Genus: Cyclotosaurus FRAAS, 1889
Etymology:
= Hemprichisaurus KUHN, 1939
Etymology:
= ?Hercynosaurus JAEKEL, 1914
Etymology:
Species: robustus (MEYER & PLIENGER, 1844) FRAAS, 1889
Etymology: Latin, robustus, "robust, strong."
= Mastodonsaurus robustus (MEYER & PLIENGER, 1844) QUENSTEDT, 1850
= Capitosaurus robustus MEYER & PLIENGER, 1844
Lectotype: GPIT PV 30086 (27-1)
Locality: Sandstone quarry in Feuerbacher Haide (former, Kochenhof Quarry, Kilesberg), just north of Stuttgart, northern Baden-Württemburg State, southwestern Germany.
Horizon: Stuttgart Formation (Schilfsandstein, the middle member of the Hauptkeuper, or Middle Keuper).
Biostratigraphy:
Age: Late Carnian Stage, Early Late Triassic Epoch, Late Triassic.
Material: Skull.
Referred material:
SMNS 4139, 4935, 5775, GPIT 27-1 – 27-16, 1801-1802, 1804: Skulls.
SCHOCH & MORENO, 2024
GPIT-PV 44620-44849: Numerous bone fragmetns of the dermal shoulder girdle.
SMNS 4139: Right part of skull.
SMNS 5775: Skull.
SMNS 7414: Posterolateral margin of skull.
SMNS 91417: Interclavicle.
SMNS 91484: Partial interclavicle.
SMNS 91492: PArtial intercalvicle.
Locality: Stutgart-Bopser (Schilfandstein), Stuttgart, northern Baden-Württemburg State, southwestern Germany.
Horizon: Stuttgart Formation (Schilfsandstein, the middle member of the Hauptkeuper, or Middle Keuper).
Biostratigraphy:
Age: Late Carnian Stage, Early Late Triassic Epoch, Late Triassic.
Material:
SMNS 97116: Partial interclavicle.
Locality: Stuttgart, Steinbruch near Waldorsfchule, northern Baden-Württemburg State, southwestern Germany.
Horizon: Stuttgart Formation (Schilfsandstein, the middle member of the Hauptkeuper, or Middle Keuper).
Biostratigraphy:
Age: Late Carnian Stage, Early Late Triassic Epoch, Late Triassic.
Material:
SMNS 91493: Interclavicle.
Locality: Wendelsheim, northern Baden-Württemburg State, southwestern Germany.
Horizon: Stuttgart Formation (Schilfsandstein, the middle member of the Hauptkeuper, or Middle Keuper).
Biostratigraphy:
Age: Late Carnian Stage, Early Late Triassic Epoch, Late Triassic.
Material:
GPIT uncatalogued: Posterolateral portion of skull roof.
CORROY, 1928Locality: Rauax, Lorraine, France.
Horizon: Muschelkalk.
Biostratigraphy:
Age: Mid Triassic Epoch, Middle Triassic.
Material:
Number: Not given: Clavicle.
Locality: Mont-sur-Meurthe Department, France.Horizon: Muschelkalk.
Biostratigraphy:
Age: Mid Triassic Epoch, Middle Triassic.
Material:
Number: Not given: Tooth.
LYDEKKER, 1890
Locality: Feuerbach-Haide, near Stuttgart, Baden-Wurttemberg State, Germany.
Horizon: Middle Keuper.
Biostratigraphy:
Age: Middle Triassic.
Material:
BMNH 37993: Cast of a slab showing the inner lamina of the bones of the cranial roof.
BMNH 37992: Cast of slab showing the palatal aspect of the skull.
BMNH R1439: Cast of slab showing the frontal aspect of muzzle.
BMNH R1439a: Cast of slab showing palatal aspect of the anterior portion of skull.
BMNH R1439b: Cast of slab showing the oral surface of the mandibular symphsis.
BMNH R1439c: Cast of slab with the ventral surface of part of the mandible.
BMNH 33074: Fragmentary skull.
BMNH 14675: Throacic plate.
BMNH 14676, BMNH 15271, BMNH R281: Fragmentary thoracic plates.
BMNH R84: Thoracic plate?
BMNH 43636: Imperfect sculptured bone.
BMNH 14675, BMNH 33073: Sculptured bones.
PATON, 1975
Locality: Stuttgart, Baden-Wurttemberg State, Germany.
Horizon: Upper Keuper.
Biostratigraphy:
Age: Carnian Stage, Lower Late Triassic Epoch, Late Triassic.
Material:
RSM: 2 clavicles and 1 interclavicle.
ZITTEL, 1890Locality:
Horizon:
Biostratigraphy:
Age:
Material:
Number: Not given: Fragmentary skull.
= Mastodonsaurus? robustus STEPHENS, 1886-1887/FLETCHER, 1971
Australian Museum, Sydney.
Locality: Cockatoe Island, Port Jackson, Sydney, New South Wales State, Australia.
Horizon:
Biostratigraphy:
Age: Triassic.
Material:
= Mastodonsaurus cyclotis QUENSTEDT, 1850
Etymology:Holotype:
Locality:
Horizon:
Biostratigraphy:
Age:
Material:
= Mastodonsaurus keuperinus FRAAS, 1880
Etymology:
= Mastodonsaurus keuperinus OCHEV, 1966 (sic)
= Mastodonsaurus kieuperinus OCHEV, 1966 (sic)Holotype: SMNS 4937
Locality: Sandstone quarry in Feuerbacher Haide (former, Kochenhof Quarry, Kilesberg), just north of Stuttgart, northern Baden-Württemburg State, southwestern Germany.
Horizon: Stuttgart Formation (Schilfsandstein, the middle member of the Hauptkeuper, or Middle Keuper).
Biostratigraphy:
Age: Late Carnian Stage, Early Late Triassic Epoch, Late Triassic.
Material: Tip of snout.
Referred material:
SMNS 833: Right palatine and choana.
LYDEKKER, 1890
NHMUK (old BMNH) 15270: Fragmentary left thoracic plate.
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Species: posthumus FRAAS, 1913
Etymology:
Holotype: SMNS 12988
Locality: Sandstone quarry (Weißer Steinbruch), south of Pfaffenhofen, northern Baden-Württemburg State, Stormberg, southwestern Germany.
Horizon: Middle Stubensandstein, km4, Upper Middle Keuper.
Biostratigraphy:
Age: Lower Norian Stage, Lower Mid Triassic Epoch, Middle Triassic.
Material: Fragmentary skull and fragmentary skeleton.
Referred material:
SCHOCH & MORENO, 2024
SMNS 12195: Interclavicle.
SMNS 18417: Partial interclavicle.
SMNS 59769: Parital jaw with teeth.
SMNS 91533: Leftquadratojugal.
Locality: Lippe region of Nordhrein-Westfalen, Germany.
Horizon: Rhaetic.
Biostratigraphy:
Age: Rhaetian Stage, Uppermost Late Triassic Epoch, Late Triassic.
Material:
Number: Not given: Skull and interclavicle.
CORROY, 1928
Locality: Ruaux, Lorraine, France.
Horizon: Muschelkalk.
Biostratigraphy:
Age: Mid Triassic Epoch, Middle Triassic.
Material:
Number: Not given: Clavicle.
= Cyclotosaurus cf. posthumus INGAVAT & JANVIER, 1981
Locality: Outcrop near the Chulabhorn Dam, Kon San Distict Chaiyaphum PRovince, Thailand.
Horizon: Huai Hin Lat Formation.
Biostratigraphy:
Age: Carnian-Norian Stage, Late Triassic Epoch, Late Traissic.
Material:
D.M.R. no Ch.D 001: Posterior part of a skull.
aff posthumus KUHN, 1938
Locality: Halberstadt, Sachsen-Anhalt State, Germany.
Horizon: Keuper Formation.
Biostratigraphy:
Age: Late Triassic.
Material:
Number: Not given: Fragment of skull.
SCHOCH & MORENO, 2024
Locality: Magstadt, Sand Pit, Baden-Württemburg State, Stormberg, southwestern Germany.
Horizon: Middle Stubensandstein, km4, Upper Middle Keuper.
Biostratigraphy:
Age: Lower Norian Stage, Lower Mid Triassic Epoch, Middle Triassic.
Material:
SMNS 50006: Interclavicle.
SMNHS 50009: Preorbital region.
SMNS 50059: Left preorbital skull.
SMNS 50063: Partial skull.
SMNS 51102: Partial skull.
SMNS 51426: Partial large skull.
SMNHS 51436: Interclavicle fragment.
= Cyclotosaurus mordax FRAAS, 1913
Etymology: Latin, mordax, "biting."Holotype: SMNS 13014
Locality: Sandstone quarry (Weißer Steinbruch), between Pfaffenhofen and Ochsenbach and quarry at Magstadt near Sindelfingen, northern Baden-Wurttemberg State, Stormberg, southwestern Germany.
Horizon: Middle Stubensandstein, km4, Upper Middle Keuper.
Biostratigraphy:
Age: Lower Norian Stage, Lower Mid Triassic Epoch, Middle Triassic.
Material: Fragmentary skull.
Referred material:
CORROY, 1928
Locality: Ruaux, Lorraine, France.
Horizon: Lower Muschelkalk.
Biostratigraphy:
Age: Anisian Stage, Lower Mid Triassic Epoch, Middle Triassic.
Material:
Number: Not given: Clavicle.
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Species: ebrachensis KUHN, 1932
Etymology: In reference to Ebrach, Franken, Bayern (Bavaria) State, Germany.
Holotype: SNSB-BSPG 1931 Xi (Bayerische Staatssammlung für Paläontologie und Historische Geologie, Munich).
Note: Destroyed during WWII (DAMIANI, 2001)
Locality: Km3, former sandstone quarry south of Ebrach, Franken, Bayern (Bavaria) State, Germany.
Horizon: Hassberge Formation, Level 5 of Kuhn's section 2.8 m above the base, in the Blasensandstein, Middle Keuper.
Biostratigraphy:
Age: Norian Stage, Middle Late Triassic, Late Triassic.
Material: Skull.
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Species: hemprichi KUHN, 1942
Etymology:
= Paracyclotosaurus hemprichi (KUHN, 1942) KUHN, 1960
Holotype: Städtischen Museum of Halberstadt, uncatalogued.
Locality: Barecke's (Formerly Heine's) clay quarry on the east edge of Halberstadt, Sachsen-Anhalt State, northeastern Germany.
Horizon: Upper Keuper Knollenmergel, km5, Middle Keuper, deepest part of the quarry.
Biostratigraphy:
Age: Upper Norian Stage, Middle Triassic Epoch, Middle Triassic.
Material: Skull.
Syntypes: MB Am 570, 572, 573, 577:
Referred material:
KUHN, 1942
Skull, clavicle, ribs, vertebrae and fragments of limbs.
= ?Hercynosaurus carinidens JAEKEL, 1914
Etymology:
= ?Cyclotosaurus carinidens (JAEKEL, 1914) ROMER, 1947Holotype:
Locality: Halberstadt, Sachsen-Anhalt State, Germany.
Horizon: Keuper.
Biostratigraphy:
Age: Late Triassic.
Material: Tooth, 2 centra and ribs.
Referred material:
KUHN, 1942
Locality: Halberstadt, Sachsen-Anhalt State, Germany.
Horizon: Keuper.
Biostratigraphy:
Age: Late Triassic.
Material:
Number: Not given:
= Hemprichisaurus keuperinus KUHN, 1939
Etymology:Holotype:
Locality:
Horizon:
Biostratigraphy:
Age:
Material:
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Species: intermedius SUEJ & MAJER, 2005
Etymology: The species name was chosen in order to emphasize that it is intermediate
between the older Cyclotosaurus robustus MEYER & PLIENINGER, 1844 and younger
C. mordax FRAAS, 1913
Holotype: ZPAL Ab III 1173
Locality: Krasiejow, Opole Silesia, Silesia Province, Poland.
Horizon: Probably Drawno beds coeval to the Lehrberg Beds of Germany.
Biostratigraphy:
Age: Late Carnian Stage, Lower Late Triassic Epoch, Late Triassic.
Material: Skull and mandible.
Referred material:
ZPAL AB III 471/8, 19, 31, 45, 113, 472/28, AB III 680: Skulls and mandibles.
ZPAL AB III 887/3: Right humerus.
ZPAL AB III 471/51: Right cleithrum.
ZPAL AB III 397: Right clavicle.
ZPAL Ab III 887/1: Interclavicle.
ZPAL Ab III 887/2: Scapulacoracoid.
ZPAL Ab III 471/51: Cleithrum.
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Species: buechneri WITZMANN, SACHS, & NYHUIS, 2016
Etymology: In honor of Dr. Martin, Büchner former director of the Museum of
Natural History in Bielefeld. Martin Büchner discovered the holotype and only
known speciimen in 1976, and has a life-long record of outstanding contributions
to the museum's geological collectioni and the popularization of geosciences in
the region.
= Cyclotosaurus robustus SCHOCH & MILNER, 2000
Holotype: Namu ES/k 36053
Locality: Bielefeld, Sieker District, North Rhine-Westphalia State, Northwestern Germany.
Horizon: Stuttgart Formation.
Biostratigraphy:
Age: Tuvalian, Late Carnian Stage, Lower Late Triassic Epoch, Late Triassic.
Material: Almost complete skull.
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Species: naraserluki MARZOLA, MATEUS, SHUBIN & CLEMENSEN, 2017
Etymology: Greenlandic, naraserluk, "amphibian, salamander."
Holotype: MGUH.VP 9522
Locality: Macknight Bjerg Quarry, 71°22.30'N, 22°33.14'W, Jameson Land, East Greenland.
Horizon: Carlsberg Fjord Beds, Ørsted Dal Member, Fleming Fjord Formation.
Biostratigraphy:
Age: Norian Stage, Late Triassic Epoch, Late Triassic.
Material: Nearly complete skull.
Referred material:
MGUH.VP 9523, MGUH.VP9524 (perviously V-2012-268 and V.2-12-269): 2 vertebral intercentra.