Genus: Trimerorhachis COPE, 1878
Etymology:

Species: insignis COPE, 1878
Etymology:

Holotype: AMNH 4565

Locality: Locality unknown?, "Wichita Basin', Texas.

Horizon: Nocona or Petrolia Formation, Wichita Group.

Biostratigraphy:

Age: Lower Permian.

Material: A partial skeleton including a palatine, a parasphenoid and occiput, vertebrae, a scapula and a humerus.

WILLISTON,

Paratype: AMNH 4557

Locality: Corn Hill (originally called Tit Mountain), Archer County, Texas.

Horizon: Petrolia Formation, Lower Wichita Group.

Biostratigraphy: Land-vertebrate faunachron upper B.

Age: Upper Wolfcampanian stage, Upper Rotliegendes Epoch, Early Permian.

Material: Skull with no parasphenoid.

Referred material:

AMNH 4558: Partial mandible.

 

CASE, 1935

Locality: A few miles south and east of Dundee, Archer County, Texas.

Horizon:

Biostratigraphy:

Age: Early Permian.

Material:

15594, 16009A, 16004, 16005, 16009B, 16009C, 16009D: Skulls and fragmentary skeletons.

plus 9 more skulls and fragmentary skeletons.

 

HUENE, 1913

Locality: Slippery Creek, Wichita Basin, Archer County, Texas.

Horizon: Petrolia Formation, Lower Wichita Group.

Biostratigraphy: Land-vertebrate faunachron upper B.

Age: Upper Wolfcampanian stage, Upper Rotliegendes Epoch, Early Permian.

Material:

AMNH 4570: Skull and dentary missing tip of snout.

 

 

MAY & HALL, 2016

Locality: Billings 4, SNOMNH site, Nobel County, Northern Oklahoma.

Horizon: Midco Member, Willington Formation.

Biostratigraphy:

Age: Leonardian Stage, Lower Rotliegendes Epoch, Early Permian.

Material:

Number: Not given.

 

OLSON, 1967

Locality: Mc Cann Quarry Site, the quarry is on Deer Creek, 8 miles west and 4 miles south of Blackwell (about 9 miles southwest in a direct line) and 2.5 miles northeast of Eddy, NE 1/4, sec. 16 and SW 1/4, sec. 9, T26N, R2W, in Kay County, Oklahoma.

Horizon: Wellington Formation (Garber Formation), Summer Group, Arbuckle limestone, (Probably equivalent to the Arroyo Formation of Texas).

Biostratigraphy:

Age: Leonardian Stage, Lower Rotliegendes Epoch, Early Permian.

Material:

OUSM 2-1-S11: Well-preserved skull.

OUSM 2-1-S15: Clavicle.

OUSM 2-1-S16: Jaw.

OUSM 2-1-S23: Large lower jaw.

OUSM 2-1-S24: Lower jaw.

OUSM 2-1-S25: Fragmentary skull, lower jaw, clavicle and scrap.

OUSM 2-1-S26: Miscellaneous fragments.

OUSM 2-1-S28: Well-preserved half skull.

OUSM 2-34-S1: Girdle fragment.


Locality: Pond Creek Site, NE 1/4, NW 1/4, sec. 14, T25N, R5W, about 5 miles east of the village of Pond Creek, Grant County, Oklahoma.

Horizon: Wellington Formation (Garber Formation), Summer Group, Arbuckle limestone, (Probably equivalent to the Arroyo Formation of Texas).

Biostratigraphy:

Age: Leonardian Stage, Lower Rotliegendes Epoch, Early Permian.

Material:

OUSM 2-6-S3: Intercentrum and palatal tooth.

 

MILNER & SCHOCH, 2013

Locality: Rattlesnake Canyon, ARcher County, Texas.

Horizon: Nocona Formation, Lower Wichita Group.

Biostratigraphy: Land-vertebrate faunachron lower B.

Age: Upper Wolfcampanian stage, Upper Rotliegendes Epoch, Early Permian.

Material:

MCZ 1167: Skull fragments, right cheek and quadrate.

MCZ 1169: Palate and occipit.

MCZ 1508: Slab with skulls.

MCZ 8118: Clavicle.

MCZ 8128: Cheek,

MCZ 8139: Cheek.

MCZ 8151: Premaxilla.

MCZ 8204: Block with two skulls.

 

Locality: East side of Godwin Creek near mouth, J. J. Lang A-833, Archer County, Texas.

Horizon: Nocona Formation, Lower Wichita Group.

Biostratigraphy: Land-vertebrate faunachron lower B.

Age: Upper Wolfcampanian stage, Upper Rotliegendes Epoch, Early Permian.

Material:

MCZ 6663: Left mandible fragment.

 

Locality: 3 miles south of Black Flat, Hooper and Wade Survey A-689, Archer County, Texas.

Horizon: Nocona Formation, Lower Wichita Group.

Biostratigraphy: Land-vertebrate faunachron lower B.

Age: Upper Wolfcampanian stage, Upper Rotliegendes Epoch, Early Permian.

Material:

MCZ 8548: Clavicle.

 

Locality:Near Dundee, Archer County, Texas.

Horizon: Petrolia Formation, Lower Wichita Group.

Biostratigraphy: Land-vertebrate faunachron upper B.

Age: Upper Wolfcampanian stage, Upper Rotliegendes Epoch, Early Permian.

Material:

FMNH UC230: Skull and postcranium.

 

 

Locality: Slippery Creek, Archer County, Texas.

Horizon: Petrolia Formation, Lower Wichita Group.

Biostratigraphy: Land-vertebrate faunachron upper B.

Age: Upper Wolfcampanian stage, Upper Rotliegendes Epoch, Early Permian.

Material:

MCZ 1168: Skull fragments.

 

Locality: Locality A-264, Archer County, Texas.

Horizon: Petrolia Formation, Lower Wichita Group.

Biostratigraphy: Land-vertebrate faunachron upper B.

Age: Upper Wolfcampanian stage, Upper Rotliegendes Epoch, Early Permian.

Material:

UCMP 27166: Scapuloracoid.

 

Locality: Parkey Ranch, Slippery Creek, Dundee, Archer County, Texas.

Horizon: Petrolia Formation, Lower Wichita Group.

Biostratigraphy: Land-vertebrate faunachron upper B.

Age: Upper Wolfcampanian stage, Upper Rotliegendes Epoch, Early Permian.

Material:

USNM 328103: Skull.

 

Locality: Main Pocket, Hay Camp, Archer County, Texas.

Horizon: Petrolia Formation, Lower Wichita Group.

Biostratigraphy: Land-vertebrate faunachron upper B.

Age: Upper Wolfcampanian stage, Upper Rotliegendes Epoch, Early Permian.

Material:

USNM 406387: Skull roof.

 

 

Locality: West of Charles Williams Ranch, Gibbs Survey A-566, Baylor County, Texas.

Horizon: Petrolia Formation, Lower Wichita Group.

Biostratigraphy: Land-vertebrate faunachron upper B.

Age: Upper Wolfcampanian stage, Upper Rotliegendes Epoch, Early Permian.

Material:

MCZ 1546: 3 snouts.

 

Locality: Wolf Creek (Red Holow),2 miles NW Boone House, H. T. and B. Survey, Sect. 11, A-160, Baylor County, Texas.

Horizon: Petrolia Formation, Lower Wichita Group.

Biostratigraphy: Land-vertebrate faunachron upper B.

Age: Upper Wolfcampanian stage, Upper Rotliegendes Epoch, Early Permian.

Material:

MCZ 2378: SKull.

 

Locality: West of Charles Williams Ranch, J. Gibbs Survey A-566, N side of North Folk, Little Wichitia River, below Fulda, Baylor County, Texas.

Horizon: Petrolia Formation, Lower Wichita Group.

Biostratigraphy: Land-vertebrate faunachron upper B.

Age: Upper Wolfcampanian stage, Upper Rotliegendes Epoch, Early Permian.

Material:

MCZ 2387: Skull.

 

Locality: Tit Mountain, Palo Pinot county, School Land, NE of Dundee, Archer County, Texas.

Horizon: Petrolia Formation, Lower Wichita Group.

Biostratigraphy: Land-vertebrate faunachron upper B.

Age: Upper Wolfcampanian stage, Upper Rotliegendes Epoch, Early Permian.

Material:

MCZ 8208: Fragments including a mandible.

MCZ 8215: Clavicle.

MCZ 8261: Skull table.

 

 

Locality: Near Redham, Baylor County, Texas.

Horizon: Petrolia Formation, Lower Wichita Group.

Biostratigraphy: Land-vertebrate faunachron upper B.

Age: Upper Wolfcampanian stage, Upper Rotliegendes Epoch, Early Permian.

Material:

MCZ 8298: Right mandible.

 

Locality: North side of North Folk, Little Wichita Below Fulda, J. Gibbs Survey A-566, Baylor County, Texas.

Horizon: Petrolia Formation, Lower Wichita Group.

Biostratigraphy: Land-vertebrate faunachron upper B.

Age: Upper Wolfcampanian stage, Upper Rotliegendes Epoch, Early Permian.

Material:

MCZ 8493: Mandible.

 

Locality: No locality data, Texas.

Horizon: Petrolia Formation, Lower Wichita Group.

Biostratigraphy: Land-vertebrate faunachron upper B.

Age: Upper Wolfcampanian stage, Upper Rotliegendes Epoch, Early Permian.

Material:

MCZ 8538: Scapulocorid.

 

Locality: South side of North Fork, Little Wichita River, below Fulda, T and NO Railway survey, Baylor, County, Texas.

Horizon: Petrolia Formation, Lower Wichita Group.

Biostratigraphy: Land-vertebrate faunachron upper B.

Age: Upper Wolfcampanian stage, Upper Rotliegendes Epoch, Early Permian.

Material:

MCZ 8598: Check fragment.

 

Locality: Locality V 5217, Thrift County, Texas.

Horizon: Petrolia Formation, Lower Wichita Group.

Biostratigraphy: Land-vertebrate faunachron upper B.

Age: Upper Wolfcampanian stage, Upper Rotliegendes Epoch, Early Permian.

Material:

UCMP 105126: Palate and pectoral girdle.

UCMP 105146: Posterior table with hyoid elements.

UCMP 105165: Partial palate.

UCMP 105175, 142027: Complete skulls.

UCMP 105224: Part of right palate.

UCMP 105239: Anterior part of skull.

UCMP 105260: Part of left mandible.

UCMP 154146: Posterior palate.

UCMP 154434: Specimen showing ceratobranchials.

 

 

OLSON, 1979

Locality: Patterson ranch in the SW 1/4, sec 436 of the King Survey, Taylor County, Texas.

Horizon: Vale Formation, Middle Clear Fork Group.

Biostratigraphy: Land-vertebrate faunachron lower to middle D.

Age: Upper Leonardian Stage,Upper Rotliegendes Epoch, Early Permian.

Material:

UCLA VP 463: Skull and nearly complete skeleton.

UCLA VP 466, 468: Fragmentary skulls and fragmentary skeleton.

UCLA VP 469: Fragments.

Plus 3 other specimens.

 

SANDER, 1989

Locality: Loftin Bonebed, Archer County, Texas.

Horizon: Nocona Formation (formerly lower to lower upper Admiral Formation, middle Wichita Group), Lower Wichita Group.

Biostratigraphy: Land-vertebrate faunachron lower B.

Age: Upper Wolfcampanian stage, Upper Rotliegendes Epoch, Early Permian.

Material:

Number: Not given: Intercentra.


Locality: Rattlesnake Canyon Bonebed 2, Archer County, Texas.

Horizon: Nocona Formation (formerly lower to lower upper Admiral Formation, middle Wichita Group), Lower Wichita Group.

Biostratigraphy: Land-vertebrate faunachron lower B.

Age: Upper Wolfcampanian stage, Upper Rotliegendes Epoch, Early Permian.

Material:

Number: Not given: Teeth and dermal bone.

 

WITZMAN, 2007

Locality: Baylor County, Texas.

Horizon: Waggoner Ranch Formation (formerly Clyde Formation, upper Wichita Group) Upper Wichita Group.

Biostratigraphy: Land-vertebrate faunachron lower to middle C.

Age: Lower Leonardian Stage, Lower Rotliegendes Epoch, Early Permian.

Material:

MCZ 1080: Pectoral girdle, vertebrae dorsal and gastral scales.

 

ZIDEK, JOHNSON, MAY & CLABORN, 2004

Locality: OMNH V 173, Northeast Frederic site of Simpson, (1976, 1979), Tillman County, Oklahoma.

Horizon: Garber Sandstone, Upper part of the Sumner Group, possibly equivalent to the Lower Arroyo Formation, Lower Clear Fork Group.

Biostratigraphy: Land-vertebrate faunachron lower D.

Age: Middle Leonardian stage, upper Rotliegendes, Early Permian.

Material:

Number: Not given:


Locality: OMNH V 247, Lake Frederick site, Tillman County, Oklahoma.

Horizon: Garber Sandstone, Upper part of the Sumner Group, possibly equivalent to the Lower Arroyo Formation, Lower Clear Fork Group.

Biostratigraphy: Land-vertebrate faunachron lower D.

Age: Middle Leonardian stage, upper Rotliegendes, Early Permian.

Material:

Number: Not given:

 

= Trimerorhachis cf. insignis DALY, 1973

Locality: SW1/4, SE1/4, SE1/4, sec. 20, T4S, R15W, 6 1/2 miles southwest of Grandfield, Tilman County, Oklahoma.

Horizon: Hennessey Formation, (Arroyo equivalent), Low Clear Fork Group.

Biostratigraphy:

Age: Leonardian Stage, Lower Rotliegendes Epoch, Early Permian.

Material:

FM UR2383: Fragmentary pectoral girdle.

FM UR2271: Fragmentary left muzzle.

FM UR2272: Fragmentary right half-muzzle.

FM UR2384: 3 small sculptured pieces.

 

OLSON, 1970

Locality: Bed 18 and 19, about 9 miles northeast of Perry, Logan County, Oklahoma, SE 1/4, SE 1/4, sec. 26, T23N, R2W.

Horizon: Wellington Formation (Garber Formation), Summer Group, Arbuckle limestone, (Probably equivalent to the Arroyo Formation of Texas).

Biostratigraphy:

Age: Leonardian Stage, Lower Rotliegendes Epoch, Early Permian.

Material:

Number: Not given.

 

Locality: Bed 16, about 9 miles northeast of Perry, Logan County, Oklahoma, SE 1/4, SE 1/4, sec. 26, T23N, R2W.

Horizon: Wellington Formation (Garber Formation), Summer Group, Arbuckle limestone, (Probably equivalent to the Arroyo Formation of Texas).

Biostratigraphy:

Age: Leonardian Stage, Lower Rotliegendes Epoch, Early Permian.

Material:

Number: Not given.


= Trimerorhachis bilobatus COPE, 1883
Etymology:

Holotype: AMNH 4562

Locality: Corn Hill (originally called Tit Mountain), Archer County, Texas.

Horizon: Petrolia Formation, Lower Wichita Group.

Biostratigraphy: Land-vertebrate faunachron upper B.

Age: Upper Wolfcampanian stage, Upper Rotliegendes Epoch, Early Permian.

Material: Partial mandible.

 

= Trimerorhachis alleni CASE, 1910
Etymology:

Holotype: AMNH 4577

Locality: Rattlesnake Canyon, Archer County, Texas.

Horizon: Nocona Formation, Lower Wichita Group.

Biostratigraphy: Land-vertebrate faunachron lower B.

Age: Upper Wolfcampanian stage, Upper Rotliegendes Epoch, Early Permian.

Material: 4 dorsal vertebrae.

 

= Trimerorhachis medius BROOM, 1913
Etymology:

Holotype: AMNH 4895

Locality: North side, Little Wichita River, Archer County, Texas

Horizon: Petrolia Formation, Lower Wichita Group.

Biostratigraphy: Land-vertebrate faunachron upper B.

Age: Upper Wolfcampanian stage, Upper Rotliegendes Epoch, Early Permian.

Material: Fragmentary skull.

_____________________________________________________________________________________
Species: mesops COPE, 1896
Etymology:

Holotype: AMNH 4568

Locality: West Coffee Creek, Baylor County, Texas.

Horizon: Arroyo Formation, Lower Clear Fork Group.

Biostratigraphy: Land-vertebrate faunachron lower D.

Age: Middle Leonardian stage, upper Rotliegendes, Early Permian.

Material: Fragmentary skull and vertebrae.

Referred material:

MCZ 1165: Skull.

MCZ 8726: Left cheek.

 

Localtiy: Baylor County, Texas.

Horizon: Arroyo Formation, Lower Clear Fork Group.

Biostratigraphy: Land-vertebrate faunachron lower D.

Age: Middle Leonardian stage, upper Rotliegendes, Early Permian.

Material:

AMNH 4591: Skull.

 

Locality: Indian Creek, Baylor County, Texas.

Horizon: Arroyo Formation, Lower Clear Fork Group.

Biostratigraphy: Land-vertebrate faunachron lower D.

Age: Middle Leonardian stage, upper Rotliegendes, Early Permian.

Material:

FMNH C688: Skull.

 

Locality:Indian Creek? Baylor County, Texas.

Horizon: Arroyo Formation, Lower Clear Fork Group.

Biostratigraphy: Land-vertebrate faunachron lower D.

Age: Middle Leonardian stage, upper Rotliegendes, Early Permian.

Material:

FMNH UC 689: Skull.

 

Locality: Coffee Creek, Baylor County, Texas.

Horizon: Arroyo Formation, Lower Clear Fork Group.

Biostratigraphy: Land-vertebrate faunachron lower D.

Age: Middle Leonardian stage, upper Rotliegendes, Early Permian.

Material:

GPIT 1800: Skull.

 

Locality: Branch West, Coffee Creek, Baylor County, Texas.

Horizon: Arroyo Formation, Lower Clear Fork Group.

Biostratigraphy: Land-vertebrate faunachron lower D.

Age: Middle Leonardian stage, upper Rotliegendes, Early Permian.

Material:

USNM 17049: Skull and partial skeleton.

 

Localtyi: H and TC Railroad Survey, Block 4, Section 7, near Grayback, Willbarger County, Texas.

Horizon: Middle Waggoner Ranch Formation.

Biostratigraphy:

Age: Early Permian.

Material:

MCZ 3221: Skull.

_____________________________________________________________________________________

Species: rogersi OLSON, 1955
Etymology:

Holotype: FMNH UR138

Locality: Pipe Site, Locality FA, Foard County, Texas.

Horizon: Choza Formation, Middle Clear Fork Group.

Biostratigraphy:

Age: Early Permian.

Material: Partial skull and postcranial elements.

Referred material:

FMNH UR73: Long slender mandible.

FMNH UR74: Skull and jaws.

_____________________________________________________________________________________
Species: sandovalensis BERMAN & REISZ, 1980
Etymology:

Holotype: CM 38025

Locality: On west side of Canyon de San Diego, west of State Highway 4 and along western edge of Jemez River, about 11.5 km south of Jemez= Springs, approximately lat 35'40"N and Long 106'45"W, Sandoval County, New Mexico.

Horizon: Cutler Formation, (originally listed as Lower part of the Abo Formation.)

Biostratigraphy:

Age: Wolfcampian Stage, Lower Rotliegendes Epoch, Early Permian.

Material: Fragmentary skull, dentaires and fragmentary skeleton.

Referred material:

= Trimerorhachis cf. T. sandovalensis CANTRELL, SUAZO, BERMAN, SPIELMANN, LUCAS, HENRICI & RINEHART, 2013

Locality: Gallina Wash locality (NMMNH locality 4668, CM locality 1057), Joyita Hills region, approximately 20 km northeast of Socorro, central Socorro County, New Mexico.

Horizon: Scholle Member, Lower Abo Formation.

Biostratigraphy:

Age: Middle Wolfcampian State (Coyotean or Seymouran age), Lower Rotliegendes Epoch, Early Permian.

Material:

CM 38008, CM38010: Partial skulls.

_____________________________________________________________________________________

Species: greggi MILNER & SCHOCH, 2013
Etymology: In honor of Mr Donald Gregg of Seymour, Texas, who not only collected the type specimen, but faciliated several productive excavations by the USNM in the Lake Kemp area.

Holotype: USNM 409692

Locality: Craddock Bone-bed, Southwest Lake Kemp, Baylor County, Texas.

Horizon: 5 meters below Red Tank Sandstone Member (informal), Arroyo Formation, Lower Clear Fork Group.

Biostratigraphy: Land-vertebrate faunachron lower D.

Age: Middle Leonardian stage, upper Rotliegendes, Early Permian.

Material: Skull.

Referred material:

Locality: 2 miles NW of Cockrell Ranch (now Powell Ranch), Brushy Creek, Baylor County, Texas.

Horizon: Arroyo Formation, Lower Clear Fork Group.

Biostratigraphy: Land-vertebrate faunachron lower D.

Age: Middle Leonardian stage, upper Rotliegendes, Early Permian.

Material: Skull.

 

Locality: Pipe Site, Locality FA, Foard County, Texas.

Horizon: Choza Formation, Middle Clear Fork Group.

Biostratigraphy: Land-vertebrate faunachron lower D.

Age: Middle Leonardian stage, upper Rotliegendes, Early Permian.

Material: Skull with broad mandible..