Genus: Protoichthyosaurus APPLEBY, 1979
Etymology: Greek, protos, “first”, and Ichthyosaurus: First Ichthyosaurus.

Species: fortimanus (OWEN, 1884) LOMAX & MASSARE, 2018
= Ichthyosaurus fortimanus OWEN, 1884
Etymology:

Holotype: NHMUK R1063

Locality: Lyme Regis, Dorset County, England, Southern United Kingdom.

Horizon:

Biostratigraphy:

Age: Hettangian-Sinemurian Stage, Lias Epoch, Early Jurassic.

Material: Paddle.

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Species: prostaxalis APPLEBY, 1979
Etymology:
= Ichthyosaurus prostaxalis (APPLEBY, 1979) MAISCH & HUNGERBUHLER, 1997

Holotype: BRLSI M3553 (old B. 1963'5/O.S.)

Locality: Street, Somerset County, England, Southern United Kingdom.

Horizon: 'Pre-Planorbis Beds,', Blue Lias Formation.

Biostratigraphy: Tilmanni Zone.

Age: Lower Hettangian Stage, Lower Lias Epoch, Early Jurassic.

Material: A partial skull, pectoral girdle and both forefins.

Paratypes:

BRLSI M3563: Skull and nearly complete skeleton.

BRLSI M3555 (formerly B. 1963'7/O.S): Fragmentary skull and skeleton.

BRLSI M3565 (formerly B. 1963'15/O.S): Fragmentary skull and skeleton.

A partial forefin (note now missing? M. Evans pers com to Lomax, et al., 2017).

OUM J. 13,799: Fragmentary skull and skeleton. Note: Skeleton may be composite (MASSARE & LOMAX, 2016).

 

Locality: Barrow-on-Soar, Leicestershire County, England, Southern United Kingdom.

Horizon: Blue Lias Formation.

Biostratigraphy: C. johnstoni sub-biozone, P. planorbis biozone.

Age: Lower Hettangian Stage, Lower Lias Epoch, Early Jurassic.

Material:

LEICT G454.1951/164 (formerly LM 454'1951/164): Fragmentary skull and skeleton.

NHMUK OR29672: A partial forfin with soft part preservation.

Referred material:

WEEDEN & CHAPMAN, 2022

LEICT G142.1991: Three-eimensional skull lacking rostrum anterior teo the external naris plus partial pectoral girdle and proximal humerus.

 

 

LOMAX, MASSARE & MISTRY, 2017

Locality: Unknown, England.

Horizon: Unknown.

Biostratigraphy:

Age: Early Jurassic.

Material:

UORCMZ FE19-2: A partial skeleton of a very small individual.

BRLSI M3562: Right forefin.

BRLSI M0653: Left forefin.

BU 5323 (formelry GIRUG 27683): Skull.

 

Locality: Lyme Regis? Dorset County, England, Southern United Kingdom.

Horizon:

Biostratigraphy:

Age: Lower Lias Epoch, Early Jurassic.

Material:

OUMNH J.26971: An isoalted proximal forefin.

 

 

Locality: Street, Somerset County, England, Southern United Kingdom.

Horizon: 'Pre-Planorbis Beds,', Blue Lias Formation.

Biostratigraphy: Tilmanni Zone.

Age: Lower Hettangian Stage, Lower Lias Epoch, Early Jurassic.

Material:

SOMAG/GEO (old AGC) 12: Skull and partial skeleton.

NMING F8747: Damaged skull and partial skeleton.

NHMUK R1-63: A large forefin.

TTNCM 41/2017 (formerly TV52): A fragmentary ventral skull and forefin.

 

Localty: Severn Valley Area, Gloucestershire County, England, Southern United Kingdom.

Horizon:

Biostratigraphy:

Age: Lias Epoch, Early Jurassic.

Material:

GLAHM V1180 part and counterpart: Partial skull in a nodule.

 

Locality: Barnstone, Nottinghamshire County, England, Southern United Kingdom.

Horizon: Hydraulic Limestone.

Biostratigraphy: Tilmanni Zone.

Age: Lower Hettangian Stage, Lower Lias Epoch, Early Jurassic.

Material:

NOTNH: FS13770: A partial skull.

UOD D1: A partial skull.

 

Locality: Binton, Warwickshire County, England, Southern United Kingdom.

Horizon: Wilmcote Limestone Member, Blue Lias Formation.

Biostratigraphy:

Age: Lower Hettangian Stage, Lower Lias Epoch, Early Jurassic.

Mateiral:

WARMS G347: A complete skull.

 

Locality: Barrow-upon-Soar, Leicestershire County, England, Southern United Kingdom.

Horizon: 'Pre-Planorbis Beds,', Blue Lias Formation.

Biostratigraphy: Tilmanni Zone.

Age: Lower Hettangian Stage, Lower Lias Epoch, Early Jurassic.

Material:

NHMUK R36958: A skull.

LEICHT G729.1889.1: A partial skull that might be associated with caudla centra.

LEICT G738.1889: Poorly preserved three-dimensional skull.

NMING F8756: Partial three-dimensional skull.

NMING F16625: Partial three-dimensional skull.

 

 

Locality: Lavernock Point, Penarth, Glamorganshire County, England, Southern United Kingdom.

Horizon: Blue Lias Formation.

Biostratigraphy:

Age: Lower Hettangian Stage, Lower Lias Epoch, Early Jurassic.

Material:

NMW 2012.23G.1: A nearly complete skull.

 

LOMAX, PORRO, & LARKIN, 2019

Localitiy: Fell Mill Farm, between Shipston-on-Stour and Honingon, Warwickshire, England, Southenr United Kingdom.

Horizon: Liassic Clay.

Biostratigraphy: Semicostatum Ammonite Zone.

Age: Lower Sinemurian Stage, Lower Lias Epoch, Early Jurassic.

Material:

BMT 1955.G35.1: An almost complete, three-dimensional skull and parital postcranial skeleton.

 

WEEDEN & CHAPMAN, 2022

Locality: Lyme Regis, Dorset County, England, Southern United Kingdom.

Horizon: Blue Lias Formation.

Biostratigraphy: A. bucklandi biozone.

Age: Sinemurian Stage, Lower Lias Epoch, Early Jurassic.

Material:

GPIT RE/09728: Three-dimensional skull.

 

= Protoichthyosaurus prosostealis APPLEBY, 1979
Etymology:
= Ichthyosaurus prosostealis (APPLEBY, 1979) MAISCH & HUNGERBUHLER, 1997

Holotype: B. 1963'24/O.S

Locality: Street, Somerset County, England, Southern United Kingdom.

Horizon:

Biostratigraphy:

Age: Lower Lias Epoch, Early Jurassic.

Material: Fragmentary skull and skeleton.

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Species: applebyi LOMAX, MASSARE & MISTRY, 2017
Etymology: In honor of paleontologist Robert M. Appleby, for his contributions to the study of ichthyosaurs. He first recognized the unique forefin structure of Protoichthyosaurus adn erected the genus.

Holotype: UNM.G.2017.1 (previously, UON FR159, UON TC3199)

Locality: Possibly from Nottinghamshire or Leicestershire,

Horizon: Unknown.

Biostratigraphy:

Age: Early Jurassic.

Material: Skull and articulated skeleton.

Referred material:

LOMAX & MASSARE, 2018

Locality: Lyme Regis, Dorset County, England, Southern United Kingdom.

Horizon:

Biostratigraphy:

Age: Hettangian-Sinemurian Stage, Lias Epoch, Early Jurassic.

Material:

NHMUK R1164: Nearly complete, isolated skull with a few articualted cervical vertebrae.

 

= Protoichthyosaurus cf. applebyi KLUG, 2024

Locality: Gruhalde, Frick, Canton Argau, Switzerland.

Horizon: Upper Beggingen Member ("Arietenkalk").

Biostratigraphy: semicostatum ammonite zone.

Age: Lower Sinemurian Stage, Early Lias Epoch, Early Jurassic.

Material:

SMF 46: Fragmetnary skull.