Genus: Dvinosaurus AMALITZKY, 1921
Etymology:
= Dwinosaurus HUENE, 1956 (sic)

Species: primus AMALITZKY, 1921
Etymology:

Lectotype: PIN 2005/43 (39)

Locality: Sokolki, North Divina River, Kotlasskii (Kotlas) District, Arkhangel’sk Province, North European Russia.

Horizon: Tatarian Series, Upper Vyatkian Substage, Vyatkian Horizon (Northern Dvina Horizon, bottom of the Vyatkian Horizon).

Biostratigraphy: Sokolki Faunal Assemblage, Sokolki Faunal Subassemblage, Scutosaurus karpinskii zone (Dicynodon Assemblage zone, LUCAS, 2005), Zone IV.

Age: Upper Tatarian Age, Lopingian Subepoch, Upper Zechstein Epoch, Late Permian.

Material: Skull and anterior portion of a skeleton.

Referred material:

SHISHKIN, 1973

2005/39:

2005/40/42,43:

2005/44:

2005/48:

2005/48b:

2005/48D:

2005/49:

2005/55:

2005/67:

2005/380:

156/10: Fragmentary skull.

156/11:

156/12: Jugal.

156/13: Squamosum.

156/16:

156/17, 18:

156/19a-d:

156/80:

156/81: Squamosum.

156/82: Squamosum.

156/83: Jugal.

156/84: Postparietal.

156/85: Postaorbital.

156/86:

156/87,88: Palatina.

156/190-193: Suprangular.

156/194-198: Dental.

156/199, 200:

156/201: Spleinal.

552/10:

552/11: Transversum.

552/12: Exoccipital.

2358/48:

2358/64:

2358/65: Jugal.

2357-7/36: Stapes.

2357-7/47:

2357-7/81:

2648/1:

2648/2: Dental.

1535: Dental.

 

= Dvinosaurus cf. primus GOLUBEV & BULANOV, 2018

Locality: Sundyr-1 locality, right bank of the Volga River (southern shore of the Cheboksary Reservoir), directly downstream from the mouth of the Sundry River, Gornomariiskii District, Mari El Republic, European Russia.

Horizon:

Biostratigraphy: Sundyr Assemblage, latest Dinocephalian superassemblage, Late Titanophoneus Tetrapod zone.

Age: Lower Upper Substage (Lower Subhorion), Lower Upper Severodvinian Stage (Lower Putyatinian Horizon, Upper Capitanian Stage), Lower Tatarian series (Late Guadalupian Series), Late Permian.

Material:

PIN, no 5388/4: Left jugal.

PIN, no. 5388/21: Right prietal.

PIN, no. 5388/32: Apical prt of tooth.

PIN< no. 5388/47: Left quadratojugal.

IVAKHNENKO, 2008b

Locality: Semin Ovrag Locality, Semin (Syomin) Ovrag Ravine, Cemin Ravin, village of Il’inskoe, right bank of the Volga River, Tetyushi District, Tatarstan, European Russia.

Horizon: Serverodvinian (Dvina) horizon, Semen Formation.

Biostratigraphy: Sokolki Faunal Assemblage, Ilinskoe Faunal Subassemblage.

Age: Upper Tatarian Age, Lopingian Subepoch, upper Zechstein Epoch, Late Permian.

Material: Not given.

 

Locality: Zavarzhye (Zavrazh’e) locality, Malaya Severnaya Dvina River, Kotlasskii (Kotlas) District, Arkhangel’sk Province, North European Russia.

Horizon: Tatarian Series, Vyatkian Substage, Vyatkian Horizon, Vyatka Regional Stage.

Biostratigraphy: Sokolki Faunal Assemblage, Sokolki Faunal Substage (Dicynodon Assemblage zone, LUCAS, 2005).

Age: Upper Tatarian Age, Lopingian Subepoch, upper Zechstein Epoch, Late Permian.

Material: Not given:

 

Locality: Agafonov locality, Viatka River Basin, Kotel’nichskii (Kotelnitch) District) Kirov Province, European Russia.

Horizon: Tatarian Series, Upper Severodvinian Substage, Severodvinian Horizon.

Biostratigraphy: Sokolki Faunal Assemblage, Ilinskoe Faunal Subassemblage.

Age: Upper Tatarian Age, Lopingian Subepoch, upper Zechstein Epoch, Late Permian.

Material:

Number: Not given:

 

NIKITAN, 1995

Locality: Malaya Severnaya Dvina River, Cis-Urals, European Russia.

Horizon:

Biostratigraphy:

Age: Upper Tatarian Age, Lopingian Subepoch, upper Zechstein Epoch, Late Permian.

Material:

PIN Nos., 2005/70, 69, 55d, 52a, 50b, 11054: Humerii.

 

Locality: Volga River, European Russia.

Horizon:

Biostratigraphy:

Age: Upper Tatarian Age, Lopingian Subepoch, upper Zechstein Epoch, Late Permian.

Material:

PIN Nos. 156/36, 35, 34, 33, 32, 31, 30, 29, 28, 202, 203: Humerii.

 

Locality: Sukhona River Basin, Vologda Province, European Russia.

Horizon:

Biostratigraphy:

Age: Upper Tatarian Age, Lopingian Subepoch, upper Zechstein Epoch, Late Permian.

Material:

PIN Nos., 4545/7: Humerus.

 

Locality: Malaya Severnaya Dvina River, Cis-Urals, European Russia.

Horizon:

Biostratigraphy:

Age: Upper Tatarian Age, Lopingian Subepoch, upper Zechstein Epoch, Late Permian.

Material:

PIN No. 2357/51: Humerus.

 

Locality: Bol’shaya Sundyr’River basin, Moscow Syncline, European Russia.

Horizon:

Biostratigraphy:

Age: Upper Tatarian Age, Lopingian Subepoch, upper Zechstein Epoch, Late Permian.

Material:

PIN No 2648/4: Humeri.

 

Locality: Sukhona River, Vologda Province, European Russia..

Horizon:

Biostratigraphy:

Age: Upper Tatarian Age, Lopingian Subepoch, upper Zechstein Epoch, Late Permian.

Material:

PIN 1650/6: Humeri.

 

SUSHKIN, 1936

Locality:

Horizon:

Horizon:

Biostratigraphy:

Age: Upper Tatarian Age, Lopingian Subepoch, upper Zechstein Epoch, Late Permian.

Material:

AM 43/103: Skull and mandibles.

AM 44/33: Right half of mandible.

AM 45/27: Part of right dentary.

AM 46: Part of left dentary.

AM 47: Part of dentary.

AM 49: An incomplete parasphenoid in 3 pieces and a basiventral.

AM 50/55: Part of the vertebral column, right and left humerus, right and left ulna and radius.

AM 51/91: Left humerus of a young individual.

AM 52/110, 1900: Left scapula and left humerus, incomplete, a cleithrum, incomplete, and a pair of ilia.

AM 53/117, 1902: Right scapula.

AM 54/31-32, 1900: Left femur.

AM 55/16, 1923: Roof of the skull, 4 caudal neural arches, 1 caudal hypocentrum, 3 trunk hypocentra and 10 ribs, clavicle, humerus, ulna and radius of the right side, both ischia and left femur.

Number not given: Left manus, incomplete with 4 metacrapals and 3 carpals in situ, and some odd phalanges.

 


TVERDOKHLEBOV, TVERDOKHLEBOVA, MINIKH, SURKOV & BENTON, 2005

Locality: European Russia.

Horizon: Malokinelskaya Svita (Severodvinian Gorizont).

Biostratigraphy:

Age: Upper Tatarian Age, Lopingian Subepoch, upper Zechstein Epoch, Late Permian.

Material:

Number: Not given:

 


Locality: Pronkino (Pron’kino) locality, 1 km SW of Pronkino village, left bank of Borovka River, Samara drainage basin, Sorochinsk District, Orenburg Region, European Russia.

Horizon: Sama Formation, Kutulukskaya Svita (Vyatkian Gorizont).

Biostratigraphy:

Age: Upper Tatarian Age, Vyatkian Regional Stage, Lopingian Subepoch, upper Zechstein Epoch, Late Permian.

Material:

Number: Not given:

 

Locality: Pokrovka, quarry on SW outskirts of Pokrovka village on right bank of Chesnokovka River, Samara drainage basin, Orenburg Region, European Russia.

Horizon: Kutulukskaya Svita (Vyatkian Gorizont).

Biostratigraphy:

Age: Upper Tatarian Age, Lopingian Subepoch, upper Zechstein Epoch, Late Permian.

Material:

Number: Not given: Vomer.

Locality: Babentzevo, 3.8 km NE of Babentzevo village, right slope of nameless gully, right tributary of Konduzla River, Samara drainage basin, Orenburg Region, European Russia.

Horizon: Malokinelskaya Svita, Severodovinian Gorizont.

Biostratigraphy:

Age: Upper Tatarian Age, Lopingian Subepoch, upper Zechstein Epoch, Late Permian.

Material:

Number: Not given:

 

= Dvinosaurus secundus AMALITZKY, 1921
Etymology:

Holotype: Am 40

Locality: Sokolki, North Divina River, Arkhangel’sk Province, Cis-Urals, European Russia.

Horizon:

Biostratigraphy:

Age: Late Permian.

Material: Skull and fragmentary skeleton.

Referred material:

AM 48/51: Roof of the skull, middle part and left side, fore part of the vertebral column in 7 pieces.

 

= Dvinosaurus tertius AMALITZKY, 1921
Etymology:

Holotype: AM 41

Locality: Sokolki, North Divina River, Arkhangel’sk Province, Cis-Urals, European Russia.

Horizon:

Biostratigraphy:

Age: Late Permian.

Material: Fragmentary skull and fragmentary skeleton.

Cotype: AM 42: Skull and mandibles in situ.


_____________________________________________________________________________________

Species: purlensis SHISHKIN, 1968
Etymology:

Holotype: PIN 1538/18

Locality: Purly, Pizhma River, Vyatka basin, Tonshaevskii District, Nizhnii Novgorod Province, European Russia.

Horizon: Vyatkian Horizon, Upper Vyatkian Substage.

Biostratigraphy: Vyazniki Faunal Assemblage.

Age: Upper Tatarian Age, Lopingian Subepoch, upper Zechstein Epoch, Late Permian.

Material: Lower jaw fragments.

_____________________________________________________________________________________
Species: egregius SHISHKIN, 1968
Etymology:

Holotype: PIN 1100/23

Locality: Vyazniki, Klyaz’ma River, Oka basin, Vladimir Province, European Russia.

Horizon:

Biostratigraphy:

Age: Upper Tatarian Age, Lopingian Subepoch, upper Zechstein Epoch, Late Permian.

Material: Skull.

Referred material:

1100/24: Squamosum.

1100/25: Supratemporal.

1100/26: Tabular.

1100/27: Postparietal.

1100/28: Postfrontal.

1100/29: Nasal.

1100/30: Quadratogjugal.

1100/31-34: Quadratojugal.

1100/35-36: Quadratojugal and quadrate.

1100/37: Exoccipital.

1100/38:

1100/40-42:

1100/43:

1100/44, 45:

1100/46: Premaxilla.

1100/48, 49: Palatina.

1100/50: Palatinum.

1100/51: Palatinum, maxillae.

1100/52: Tranversum and maxilla.

1100/53:

1100/54: Dentalia.

1100/56:

1100/57:

1100/58:

1100/59:

1100/60-64:

1100/65: Dental.

1100/70: Postsplenial.

1100/71, 72: Angular.

1100/73-75: Articular and supraangular.

1100/76: Dentale.

 

NIKITIN, 1995

Locality: Klyaz’ma River Basin, European Russia.

Horizon:

Biostratigraphy:

Age: Upper Tatarian Age, Lopingian Subepoch, upper Zechstein Epoch, Late Permian.

Material:

PIN Nos., 1100/148-152: Complete or partly damaged (the later with pathological changes.

PIN No. 1100/153: Proximal fragment of a humerus.

_____________________________________________________________________________________

Species: campbelli GUBIN, 2004
Etymology: In honor of K. E. Campbell, who gave essential help in teh organization of the works on the study of Russian Late Permian tetrapods.

Holotype: PIN 4881/426

Locality: Gorokhovets locality, Gorokhovetskii District, Vladimir Region, European Russia.

Horizon: Vyatka Horizon.

Biostratigraphy:

Age: Upper Tatarian Age, Lopingian Subepoch, upper Zechstein Epoch, Late Permian.

Material: Left ramus of the lower jaw.

Referred material:

PIN 490, 909: Premaxillae.

PIN 410: Ceratobranchial.

PIN 412: Supratemporal.

PIN 413: Premaxilla and vomer.

PIN 420, 958: Ilia.

PIN 429, 431: Occipitals.

PIN 442, 575, 577, 912, 916, 920: Intercentra.

PIN 455-461, 900-901: humeri.

PIN 462, 465, 530, 929: Lower jaw fragments.

PIN 502: Postparietal.

PIN 509: Nasal.

PIN 515: Parasphenoid.

PIN 516: Parietal.

PIN 518: Jugal.

PIN 667: Scapulocoracoid.

PIN 678, 683: Femora.

PIN 885: Clavical.

PIN 887: Vomer.

PIN 889: Palatine.

PIN 897: Ectopterygoid.

PIN 906: Interclavicle.

PIN 945: Quadratojugal.

PIN 948: Squamosal.

PIN 949: Frontal.