Species: ? FREDE, SUMIDA & BERMAN, 1993

Holotype: Not given.

Locality: Valley of the Gods (i), NE ¼, NW ¼, sec. 29, T40S, R19E, San Juan County, Utah.

Horizon: Halgaito Shale, equivalent of Lower Cutler Formation.

Biostratigraphy:

Age: Virgilian, upper Stephanian, Upper Silesian Series, Gzelian Epoch, Late Pennsylvanian (Late Carboniferous).

Material:
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Sp

BERMAN, 1993

Locality: Jemez Springs Locality, 10 km north to about 15 km south of Jemez Springs, Sandoval County, New Mexico.

Horizon: Lower Cutler Formation.

Biostratigraphy:

Age: Virgilian, Gzelian Epoch, Late Pennsylvanian.

Material:

Number: Not given.

 

Locality: Pecos River Valley Locality, drainage from Glorieta Pass at Glorieta on the eastern border of Santa Fe County, southeastward to about 8 km South of Ribera in San Miguel County, New Mexico.

Horizon: Sangre de Cristo Formation.

Biostratigraphy:

Age: Probably Missourian, Lower Stephanian, upper Silesian Stage, Kasimovian Epoch? Late Pennsylvanian (Late Carboniferous).

Material:

Number: Not given:

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CASE, 1908

Locality: Pitcairn, Alleghany County, Pennsylvania.

Horizon: Round Knob Formation, Conemaugh Group.

Biostratigraphy:

Age: Late Pennsylvanian.

Material:

Number: Not given: A nearly complete vertebra, pleurocentra and 2 ribs, somewhat distorted.

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CASE, 1915

Locality: Brier Creek bone bed, on the ranch of Mr. Estell, about 12 miles south of Dundee, a little east of the road between Dundee and Archer City, Archer County, Texas.

Horizon:

Biostratigraphy:

Age: Early Permian.

Material:

UCMP 3432: Femur of a juvenile.

UCMP 3406: Interclavicle.

UCMP 3385: Clavicle.

UCMP 3408: Cleithrum.

UCMP 3316: Humerus of a juvenile.

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DEMPSTER, 1935

Locality:

Horizon:

Biostratigraphy:

Age:

Material:

UCMP 3060: Skull.

UCMP 3000: Posterior portion of skull.

UCMP 14408: Posterior portion of skull.

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HARRIS, LUCAS, BERMAN & HENRICI, 2004

Locality: NMMNH Locality 4509, Lucero uplift, Carrizo Arroyo, Valencia County, New Mexico.

Horizon: Red Tanks Member, Bursum Formation.

Biostratigraphy:

Age: Upper Virgilian, upper Stephanian, Upper Silesian Series, Gzelian Epoch, Late Pennsylvanian (Late Carboniferous).

Material:

NMMNH P-40430: 4 articulated vertebrae with attached dorsal portion of right scapular blade, 3 segments of articulated central elements, associated neural spines, associated rib fragments, dorsal portion of left scapular blade, left coracoid, left radius, proximal left ulna, associated mesapodials, and associated indeterminate bone fragments.

NMMNH P-41900: Left and right scapulae (?).


Locality: NMMNH Locality 3423, Lucero uplift, Carrizo Arroyo, Valencia County, New Mexico.

Horizon: Red Tanks Member, Bursum Formation.

Biostratigraphy:

Age: Upper Virgilian, upper Stephanian, Upper Silesian Series, Gzelian Epoch, Late Pennsylvanian (Late Carboniferous).

Material:

CM 70381: Ilium.

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JOHNSON, 1992

Locality: Archer City Bone Bed 3, Archer County, Texas.

Horizon: Upper Archer City Formation (formerly Putnam Formation, Wichita Group), upper Bowie Group.

Biostratigraphy: Land-vertebrate faunachron upper A.

Age: Wolfcampanian stage, Lower Rotliegendes Epoch, Early Permian.

Material:

Number: Not given:

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JOHNSON, 2012

Locality: Conner Ranch Bonebed (Conner Ranch Shark Tooth Bed, SMU locality 462), near Romer’s Cottonwood Creek Locality, Archer County, Texas.

Horizon: Upper Archer City Formation (formerly Putnam Formation, Wichita Group), upper Bowie Group.

Biostratigraphy: Land-vertebrate faunachron upper A.

Age: Wolfcampanian stage, Lower Rotliegendes Epoch, Early Permian.

Material:

SMU 69457, 69458, 76688:

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KISSLER & LEHMAN, 2000, KISSLER & LEHMAN, 2002

Locality: OMNH locality V1005, Seminole County, Oklahoma.

Horizon: Ada Formation.

Biostratigraphy:

Age: Late Pennsylvanian.

Material:

OMNH 55361: Neural arch.

OMNH 55362: Intercentrum.

OMNH 55360: Dentary with associated skull? fragments.

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LANGSTON, 1952

Locality: Harvard locality, 4.5 miles south of Jamez Springs, Sandoval County, New Mexico.

Horizon: Abo Formation.

Biostratigraphy:

Age: Wolfcampian Stage, Lower Rotliegendes Epoch, Early Permian.

Material:

Number: Not given: Fragmentary skull.

Note: Found underneath the fragmentary Embolomerous amphibian skeleton UCMP 39179.

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LANGSTON & REIZ, 1981

Locality: UCMP Loc. v-2813, Camp Quarry, on the southeast slope of a small butte called Loma Salazar, 440 m South of New Mexico State Highway 96, about 840 m southeast of the Rio Puerco bridge at Arroyo
del Agua, section 8, T22N, R3E, Rio Arriba County, New Mexico.

Horizon: Abo/Cutler Formation.

Biostratigraphy:

Age: Wolfcampian, Lower Rotliegendes Epoch, Early Permian.

Material:

Number: Not given:

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LUCAS, RINEHART & HUNT, 2005, WERNEBURG, LUCAS, SCHNEIDER & RINEHART, 2010

Locality: NMMNH locality 6121, sec. 18, T24N, R6E, El Cobre Canyon, Rio Arribia County, New Mexico.

Horizon: Lower El Cobre Canyon Formation, Cutler Group.

Biostratigraphy:

Age: Cobrean, Gzelian Epoch, Early Late Pennsylvanian.

Material:

NMMNH P-46379: Fragmentary skull.

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OLSON, 1936

Locality:

Horizon:

Biostratigraphy:

Age:

Material:

MCZ 1542: Sacral rib.

MCZ 460: Ilium.

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OLSON, 1967

Locality: East Manitou site, SW 1/4, T1N, R16W, Tillman County, Oklahoma.

Horizon:

Biostratigraphy:

Age: Early Permian.

Material:

Number: Not given: Fragments.

 

Locality: South Pauls Valley Site, SW 1/4, sec. 19, T3N, R1E, Garvin County, Oklahoma.

Horizon:

Biostratigraphy:

Age: Early Permian.

Material:

OUSM 2-12-S1: Scapula, and rib fragments.

 

Locality: Villines' site south of Rosedale, Oklahoma.

Horizon:

Biostratigraphy:

Age: Early Permian.

Material:

OUSM 2-40-S4: Fragments of skull and vertebrae.

 

Locality: Orlando Site, 2 miles northeast of Orlando, SE 1/4, sec. 31, T20N, R1W, 2 miles northeast of Orlando in Noble County, Oklahoma.

Horizon: Wellington Formation (Garber Formation), Summer Group, Arbuckle limestone, (Probably equivalent to the Arroyo Formation of Texas).

Biostratigraphy:

Age: Leonardian Stage, Lower Rotliegendes Epoch, Early Permian.

Material:

OUSM 2-1-S38: Skull fragment.

 

Locality: Lucien Site 1, NE 1/4, NE 1/4, sec. 11, T21N, R3W, about 4 miles north of Lucien, Noble County, Oklahoma.

Horizon: Wellington Formation (Garber Formation), Summer Group, Arbuckle limestone, (Probably equivalent to the Arroyo Formation of Texas).

Biostratigraphy:

Age: Leonardian Stage, Lower Rotliegendes Epoch, Early Permian.

Material:

OUSM 2-7-S3: Intercentrum.


Locality: Lucine Site 3, NW 1/4, SW 1/4, sec. 18, T21N, R2W, Oklahoma.

Horizon: Wellington Formation (Garber Formation), Summer Group, Arbuckle limestone, (Probably equivalent to the Arroyo Formation of Texas).

Biostratigraphy:

Age: Leonardian Stage, Lower Rotliegendes Epoch, Early Permian.

Material:

OUSM 2-4-S4: Partial jaw and skull fragment.

 

Locality: Perry Site 3, south of Perry, SW 1/4, SE 1/4, sec. 15, T20N, R1W, Noble County, Oklahoma.

Horizon:

Biostratigraphy:

Age: Early Permian.

Material:

OUSM 2-1-S32: Fragment of skull.

 

Locality: South Covington Site, Garfield County, Oklahoma.

Horizon:

Biostratigraphy:

Age: Early Permian.

Material:

OUSM unnumbered


Locality: Pond Creek Site, NE 1/4, NW 1/4, sec. 14, T25N, R5W, about 5 miles east of the village of Pond Creek, Grant County, Oklahoma.

Horizon: Wellington Formation (Garber Formation), Summer Group, Arbuckle limestone, (Probably equivalent to the Arroyo Formation of Texas).

Biostratigraphy:

Age: Leonardian Stage, Lower Rotliegendes Epoch, Early Permian.

Material:

OUSM 2-0-S18: Skull fragments.

OUSM 2-6-S7: Large tooth.

OUSM 2-10-S1: Intercentrum.

OUSM 2-10-S2: Caudal intercentrum.

OUSM 2-16-S2: Large tooth.

 

Locality: Lucien Site 4, NW 1/4, sec. 19, T21N, R2W, Noble County, Oklahoma.

Horizon: Wellington Formation (Garber Formation), Summer Group, Arbuckle limestone, (Probably equivalent to the Arroyo Formation of Texas).

Biostratigraphy:

Age: Leonardian Stage, Lower Rotliegendes Epoch, Early Permian.

Material:

OUSM 2-7-S4: Intercentrum.

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Eryops? OLSON, 1972

Locality: "Diplocaulus site", FA, Foard County, Texas.

Horizon: Vale Formation, Middle Clear Fork Group.

Biostratigraphy: Land-vertebrate faunachron lower to middle D.

Age: Upper Leonardian Stage,Upper Rotliegendes Epoch, Early Permian.

Material:

UR 171: Femur.

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OLSON, 1991

Locality: Fort Sill, OMNH Locality V51, Dolese Brothers limestone quarry at Richards Spur, Fissure Fills, just west of U.S. Highways 62 & 281, 10.5 miles south of Apache, 6 miles north of Fort Sill, SW 1/2, Sec. 31, T4N, R11W, Comanche County, Oklahoma.

Horizon: Wellington Formation (Garber Formation), Summer Group, Arbuckle limestone, (Probably equivalent to the Arroyo Formation of Texas).

Biostratigraphy:

Age: Leonardian Stage, Lower Rotliegendes Epoch, Early Permian.

Material:

UCLA 1751: Skull fragment, Medio-posterior margin at base.

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ROMER & WITTER, 1943

Locality: Near Fulda, Baylor County, Texas.

Horizon: Petrolia Formation (formerly Belle Plains Formation, middle Wichita Group), Lower Wichita Group.

Biostratigraphy: Land-vertebrate faunachron upper B.

Age: Upper Wolfcampanian stage, Upper Rotliegendes Epoch, Early Permian.

Material:

MCZ 1539: A nearly complete articulated skeleton with skin impressions.

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ROMER & WITTER, 1942

Locality: Terrapin School, Archer County, Texas.

Horizon: Lower Archer City Formation (formerly Moran Formation, Lower Wichita Group), middle Bowie Group.

Biostratigraphy: Land-vertebrate faunachron middle A.

Age: Lower Wolfcampanian stage, Lower Rotilegendes Epoch, Early Permian.

Material:

Number: Not given: Remains mixed in with the type of Edops craigi.

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ROWLAND, LUCAS, KUES & ESTEP, 1997

Locality: New Mexico.

Horizon: Madera Formation, Red Tanks Member.

Biostratigraphy:

Age: Virgilian, Gzelian Epoch, Late Pennsylvanian (Late Late Carboniferous).

Material:

Number: Not given: Interentra and a partial neural spine.

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SANDER, 1987

Locality: Geraldine Bonebed, Archer County, Texas.

Horizon: Nocona Formation (formerly lower to lower upper Admiral Formation, middle Wichita Group), Lower Wichita Group.

Biostratigraphy: Land-vertebrate faunachron lower B.

Age: Upper Wolfcampanian stage, Upper Rotliegendes Epoch, Early Permian.

Material:

MCZ 1903: Small skull.

MCZ 188: Partial skeleton.

MCZ 1885: Skeleton.

MCZ 1900: Skeleton.

Skeleton, skeleton, hind leg, jaws, skull, string of vertebrae, girdle and limb bones, several jaws, miscellaneous small bones, disartuclated skull, skull, disartuculated, skeleton, well articulated skeleton at Yale University, skull and miscellaneous bones.

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SUMIDA, WALLISER & LOMBARD, 1999

Locality: John’s Canyon (iii), NE ¼, NE ¼, sec. 34, T40S, R18E, Utah.

Horizon: Halgaito Shale, equivalent of Lower Cutler Formation.

Biostratigraphy:

Age: Virgilian, upper Stephanian, Upper Silesian Series, Gzelian Epoch, Late Pennsylvanian (Late Carboniferous).

Material:

CMNH


Locality: Lisbon Valley, SE ¼, SE ¼, sec. 30, T30S, R25E, about 12 miles south-southeast of the town of La Sal, San Juan County, Utah.

Horizon: Halgaito Shale, equivalent of Lower Cutler Formation.

Biostratigraphy:

Age: Virgilian, upper Stephanian, Upper Silesian Series, Gzelian Epoch, Late Pennsylvanian (Late Carboniferous).

Material:

UCLA VP 1668:


Locality: Valley of the Gods 1997, NE ¼, Sec. 30, T40S, R19E, San Juan County, Utah.

Horizon: Halgaito Shale, equivalent of Lower Cutler Formation.

Biostratigraphy:

Age: Virgilian, upper Stephanian, Upper Silesian Series, Gzelian Epoch, Late Pennsylvanian (Late Carboniferous).

Material:

CSUSB VP:


Locality: East of Canyonlands, Utah.

Horizon: Cedar Mesa Sandstone.

Biostratigraphy:

Age: Early Permian.

Material:

CSUSB VP

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VAUGHN, 1964

Locality: East of Mitchell Butte, Monument Valley, Navajo County, Arizona.

Horizon: Orange Rock Shale.

Biostratigraphy:

Age: Early Permian.

Material:

UCLA VP 1642: A portion of neural spine.

UCLA VP 1643: Fragment of lower jaw.

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VAUGHN, 1965

Locality: Lisbon Valley, SE ¼, SE ¼, sec. 30, T30S, R25E, about 12 miles south-southeast of the town of La Sal, San Juan County, Utah.

Horizon: Halgaito Shale, equivalent of Lower Cutler Formation.

Biostratigraphy:

Age: Virgilian, Gzelian Epoch, Late Pennsylvanian.

Material:

UCLA VP 1668: Nasal bone.

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VAUGHN, 1968

Locality: Pennsylvania.

Horizon: Dunkard.

Biostratigraphy:

Age: Early Permian.

Material:

USNM 21860: Fragmentary skull.
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VAUGHN, 1969

Locality: Domingo Canyon locality NW 1/4, Sec. 6, T15S, R11E, Otero County, New Mexico.

Horizon: Abo Formation.

Biostratigraphy:

Age: Wolfcampian Stage, Lower Rotliegendes Epoch, Early Permian.

Material:

UCLA VP 1727: Fragmentary skull.

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WERNEBURG, LUCAS, SCHNEIDER & RINEHART, 2010

Locality: Canon del Cobre, Rio Arriba County, New Mexico.

Horizon: El Cobre Canyon Formation, Cutler Group.

Bisotratigraphy:

Age: Cobrean, Gzelian Epoch, Early Late Pennsylvanian.

Material:

FMNH UC 751: Fragmentary skull.