAMSM J35179: Displaced vertebrae from an adult overlap the posterior vertebrae of the foetus.

Note: Ichthyosaurus sp Seeley, 1889, Woods, 1891, Howe et al 1981, Leptonectes tenuirostris Lomax & Massare, 2012, Massare et al, 2021 (from Weeden & Chapman, 2022).

NHMUK PV (old BMNH) R 1121: Tail vertebrae right forefin and hind fin.

Note: Ichthyosaurus tenuirostris Lydekker, 1889, Leptonectes tenuirostris McGowan, 1996.

NHMUK PF (old BMNH) R 1127: Left humerus, radius and ulna.

Note: Ichthyosaurus tenuirostris Lydekker, 1889, Leptopterygius tenuirostris McGowan, 1989, Leptonectes tenuirostris McGowan, 1996, Motani et al, 1999 (from Weeden & Chapman, 2022).

SOMAG/GEO 1: Skull with disarticulated anterior trunk and right forefin.

Note: Temnodontosaurus longirostris McGowan, 1974, Leptonectes tenuirostris Taylor et al., 2009 (from Weeden & Chapman, 2022).

SOMAG/GEO 10: Skull with anterior frib cage and part of left forefin.

SOMAG/GEO 18: Skull distintegrated and skeleton.

Note: Gut contents with coleoid hooklets.

 

Locality: Unknown, England, Southern United Kingdom.

Horizon: Unknown.

Biostratigraphy:

Age: Early Jurassic.

Material:

BRLSI (old BATGM) M3566, BRLSI (old BATGM)) M3565: Partial skeletons.

Note: Leptopterygius tenuirostris McGowan, 1989 1993, Leptonectes tenuirostris McGowan, 1996 (from Weeden & Chapman, 2022).

BRSLI M3352 (old BATGM M3552): Skull and skeleton.

Note: Leptopterygius tenuirostris McGowan, 11989, 1993, Leptonectes tenuirostris McGowan, 1996 (from Weeden & Chapman, 2022).

 

 

Locality: Glastonbury or more likely from Street or Edgarley, Somerset County, England, Southern United Kingdom.

Horizon:

Biostratigraphy:

Age: Lias Epoch, Early Jurassic.

Material:

ANSP (PAS) 17307 (BRSMG Db2486 a cast that is presumed destroyed during WWII, and ANSP 20668 is another cast: Skulls and skeletons.

Note: Ichthyosaurus tenuirostris or Ichthyosaurus communis Taylor, 2016 (from Weeden & Chapman, 2022).

 

Locality: Probably Lyme Regis, Dorset County, England, Southern United Kingdom.

Horizon:

Biostratigraphy:

Age: Early Jurassic.

Material:

NHMUK PV (old BMNH) R 1120: Skull and skeleton.

Note: Ichthyosaurus tenuirostris Owen, 1881, Lydekker, 1889, Woodward & Sherborn, 1890, Leptopterygius tenuirostris Huene, 1922, Temnodontosaurus longirostris McGowan, 1974, Rolfe et al, 1988, Leptopterygius tenuirostris McGowan, 1989, Leptonectes tenuirostris McGowan, 1996 (from Weeden & Chapman, 2022).

 

Locality: Below high tide, Lyme Regis, Dorset County, England, Southern United Kingdom.

Horizon: Blue Lias Formation,

Biostratigraphy: A. bucklandi biozone.

Age: Sinemurian Stage, Lias Epoch, Early Jurassic.

Material:

ROM VP 47698: Skull and skeleton missing rigth fin.

Note: Un-named Fowles, 1986, Pamplin, 1987, Leptoptergius tenuirostris McGowan, 1989, 1993, Leptonectes tenuirostris McGowna, 1996, 2001, Sues, 2019 (from Weeden & Chapman, 2022).

 

McGOWAN, 1974

Locality: Street, Somerset County, England, Southern United Kingdom.

Horizon:

Biostratigraphy:

Age: Lower Lias Epoch, Lower Early Jurassic.

Material:

NHMUK PV (old BMNH) R498: Fragmentary skeleton possibly tampered with.

MASSARE, LOMAX, WILLIAMS & HOWELLS, 2024

BRLSI M3556: Skull, and partial skeleton.

 

Locality: Stogursey, Somerset County, England, Southern United Kingdom.

Horizon:

Biostratigraphy:

Age: Lower Lias Epoch, Early Early Jurassic.

Material:

BMB C2: Fragmentary skeleton possibly tampered with.

 

Locality: Keinton Mandeville, near Street, Somerset County, England, Southern United Kingdom.

Horizon:

Biostratigraphy:

Age: Lower Lias Epoch, Early Jurassic.

Material:

WM 527: Skull and skeleton.

Note: Leptopterygius tenuirostris McGowan, 1989, 1993, Leptonectes tenuirostris McGowan, 1996 (from Weeden 8 Chapman, 2022).

 

Locality: Barow-on-Soar, Leicestershire County, England, Southern United Kingdom.

Horizon:

Biostratigraphy:

Age: Lower Lias Epoch, Early Jurassic.

Material:

LEICS OS.90.1953: Skull and skeleton.

 


Locality: Pinhay Bay, just west of Lyme Regis, in the adjoining county of Devon, Dorset County, England, Southern United Kingdom.

Horizon:

Biostratigraphy:

Age: Lower Lias Epoch, Early Jurassic.

Material:

DLR '002': Skull and skeleton.

 

OWEN, 1854

Locality: Lyme Regis coast, Pinhay Bay- Charmouth, (SY 3291-SY 3794), Dorset County, England, Southern United Kingdom.

Horizon:

Biostratigraphy:

Age: Hettangian-Pliensbachian Stage, Lower Lias Epoch, Early Early Jurassic.

Material:

ROYAL COLLEGE OF SURGEONS: 172: Skull and nearly complete skeleton.

ROYAL COLLEGE OF SURGEONS: 174: Tooth.

 


Locality: I miles for the limestone quarry in Bath, on the lower Bristol road, about the 6 feet below the turf in a stratum of rubble, Gloucestershire County, England, Southern United Kingdom.

Horizon: Oolite.

Biostratigraphy:

Age: Early Jurassic.

Material:

ROYAL COLLEGE OF SURGEONS: 173: Nearly complete lower jaw.

 

OWEN, 1881

Locality: Lyme Regis coast, Pinhay Bay- Charmouth, (SY 3291-SY 3794), Dorset County, England, Southern United Kingdom.

Horizon:

Biostratigraphy:

Age: Hettangian-Pliensbachian Stage, Lower Lias Epoch, Early Early Jurassic.

Material:

NHMUK PV (old BMNH) R1120: Skull and skeleton.

 

REIFF, 1935, MAISCH, 1999

Locality: Alpha-Olschiefer, Dusslingen, Baden-Wurttemberg State, Germany.

Horizon:

Biostratigraphy: Tuberculateus zone.

Age: Early Jurassic.

Material:

Tubingen Geol. Uinv. Inst; GPIT/Re/400: Skull, right humerus, coracoid, interclavicle, clavicle fragment ribs and teeth.

Tubingien Geol. Univ. Inst: Dorsal vertebrae.

 


Locality: Ofterdingen, Baden-Wurttemberg State, Germany.

Horizon:

Biostratigraphy: Tubrculatus zone.

Age: Early Jurassic.

Material:

Tubingen Geol. Uinv. Inst: Dorsal vertebrae.

 


Locality: Malsch, Baden-Wurttemberg State,, Germany.

Horizon: Doppelfels.

Biostratigraphy: Kuperfel zone.

Age: Late Triassic?

Material:

Karlsruhe bad. Landessammlung F. Naturde: Dorsal and caudal vertebrae.

 

WEEDEN & CHAPMAN, 2023

Locality: Glastonbury, Somerset County, England, Southern United Kingdom.

Horizon:

Biostratigraphy:

Age: Early Jurassic.

Material:

NHMUK PV OR (old BMNH) 24300: Crushed skull.

Note: Ichthyosaurus tenuirostris Lydekker, 1889, Massare, 1987, Leptonectes tenuirostris Montani et al., 2013 (from Weeden & Chapman, 2022).

 

Locality: Tewkesbury, Gloucestershire County, England, Southern United Kingdom.

Horizon:

Biostratigraphy:

Age: Early Jurassic.

Material:

NHMUK PV OR (old BMNH) 41253; Patial forefins.

Note: Ichthyosaurus tenuirostris Lydekker, 1889, Leptopterygius tenuirostris McGowan, 1989, Leptonectes tenuirostris McGowan, 1996 (from Weeden & Chapman, 2022).

 

Locality: Southam Quarry, Southam, Warwickshire County, England, Southern United Kingdom.

Horizon: Blue Lias Formation.

Biostratigraphy: A. liassicus or S. angulata biozone.

Age: Lias Epoch, Early Jurassic.

Material:

WARMS G15647: Posterior three-dimensional skull and posterior vertebral columen including tail bend.

 

Locality: Possibly Somerset, Somerset County, England, Southern United Kingdom.

Horizon:

Biostratigraphy:

Age: Early Jurassic.

Material:

ANSP 17428: Skull missing front of rostrum, rib cage pelvic girdle and most of the backbone disarticulated.

 

Locality: Lyme Regis, Dorset County, England, Southern United Kingdom.

Horizon:

Biostratigraphy:

Age: Early Jurassic.

Material:

MBI 2099: Three-dimensional skull lacking most of snout and anterior mandible.

Note: Ichthyosaurus sp Jaekel, 1903, Leptonectes cf. tenuirostris Maisch & Matzke, 2003 (from Weeden & Chapman, 2022).

 

Locality: Between Lyme Regis and Charmouth, Dorset County, England, Southern United Kingdom.

Horizon:

Biostratigraphy: A. semicostatum biozone.

Age: Early Jurassic.

Material:

UMZC T.1038: Skull in 3 mimensions lacking snout with anterior vertebrae, ribs and partial pectoral girdle.

Note: Ichthyosaurus cf. latifrons Whitear, 1956, Unnamed Romer, 1968, Leptonectes cf. tenuirostris (Leptonectes latifrons specimens included within Leptonectes tenuirostris McGowan, 1989) Mcgowan, 1996 (from Weeden & Chapman, 2022).

= Leptopterygius cf. tenuirostris REIF

Locality: Vaihingen, Baden-Wurttemberg State, Germany.

Horizon: Angulatenton, Unter.

Biostratigraphy:

Age: Early Jurassic.

Material:

Stuttgart Naturaliensammlung: Femur.

Stuttgart Naturaliensammlung: Dorsal vertebrae.

 

= Ichthyosaurus grandipes SHARPE, 1830
Etymology:

Holotype:

Locality: 4 miles from Stratford-upon-Avon, Warwickshire County, England, Southern United Kingdon.

Horizon:

Biostratigraphy:

Age: Lias Epoch, Early Jurassic.

Material: Partial skeleton.

 

= Ichthyosaurus chirostrongulostinus HAWKINS, 1834
Etymology:

Holotype: NHMUK PV OR (old BMNH) 2009

Locality: Street, Somerset County, England, Southern United Kingdom.

Horizon: Blue Lias Formation.

Biostratigraphy: P. tilmanni or P. planorbis Biozone.

Age: Lower Hettangian Stage, Lower Lias Epoch, Early Jurassic.

Material: Skull.

Note: Ichthyosaurus chirostrongulostinus Hawkins, 1934, Strongylostinus Hawkins, 1840, Ichthyosaurus tenuirostris OWen, 1881, Lydekker, 1889, Woodward & Sherborn, 1890, Leptopterygius tenuirostris Huene, 1922, McGowan, 1898, 1991, Leptonetes tenuirostris McGowan & Milner, 1999 (from Weeden & Chapman, 2022).
= Ichthyosaurus strongylostinus HAWKINS, 1834
Etymology:
= Strongylostinus HAWKINS, 1840

_____________________________________________________________________________________

Species: margaritatus (HUENE, 1922) (nomen dubium)
Etymology:
= Leptopterygius margaritatus HUENE, 1922 (nomen dubium)

Holotype: SMNS 4147

Locality: Eislingen, Baden-Wurttemberg State, Germany.

Horizon:

Biostratigraphy:

Age: Upper Pliensbachian Stage, Middle Lias Epoch, Middle Early Jurassic.

Material: 10 vertebrae.

Referred material:

GPIT 10757: Tooth.

_____________________________________________________________________________________

Species: solei (McGOWAN, 1993) McGOWAN, 1996
Etymology:
= Leptopterygius solei McGOWAN, 1993

Holotype: BRSMG Ce 9856

Locality: Western flank of Stonebarrow, the coastal cliff that lies just east of the River Char (national Grid Reference SY 370929), Lyme Regis coast, Pinhay Bay- Charmouth, Dorset County, England, Southern United Kingdom.

Horizon: Black Ven Marls.

Biostratigraphy: A. stellare sub-zone, A. obtusum biozone (old Caenisites obtusum zone).

Age: Hettangian-Pliensbachian Stage, Lower Lias Epoch, Early Early Jurassic.

Material: Skull and nearly complete skeleton.

Note: An unusual central prominence and associated groove extending to the periphery of the flattened humeral head may be pathological (PARDO-PEREZ, KEAR, GOMEZ, MORONI & MAXWELL, 2018).

Note: Unnamed Clark, 1989, Leptopterygius solei McGowan, 1993, Benton & Spencer, 1995, Leptonectes solei McGowa, 1996,2001, McGowan & Motani, 2003, Maich & Matzke, 2003 (from Weeden & Chapman, 2022).

Referred material:

Locality: Church Cliff, Lyme Regis, Dorset County, England, Southern United Kingdom.

Horizon:

Biostratigraphy: Arnioceras semicostatum zone.

Age: Lower Sinemurian Stage, Lowermost Lias Epoch, Early Jurassic.

Material:

MHN 96270 (DLR "001" McGOWAN, 1989): Skull and skeleton.

Note: Leptopterygius sp McGowan, 1989, Leptonectes solei McGowan, 1993, Maisch & Matzke,2003 (from Weeden & Chapman, 2022).

_____________________________________________________________________________________

Species: moorei McGOWAN & MILNER, 1999
Etymology: In honor of Mr. Chris Moore, who discovered the specimen.

Holotype: NHMUK PF (old BMNH) R 14370

Locality: Foreshore , just to the west of Seatown, Dorset County, England, Southern United Kingdom.

Horizon:

Biostratigraphy: Uppermost part of the Belemnite Marls, 1 meter below the Beleminite Stone Band.

Age: Lower Pliensbachian, close to the top of the Lower Lias Epoch, Early Jurassic.
Material: An incomplete skeleton comprising the skull, fore paddles and anterior trunk.