Genus: Benthosuchus EFREMOV, 1936
Etymology:
= Benthosaurus EFREMOV, 1929 non GOODE & BEAN, 1886
Etymology:
= Parabenthosuchus OCHEV, 1958
Etymology:
= Rasaurus KUZMIN, 1938 (nomen nudum)
Etymology:

Species: sushkini (EFREMOV, 1929) EFREMOV, 1936
Etymology:
= Benthosaurus sushkini EFREMOV, 1929
= Rasaurus KUSMIN, 1937 (no species name given, possibly a replacment name for Benthosaurus, SCHOCH, & MILNER, 2000)

Holotype: PIN 9/2243/1

Locality: Vakhnevo locality, Banks of Sharzhenga river, a left tributary of the Yug, Nikol'skii District (Nikol’skij Rajon), Vologodska Oblast, east European Russia.

Horizon: Rybinskian Horizon, Vetluga Series, Shilikha Formation..

Biostratigraphy:

Age: Olenekian Stage?, Upper Scythian Epoch, Early Triassic.

Material: Skull.
Note: Stolen.

NOVIKOV, 2012: In addition to the holotype, more than 130 specimens, collection PIN, no. 2243.

Referred material:

BYSTROW & EFREMOV, 1940

Localities:

Horizon:

Biostratigraphy:

Age:

Material:

No. 4-19/22452: Skull of a juvenile.

No. 4-19/2252: Skull of a juvenile and dentaries.

No. 5-9/2243, No. 1-9/2443: Skulls.

No. 23-32/2244, No. 2-19/2252, No. 3-19/2252, No. 9-19/2252, No. 32/2354, 12-19/2252, 12-32/2354: Fragmentary skulls.

No. 45-19/2252, 34-9/2243: Lower jaw.

No. 50-32/2354, 50-9/2243, 76-19/2252, 88-19/2252, 49-9/2243, 73-19/2252: Vertebrae.

No. 134-19/2252, 135-19/2252, 82-32/2354, 67-19/2252, 196-19/2252: Scapulae.

No. 66-32/2354, 139-19/2252, 69-32/2354: Clavicles.

No. 74-19/2252, 62-322/2354, 165-19/2252, 72-32/2354, 190-19/2252: Praescapularis clavicle.

No. 148-19-2252: Interclavicle.

No. 200-19/2252, 194-19/2252, 83-9/2243, 201-19/2252, 199-19/2252, No. 209-19/2252, 85-9/2243, 198-19/2252, 207-19/2252: Humeri.

No. 217-19/2252, 214-19/2252, 219-19/2252, 215-19/2252, 213-19/2252, 211-29/2252, 152-9/2243, 216-19/2252, 212-19/2252, 290-19/2252, 88-9/2243, 110-9/2243: Ilia.

No. 246-19/2252, 233-29/2252, 245-19/2252, 244-19/2252, 95-9/2243, 98-9/2243, 94-32/2354, 95 32/2354: Femora.

Plus numerous other skulls and parts of skeletons.

 

GETMANOV, 1979

Locality:

Horizon:

Biostratigraphy:

Age:

Material:

19/2243, 32/2354: Lower jaw.

 

NOVIKOV, 2012

Locality: Kudrino Locality, Nikol'skii District, Vologda Region, European Russia.

Horizon: Rybinskian Horizon, Vetluga Series, Shilikha Formation..

Biostratigraphy:

Age: Olenekian Stage?, Upper Scythian Epoch, Early Triassic.

Material:

PIN, no. 2355: 24 specimens.

 

Locality: Bol'Shoe Aksilovo Locality, Nikol'skii District, Vologda Region, European Russia.

Rybinskian Horizon, Vetluga Series, Shilikha Formation..

Biostratigraphy:

Age: Olenekian Stage?, Upper Scythian Epoch, Early Triassic.

Material:

PIN 5410/1:

 

Locality: Makar'ev (?) locality, Neiskii District, ?Kostroma Region, European Russia.

Rybinskian Horizon, Vetluga Series, Shilikha Formation..

Biostratigraphy:

Age: Olenekian Stage?, Upper Scythian Epoch, Early Triassic.

Material:

PIN no. 3783/1, 2, 65:

 

Locality: A group of localities of Mechet', Borskii District, Samara Region, European Russia.

Rybinskian Horizon, Vetluga Series, Shilikha Formation..

Biostratigraphy:

Age: Olenekian Stage?, Upper Scythian Epoch, Early Triassic.

Material:

PIN 2424: More than ten specimens.

 

Locality: Kamennyi Dol Locality, Borskii District, Samara Region, European Russia.

Horizon: Basal Rybinskian Horizon.

Biostratigraphy:

Age: Early Triassic.

Material:

PIN 4419: Five specimens.

 

Locality: Kamennyi Yar, Sorochinskii District, Orenburg Region, European Russia.

Horizon: Basal Rybinskian Horizon.

Biostratigraphy:

Age: Early Triassic.

Material:

PIN 4197: More than 15 specimens.

 

Locality: Kamennyi 2 locality, Totskii District, Orenburg Region, European Russia.

Horizon: Basal Rybinskian Horizon.

Biostratigraphy:

Age: Early Triassic.

Material:

PIN 4195/1,9, 12: 3 specimens.

 

Locality: Shilkha 2 locality, Vetulzhskii District, Nizhni Novgorod Region, European Russia.

Horizon: Basal Rybinskian Horizon.

Biostratigraphy:

Age: Early Triassic.

Material:

PIN 2484/1:

 

Locality: Elva Vymskaya locality, Ust'-Vymskii District, Komi Republic, European Russia.

Horizon: Basal Rybinskian Horizon.

Biostratigraphy:

Age: Early Triassic.

Material:

PN 3998/1:

 

Locality: Blumental Locality, Belyaevskii District, Orenburg Region, European Russia.

Horizon: Basal Rybinskian Horizon.

Biostratigraphy:

Age: Early Triassic.

Material:

PIN 4167/1, 2: 2 specimens.

 

SHISHKIN, 1979

Locality:

Horizon:

Biostratigraphy:

Age:

Material:

3783/1: Skull.

 

= Benthosuchus cf. sushkini TVERKOKHLEBOV, TVERDOKHLEBOVA, SURKOV & BENTON, 2002

Locality: Kamenka, 5 km NE of Kamenno-Ostrovnoe village (Ural drainage basin), 30 km east of Orenburg, Orenburg Region, eastern European Russia.

Horizon: Staritskaya Svita, Rybinskian Groizont, Middle Vetlugian supergorizont.

Biostratigraphy:

Age: Middle Buntsandstein, Lowermost Olenekian Stage, upper Scythian Epoch, Early Triassic.

Material:

Number: Not given:


Locality: Markovka II, left bank of the Soroka River, left tributary of the Samara River, 0.4 km west of south outskirts of Markovka village, Orenburg Region, Eastern European Russia.

Horizon: Staritskaya Svita, Rybinskian Groizont, Middle Vetlugian Supergorizont.

Biostratigraphy:

Age: Middle Buntsandstein, Lowermost Olenekian Stage, upper Scythian Epoch, Early Triassic.

Material:

Number: Not given: Lower jaws.

 

Locality: Kamenny II, Kamenny ravine, right tributary of Soroka River, 0.6 km east of Markovka village, Orenburg Region, Eastern European Russia.

Horizon: Staritskaya Svita, Rybinskian Groizont, Middle Vetlugian Supergorizont.

Biostratigraphy:

Age: Middle Buntsandstein, Lowermost Olenekian Stage, upper Scythian Epoch, Early Triassic.

Material:

Number: Not given: Skull.

 

Locality: Panik creek (Prokayevka), right bank of the Buzuluk River, 5.2 km south of Prokaevka village, Buzulukskij district, Orenburg region, southeastern European Russia.

Horizon: Staritskaya Svita, Rybinskian Groizont, Middle Vetlugian Supergorizont (Kyzylsay Formation, Rybinskian Horizon, Vetluga Series).

Biostratigraphy:

Age: Middle Buntsandstein, Lowermost Olenekian Stage, upper Scythian Epoch, Early Triassic.

Material:

Number: Not given: Fragment of skull.

 

Locality: Mechet V, NE outskirts of Usmanka village, near the mouth of Mechet ravine, Orenburg Region, Eastern European Russia.

Horizon: Staritskaya Svita, Rybinskian Groizont, Middle Vetlugian Supergorizont.

Biostratigraphy:

Age: Middle Buntsandstein, Lowermost Olenekian Stage, upper Scythian Epoch, Early Triassic.

Material:

Number: Not given: Occiptial.

Locality: Usmanka, unnamed ravine flowing into the Tavolzhanka River from the right, near Usmanka village, Samara Region, Eastern European Russia.

Horizon: Staritskaya Svita, Rybinskian Groizont, Middle Vetlugian Supergorizont.

Biostratigraphy:

Age: Middle Buntsandstein, Lowermost Olenekian Stage, upper Scythian Epoch, Early Triassic.

Material:

Number: Not given: Skull.

_____________________________________________________________________________________

Species: uralensis (OCHEV, 1958)
Etymology:
= Parabenthosuchus uralensis OCHEV, 1958

Holotype: PIN 4167/1 (was SGU 104/3810)

Locality: Blyumental’ I site, 0.5 km east of Blyumental village (drainage basin of left bank of the Ural River), Belyaev District, Orenburg Region (Orenburgskaya Oblast), Southern Priural Region, eastern European Russia.

Horizon: Kopanskaya Svita (Kopanskoye Formation), Vokhmian Gorizont, Lower Vetulgian Supergorizont.

Biostratigraphy:

Age: Lower Buntsandstein, Induan stage, upper Scythian Epoch, Early Triassic.

Material: Skull.

Referred material:

PIN 4617/1-2: Left pterygoid.

_____________________________________________________________________________________

Species: korobkovi IVACHENKO, 1972
Etymology:

Holotype: PIN 3200/1

Locality: Tikhvinskoe (Tikvin site), 40 km below Rybinsk, right bank of Volga at Tikhvinskoye village near Rynbinsk, Andropovskij Rajon, Rybinskaya district, Yaroslavl’ Province, European Russia.

Horizon: Rybinskian Horizon, Vetluga subhorizone, Parshion Subformation, Rybinsk Formation.

Biostratigraphy:

Age: Lower Olenekian, Early Triassic.

Material: Skull lacking parts in front of nostrils.

Referred material:

PIN 3200/2, 3200/3: Skulls.

PIN 3200/4, 3200/5: Fragmentary skulls.

NOVIKOV, 2012: About 100 specimens collection number 3200.


NOVIKOV & SENNIKOV, 1995

Locality: Tikhvinskoye locality, right bank of the Volga River, 12 km from the city of Rybinsk, north of Moscow, Rybinskaya district, Yaroslavl’ Province, European Russia.

Horizon: Rybinskian Horizon.

Biostratigraphy:

Age: Early Triassic.

Material:

Number: Not given:

 

SHISHKIN, 1979

Locality:

Horizon:

Biostratigraphy:

Age:

Material:

3200/66: Skull.

3200/197, 3200/165, 32000/6, 3200/16, 3200/81, 3200/112: Fragmentary skulls.

3200/27-29, 49-58, 122-125: Lower jaws.

 

ROCEK, 1991

Locality:

Horizon:

Biostratigraphy:

Age:

Material:

PIN 1300/167, 179, 181, 207, 238, 240: Fragmentary skulls.

 

NOVIKOV, 2012

Localty: Yakovka Locality, Rybinskii District, Yaroslavi Region, European Russia.

Horizon: Rybinsk Formation.

Biostratigraphy:

Age: Early Triassic.

Material: 5400/1:

 

Locality: Tutaev Locality, Rybinskii District, Yaroslavi Region, European Russia.

Horizon: Rybinsk Formation.

Biostratigraphy:

Age: Early Triassic.

Material:

PIN n. 4260:

 

= Wetlugasaurus cf. groenlandicus BLOM, 1972

Holotype:

Locality:

Horizon:

Biostratigraphy:

Age:

Material:

_____________________________________________________________________________________

Species: bystrowi GETMANOV, 1989
Etymology:

Holotype: PIN N3783/1

Locality: Muraptalovo 1 locality (Unzha site near Makaryev), Nejskij Rajon, Kuyurgazinskii District, Kostroma region (Kostromskoj Oblast), eastern European Russia.

Horizon: Kzylsai Formation, Vetluga Subhorizon, Sludkian Horizon (old Rybinskian Horizon).

Biostratigraphy:

Age: Lower Olenekian, Early Triassic.

Material: Complete skull.
Note: Stolen.

Referred material:

PIN 3783/2: Right pterygoid.

_____________________________________________________________________________________

Species: gusevae NOVIKOV, 2012
Etymology: In honor of L. V. Guseva, an employee of SOIKM.

Holotype: SOIKM, no. KMK KP-23420

Locality: Alekseevka 1 locality, Tavolzhanka River Basin, Borskii District, Samara Region, European Russia.

Horizon: Basal Rybinskian Horizon, Basal Kamennyi Yar Formation.

Biostratigraphy:

Age: Early Triassic.

Material: Incomplete skull.

Referred material:

PIN no. 4513/6: Posterior fragment of a left lower jaw ramus.

 

Locality: Krasnaya Yaruga 1 locality, Orenburg Region, Sorochinskii District, European Russia.

Horizon: Terminal Vokhmian Horizon.

Biostratigraphy:

Age: Early Triassic.

Material:

PIN no. 4495/2: Posteior region of a right lower jaw ramus.

 

Locality: Krasnaya yaruga 2 locality, Sorochinskii District, Orenburg Region, European Russia.

Horizon: Basal Rybinskian Horizon.

Biostratigraphy:

Age: Early Triassic.

Material:

PIN 4495/11: Left lower jaw ramus.

 

Locality: Rodnikovyi Dol Locality: Sorochinskii District, Orenburg Region, European Russia.

Horizon: Basal Rybinskian Horizon.

Biostratigraphy:

Age: Early Triassic.

Material:

PIN 4534/1: Posteior half of a right lower jaw ramus.

 

Locality: Kamennyi Dol locality, Borskii District, Samara Region, European Russia.

Horizon: Basal Rybinskian Horizon.

Biostratigraphy:

Age: Early Triassic.

Material:

PIN 4419/49: Anterior end of a right lower jaw ramus.

 

Locality: Alekseevka 2 locality, Borskii District, Samara Region, European Russia.

Horizon: Basal Rybinskian Horizon.

Biostratigraphy:

Age: Early Triassic.

Material:

PIN 4513/42: Anterior end of a right lower jaw ramus.