Species: primus AMALITZKY, 1921
Etymology: Latin, primus, "primer, first."
Lectotype: PIN 2005/43 (39)
Locality: Sokolki, North Divina River, Kotlasskii (Kotlas) District, Arkhangel’sk Province, North European Russia.
Horizon: Tatarian Series, Upper Vyatkian Substage, Vyatkian Horizon (Northern Dvina Horizon, bottom of the Vyatkian Horizon).
Biostratigraphy: Sokolki Faunal Assemblage, Sokolki Faunal Subassemblage, Scutosaurus karpinskii zone (Dicynodon Assemblage zone, LUCAS, 2005), Zone IV.
Age: Upper Tatarian Age, Lopingian Subepoch, Upper Zechstein Epoch, Late Permian.
Material: Skull and anterior portion of a skeleton.
Referred material:
SHISHKIN, 1973
2005/39:
2005/40/42,43:
2005/44:
2005/48:
2005/48b:
2005/48D:
2005/49:
2005/55:
2005/67:
2005/380:
156/10: Fragmentary skull.
156/11:
156/12: Jugal.
156/13: Squamosum.
156/16:
156/17, 18:
156/19a-d:
156/80:
156/81: Squamosum.
156/82: Squamosum.
156/83: Jugal.
156/84: Postparietal.
156/85: Postaorbital.
156/86:
156/87,88: Palatina.
156/190-193: Suprangular.
156/194-198: Dental.
156/199, 200:
156/201: Spleinal.
552/10:
552/11: Transversum.
552/12: Exoccipital.
2358/48:
2358/64:
2358/65: Jugal.
2357-7/36: Stapes.
2357-7/47:
2357-7/81:
2648/1:
2648/2: Dental.
1535: Dental.
IVAKHNENKO, 2008b
Locality: Semin Ovrag Locality, Semin (Syomin) Ovrag Ravine, Cemin Ravin, village of Il’inskoe, right bank of the Volga River, Tetyushi District, Tatarstan, European Russia.
Horizon: Serverodvinian (Dvina) horizon, Semen Formation.
Biostratigraphy: Sokolki Faunal Assemblage, Ilinskoe Faunal Subassemblage.
Age: Upper Tatarian Age, Lopingian Subepoch, upper Zechstein Epoch, Late Permian.
Material: Not given.
Locality: Zavarzhye (Zavrazh’e) locality, Malaya Severnaya Dvina River, Kotlasskii (Kotlas) District, Arkhangel’sk Province, North European Russia.
Horizon: Tatarian Series, Vyatkian Substage, Vyatkian Horizon, Vyatka Regional Stage.
Biostratigraphy: Sokolki Faunal Assemblage, Sokolki Faunal Substage (Dicynodon Assemblage zone, LUCAS, 2005).
Age: Upper Tatarian Age, Lopingian Subepoch, upper Zechstein Epoch, Late Permian.
Material: Not given:
Locality: Agafonov locality, Viatka River Basin, Kotel’nichskii (Kotelnitch) District) Kirov Province, European Russia.
Horizon: Tatarian Series, Upper Severodvinian Substage, Severodvinian Horizon.
Biostratigraphy: Sokolki Faunal Assemblage, Ilinskoe Faunal Subassemblage.
Age: Upper Tatarian Age, Lopingian Subepoch, upper Zechstein Epoch, Late Permian.
Material:
Number: Not given:
NIKITAN, 1995
Locality: Malaya Severnaya Dvina River, Cis-Urals, European Russia.
Horizon:
Biostratigraphy:
Age: Upper Tatarian Age, Lopingian Subepoch, upper Zechstein Epoch, Late Permian.
Material:
PIN Nos., 2005/70, 69, 55d, 52a, 50b, 11054: Humerii.
Locality: Volga River, European Russia.
Horizon:
Biostratigraphy:
Age: Upper Tatarian Age, Lopingian Subepoch, upper Zechstein Epoch, Late Permian.
Material:
PIN Nos. 156/36, 35, 34, 33, 32, 31, 30, 29, 28, 202, 203: Humerii.
Locality: Sukhona River Basin, Vologda Province, European Russia.
Horizon:
Biostratigraphy:
Age: Upper Tatarian Age, Lopingian Subepoch, upper Zechstein Epoch, Late Permian.
Material:
PIN Nos., 4545/7: Humerus.
Locality: Malaya Severnaya Dvina River, Cis-Urals, European Russia.
Horizon:
Biostratigraphy:
Age: Upper Tatarian Age, Lopingian Subepoch, upper Zechstein Epoch, Late Permian.
Material:
PIN No. 2357/51: Humerus.
Locality: Bol’shaya Sundyr’River basin, Moscow Syncline, European Russia.
Horizon:
Biostratigraphy:
Age: Upper Tatarian Age, Lopingian Subepoch, upper Zechstein Epoch, Late Permian.
Material:
PIN No 2648/4: Humeri.
Locality: Sukhona River, Vologda Province, European Russia..
Horizon:
Biostratigraphy:
Age: Upper Tatarian Age, Lopingian Subepoch, upper Zechstein Epoch, Late Permian.
Material:
PIN 1650/6: Humeri.
SUSHKIN, 1936
Locality:
Horizon:
Horizon:
Biostratigraphy:
Age: Upper Tatarian Age, Lopingian Subepoch, upper Zechstein Epoch, Late Permian.
Material:
AM 43/103: Skull and mandibles.
AM 44/33: Right half of mandible.
AM 45/27: Part of right dentary.
AM 46: Part of left dentary.
AM 47: Part of dentary.
AM 49: An incomplete parasphenoid in 3 pieces and a basiventral.
AM 50/55: Part of the vertebral column, right and left humerus, right and left ulna and radius.
AM 51/91: Left humerus of a young individual.
AM 52/110, 1900: Left scapula and left humerus, incomplete, a cleithrum, incomplete, and a pair of ilia.
AM 53/117, 1902: Right scapula.
AM 54/31-32, 1900: Left femur.
AM 55/16, 1923: Roof of the skull, 4 caudal neural arches, 1 caudal hypocentrum, 3 trunk hypocentra and 10 ribs, clavicle, humerus, ulna and radius of the right side, both ischia and left femur.
Number not given: Left manus, incomplete with 4 metacrapals and 3 carpals in situ, and some odd phalanges.
TVERDOKHLEBOV, TVERDOKHLEBOVA, MINIKH, SURKOV & BENTON, 2005Locality: European Russia.
Horizon: Malokinelskaya Svita (Severodvinian Gorizont).
Biostratigraphy:
Age: Upper Tatarian Age, Lopingian Subepoch, upper Zechstein Epoch, Late Permian.
Material:
Number: Not given:
Locality: Pronkino (Pron’kino) locality, 1 km SW of Pronkino village, left bank of Borovka River, Samara drainage basin, Sorochinsk District, Orenburg Region, European Russia.Horizon: Sama Formation, Kutulukskaya Svita (Vyatkian Gorizont).
Biostratigraphy:
Age: Upper Tatarian Age, Vyatkian Regional Stage, Lopingian Subepoch, upper Zechstein Epoch, Late Permian.
Material:
Number: Not given:
Locality: Pokrovka, quarry on SW outskirts of Pokrovka village on right bank of Chesnokovka River, Samara drainage basin, Orenburg Region, European Russia.
Horizon: Kutulukskaya Svita (Vyatkian Gorizont).
Biostratigraphy:
Age: Upper Tatarian Age, Lopingian Subepoch, upper Zechstein Epoch, Late Permian.
Material:
Number: Not given: Vomer.
Locality: Babentzevo, 3.8 km NE of Babentzevo village, right slope of nameless gully, right tributary of Konduzla River, Samara drainage basin, Orenburg Region, European Russia.
Horizon: Malokinelskaya Svita, Severodovinian Gorizont.
Biostratigraphy:
Age: Upper Tatarian Age, Lopingian Subepoch, upper Zechstein Epoch, Late Permian.
Material:
Number: Not given:
= Dvinosaurus secundus AMALITZKY, 1921
Etymology:Holotype: Am 40
Locality: Sokolki, North Divina River, Arkhangel’sk Province, Cis-Urals, European Russia.
Horizon:
Biostratigraphy:
Age: Late Permian.
Material: Skull and fragmentary skeleton.
Referred material:
AM 48/51: Roof of the skull, middle part and left side, fore part of the vertebral column in 7 pieces.
= Dvinosaurus tertius AMALITZKY, 1921
Etymology:Holotype: AM 41
Locality: Sokolki, North Divina River, Arkhangel’sk Province, Cis-Urals, European Russia.
Horizon:
Biostratigraphy:
Age: Late Permian.
Material: Fragmentary skull and fragmentary skeleton.
Cotype: AM 42: Skull and mandibles in situ.
_____________________________________________________________________________________
Species: purlensis SHISHKIN, 1968
Etymology: In reference to Purly, Pizhma River, Vyatka basin, Tonshaevskii District, Nizhnii
Novgorod Province, European Russia.
Holotype: PIN 1538/18
Locality: Purly, Pizhma River, Vyatka basin, Tonshaevskii District, Nizhnii Novgorod Province, European Russia.
Horizon: Vyatkian Horizon, Upper Vyatkian Substage.
Biostratigraphy: Vyazniki Faunal Assemblage.
Age: Upper Tatarian Age, Lopingian Subepoch, upper Zechstein Epoch, Late Permian.
Material: Lower jaw fragments.
_____________________________________________________________________________________
Species: egregius SHISHKIN, 1968
Etymology:
Holotype: PIN 1100/23
Locality: Vyazniki, Klyaz’ma River, Oka basin, Vladimir Province, European Russia.
Horizon:
Biostratigraphy:
Age: Upper Tatarian Age, Lopingian Subepoch, upper Zechstein Epoch, Late Permian.
Material: Skull.
Referred material:
1100/24: Squamosum.
1100/25: Supratemporal.
1100/26: Tabular.
1100/27: Postparietal.
1100/28: Postfrontal.
1100/29: Nasal.
1100/30: Quadratogjugal.
1100/31-34: Quadratojugal.
1100/35-36: Quadratojugal and quadrate.
1100/37: Exoccipital.
1100/38:
1100/40-42:
1100/43:
1100/44, 45:
1100/46: Premaxilla.
1100/48, 49: Palatina.
1100/50: Palatinum.
1100/51: Palatinum, maxillae.
1100/52: Tranversum and maxilla.
1100/53:
1100/54: Dentalia.
1100/56:
1100/57:
1100/58:
1100/59:
1100/60-64:
1100/65: Dental.
1100/70: Postsplenial.
1100/71, 72: Angular.
1100/73-75: Articular and supraangular.
1100/76: Dentale.
NIKITIN, 1995
Locality: Klyaz’ma River Basin, European Russia.
Horizon:
Biostratigraphy:
Age: Upper Tatarian Age, Lopingian Subepoch, upper Zechstein Epoch, Late Permian.
Material:
PIN Nos., 1100/148-152: Complete or partly damaged (the later with pathological changes.
PIN No. 1100/153: Proximal fragment of a humerus.
_____________________________________________________________________________________
Species: campbelli GUBIN, 2004
Etymology: In honor of K. E. Campbell, who gave essential help in teh organization
of the works on the study of Russian Late Permian tetrapods.
Holotype: PIN 4881/426
Locality: Gorokhovets locality, Gorokhovetskii District, Vladimir Region, European Russia.
Horizon: Vyatka Horizon.
Biostratigraphy:
Age: Upper Tatarian Age, Lopingian Subepoch, upper Zechstein Epoch, Late Permian.
Material: Left ramus of the lower jaw.
Referred material:
PIN 490, 909: Premaxillae.
PIN 410: Ceratobranchial.
PIN 412: Supratemporal.
PIN 413: Premaxilla and vomer.
PIN 420, 958: Ilia.
PIN 429, 431: Occipitals.
PIN 442, 575, 577, 912, 916, 920: Intercentra.
PIN 455-461, 900-901: humeri.
PIN 462, 465, 530, 929: Lower jaw fragments.
PIN 502: Postparietal.
PIN 509: Nasal.
PIN 515: Parasphenoid.
PIN 516: Parietal.
PIN 518: Jugal.
PIN 667: Scapulocoracoid.
PIN 678, 683: Femora.
PIN 885: Clavical.
PIN 887: Vomer.
PIN 889: Palatine.
PIN 897: Ectopterygoid.
PIN 906: Interclavicle.
PIN 945: Quadratojugal.
PIN 948: Squamosal.
PIN 949: Frontal.
_____________________________________________________________________________________
Species: gubini ULIAKHIN & GOLUBEV, 2024
Etymology: In honor of Yu. M. Gubin (1950-2018), who specialized in Permina temnospondyl amphibians.
= Dvinosaurus sp BULANOV & GOLUBEV, 2011
= Dvinosaurus cf. primus GOLUBEV & BULANOV, 2018
= Dvinosaurus aff. primus SUCHKNOVA & GOLUBEV, 2019
Holotype: PIN, n. 5388/445
Locality: Sundyr-1 locality, right bank of the Volga River (southern shore of the Cheboksary Reservoir), directly downstream from the mouth of the Sundry River, Gornomariiskii District, Mari El Republic, European Russia.
Horizon:
Biostratigraphy: Sundyr Assemblage, latest Dinocephalian superassemblage, Late Titanophoneus Tetrapod zone.
Age: Lower Upper Substage (Lower Subhorion), Lower Upper Severodvinian Stage (Lower Putyatinian Horizon, Upper Capitanian Stage), Lower Tatarian series (Late Guadalupian Series), Late Permian.
Material: Right detnary.
Referred material:
PIN, no 5388/4, 48, 439: Zygomatic.
PIN, no. 5388/21: Right prietal.
PIN, no. 5388/32: Apical prt of tooth.
PIN, no. 5388/45, 393, 395, 397, 399, 404, 409, 411, 412, 413, 414, 443, 444, 447, 454, 455, 474, 475, 478, 480, 493: Teeth.:
PIN, no. 5388/46, 396, 398, 403, 405, 415, 441, 472, 476, 513: Clavicles.
PIN, no. 5388/47, 402: Left quadratojugals.
PIN, no. 5388/203, 407, 479: Squamosal.
PIN, no. 5388/223, 323, 473, 545: hypocentra.
PIN, no. 5388/324, 392, 452, 492: Supratemporals.
PIN, no. 5388/388, 391, 450:Parasphenoids with basisphenoids.
PIN, no. 5388/394, 300, 410: Maxilary bones.
PIN, no. 5388/440, 463, 464, 465, 477, 544: interclavicles.
PIN, no. 5388/452: Lacrimal.
PIN, no. 5388/453: Postorbital.
PIN, no. 5388/456: vomer.
PIN, no. 5388/457: Occipital.
PIN, no. 5388/458, 459, 460, 461, 462: Ilia.
PIN, no. 5388/466, 467, 467, 469: Humeri.
PIN, no. 5388/470, 471, 491: Frontals.
PIN, no. 5388/481: Nasal bone.
PIN, no. 5388/490: Transverse.
PIN, no. 5388/546, 547: Pleurocentra.