Genus: Clidastes COPE, 1868
Etymology:
= Edestosaurus MARSH, 1871
Etymology:

Species: propython COPE, 1869
Etymology:

Holotype: ANSP 10193

Locality: Uniontown, Perry County, Alabama.

Horizon: Selma Formation.

Biostratigraphy:

Age: Campanian Stage, Senonian subepoch, Upper Gulf epoch, Late Cretaceous.

Material: Skull and nearly complete skeleton.

Frontal from JIMENEZ-HUIDOBRO, SIMOES & CALDWELL, 2016.

 

Clidastes skull after Carroll & Debraga, 1992.

Referred material:

BEHLKE & BOYD, 2003

Locality: Logan County, Kansas.

Horizon: Smoky Hill Chalk Member, Niobrara Formation.

Biostratigraphy:

Age: Santonian or Campanian Stage, Middle Senonian Subepoch, Uppermiddle Gulf Epoch, Late Cretaceous.

Material:

Number: Not given: Articulated cervical and dorsal series, several caudals, ribs and a nearly complete left front paddle.

 

= Clidastes cf. propython GERMANN, 2018

Locality: Alsen near Lägerdorf, Schleswig-Holstein State, Germany.

Horizon:

Biostratigraphy: Papillosa-Zone, Conica/gracilis zone.

Age: Early Campanian Stage, Senonian Subepoch, Gulf Epoch, Late Cretaceous.

Material:

Number: Not given: Tooth with fragment of jaw, a fragmentary tooth in a jaw fragment and a fragment of a sacral vertebra.

GRIGORIEV, ARKHANGELSKY, & MERKULOV, 2015

Locality: Beloe Ozero locality, Lysogorskii District, Saratov Region, European Russia.

Horizon: Rybushka Formation.

Biostratigraphy:

Age:

Campanian Stage, Middle Senonian Subepoch, Uppermiddle Gulf Epoch, Late Cretaceous.

Material:

ZIN PH, no. 5/48: Fused frontals.

 

JIMENEZ-HUIDOBRO, SIMOES & CALDWELL, 2016.

Locality:

Horizon:

Bisotratigraphy:

Age:

Material:

FHSM VP 985.0012: Frontal.

FHSM VP 985.0012 frontal from JIMENEZ-HUIDOBRO, SIMOES & CALDWELL, 2016.

 

KUEHNE & KUEHNE, 2018

Locality: Stone Bidge Site, Camden County, New Jersey.

Horizon: Englishtown Formation.

Biostratigraphy:

Age: Late Campanian Stage, Senonian Subepoch, Late Gulf Epoch, Late Cretaceous.

Material:

NJSM GP 21913: Caudal vertebral centrum.

NJSM GP 21609: Tooth.

 

 

LINDGREN & SIVERSON, 2004

Locality: Asen, Kristianstad Basin, N 56°08’56.1”, E 14°29’56.0”, Skane (Scania) Province, Sweden.

Horizon:

Biostratigraphy: Belemnellocamax mammillatus zone.

Age: Early Campanian Stage, Senonian subepoch, Upper Gulf epoch, Late Cretaceous.

Material:

RM PZ R1778: The 3rd? cervical vertebra from a juvenile.

RM PZ R1779: 6th or 7th cervical vertebra.

RM PZ R1797: Second premaxillary or second dentary tooth crown.

RM PZ R1798: A large tooth-crown from the mid portion of jaw.

RM PZ R1799: A large tooth-crown from the anterior part of a mid-portion of the jaw.

RM PZ R1800: Crown from the anterior part of the mid-portion of the jaw of a juvenile.

RM PZ R1801: Crown from the posterior portion of the jaw.

RM PZ R1803: Crown from the posterior portion of the jaw.

RM PZ R1804, R1805: Pterygoid teeth crowns.

LO 82312: Tooth-crown of a juvenile.

LO 8232t: Tooth-crown from mid-portion of the jaw.

Number: Not given: 12 pterygoid teeth, 93 marginal teeth, 2 cervical vertebrae, 2 dorsal vertebrae, 6 cervical/dorsal vertebrae and 10 caudal vertebrae.

 

Locality: Axeltorp Kristianstad Basin, Skane (Scania) Province, Sweden.

Horizon:

Biostratigraphy: Belemnellocamax mammillatus zone.

Age: Early Campanian Stage, Senonian subepoch, Upper Gulf epoch, Late Cretaceous.

Material:

RM PZ R1752a, b: 2 fragments of an axis.

 

 

Locality: Ignaberga, Kristianstad Basin, Skane (Scania) Province, Sweden.

Horizon:

Biostratigraphy: Belemnellocamax mammillatus zone.

Age: Early Campanian Stage, Senonian subepoch, Upper Gulf epoch, Late Cretaceous.

Material:

Number: Not given: 3 marginal teeth.

 

Locality: Ivo Klack, Kristianstad Basin, Skane (Scania) Province, Sweden.

Horizon:

Biostratigraphy: Belemnellocamax mammillatus zone.

Age: Early Campanian Stage, Senonian subepoch, Upper Gulf epoch, Late Cretaceous.

Material:

RM PZ R1027: Terminal caudal vertebra.

Number: Not given: 1 marginal tooth.

 

 

Locality: Ugnsmunnarna, Kristianstad Basin, Skane (Scania) Province, Sweden.

Horizon:

Biostratigraphy: Belemnellocamax mammillatus zone.

Age: Early Campanian Stage, Senonian subepoch, Upper Gulf epoch, Late Cretaceous.

Material:

Number: Not given: 3 marginal teeth.


 

MARTIN, SCHUMACHER PARRIS & GRANDSTAFF, 1998

Locality: Marindahl Dam Spillway, Yankton county, South Dakota.

Horizon: 4.6 meters below the Niobrara/Pierre Contact.

Biostratigraphy:

Age: Late Cretaceous.

Material:

WHO unnumbered: A block of chalk containing frontal, prefrontal, pterygoid, 2 dentaries, jugal, 4 cervical vertebrae, trunk vertebrae and 10 ribs.

 

OM, MANNING, BADGER & WHITLEY, 2016

Locality: Stone Bridge Locality, Cherry Hill, Camden County, southern New Jersey.

Horizon: Upper Woodbury Formation.

Biostratigraphy:

Age: Early-Middle Campanian Stage, Senonian Subepoch, Gulf Epoch, Late Cretaceous.

Material:

Number: Not given: Teeth and caudal vertebra.

RUSSELL, 1970

Locality: Harrell Station Area, Southeast of Marion junction, Dallas County, Alabama.

Horizon: Selma Formation.

Biostratigraphy:

Age: Late Cretaceous.

Material:

P 27324: Disarticulated skull, presacral vertebra.

P 27828: Pterygoid and 6 vertebrae.

P 27439: Parts of skull, atlas and cervical vertebrae.

P 27440: Scapula and coarcoid fragments.

P 27449, 27545: Humerii.

P 27478: Tooth and metacarpal.

P 27484: Fragmentary disarticulated skull and presacral vertebrae.

P 27546: Quadrate.

PR 37: Cranial elements and humerus.

PR 38: Disarticulated skull and vertebrae.

PR 39: Cranial elements and presacral vertebrae.

PR 40: Disarticulated skull, most of vertebral column and appendicular elements.

PR 41: Cranial elements, vertebrae and hind limbs.

PR 42, 43, 46: Vertebrae.

PR 45: Cranial elements, presacral vertebrae and pectoral elements.

PR 47: Presacral vertebrae.

PR 48: Cranial elements, presacral vertebrae and humerus.

PR 49: Pygal vertebrae.

PR 50: Dorsal vertebrae.

PR 52: Disarticulated skull and atlas.

PR 53: Caudal vertebrae.

PR 138: Quadrate, marginal tooth and caudal vertebra.

PR 142: Cranial elements and vertebrae.

PR 192: Pectoral elements.

PR 200: Pygal vertebra and phalanges.

PR 211: Postorbitofrontal.

PR 238: Dentary.

 

Locality: Cedarville, Hale County, Alabama.

Horizon: Selma Formation.

Biostratigraphy:

Age: Late Cretaceous.

Material:

PR 51: Vertebrae.

 

Locality: Boligee, Greene County, Alabama.

Horizon: Selma Formation.

Biostratigraphy:

Age: Late Cretaceous.

Material:

PR 63: Humerus.

 


Locality: Mount Hebron, Greene County, Alabama.

Horizon: Selma Formation.

Biostratigraphy:

Age: Late Cretaceous.

Material:

PR 150: Cranial elements and cervical vertebrae.

 

Locality: West Greene, Greene County, Alabama.

Horizon: Selma Formation.

Biostratigraphy:

Age: Late Cretaceous.

Material:

PR 148: Cranial elements and vertebrae.

PR 156: Cranial elements, vertebrae and humerus.

PR 162: Quadrate and vertebrae.

PR 164: Disarticulated skull, most of vertebral column, and appendicular elements.

PR 175: Dorsal vertebrae.

PR 176: Caudal vertebrae.

PR 183: Radius.

 


Locality: Unknown.

Horizon: Selma Formation?

Biostratigraphy:

Age: Late Cretaceous.

Material:

P 27355 : Dentary and splenial.

 

RUSSELL, 1967

Locality:

Horizon:

Biostratigraphy:

Age:

Material:

AMNH 182:

AMNH R173:

AMNH 1507:

AMNH 1525:

AMNH 1546:

AMNH 1593:

AMNH ND2: Fragmentary skull.

YPM 1308:

YPM 1315:

YPM 1328:

and 13 unnumbered specimens.

USNM 3765:

USNM 3778:

MCZ 1625:

KU 1000:

KU 1005:

KU 1058:

KU 1108:

KU 1124:

KU 1134:

CNHM UR901:

CNHM P 12856:

CM 1190:

BELL, 1997

ANSP 10193:

UCMP 34535:

YPM 1105:

YPM 1310:

SHELDON, 1997

RMM 5830, 1788:

 


Locality: From bluff near A. Finney's ranch, Cottonwood Creek, SSW of Edgemont, Fall River County, South Dakota.

Horizon: Lower Pierre Formation.

Biostratigraphy:

Age: Late Cretaceous.

Material:

AMNH 5812:

 

NICHOLLS, 1988

Locality: South Dakota.

Horizon: Sharon Springs Member, Pierre Shale.

Biostratigraphy:

Age: Campanian Stage, Senonian subepoch, Upper Gulf epoch, Late Cretaceous.

Material:

As per NICHOLLS, 1988

AMNH 5812, FMNH PR 560, 671, KUVP uncatalogued, SDSM PC 8026 (field number), RTMP P.83.19.2, UCM 20568, UCM uncatalogued, USNM 2778-66, 2859-66:


SHELDON, 1996

Locality: Logan County, Kansas.

Horizon: Smoky Hill Chalk Member, Niobrara Formation.

Biostratigraphy:

Age: Santonian or Campanian Stage, Middle Senonian Subepoch, Uppermiddle Gulf Epoch, Late Cretaceous.

Material:

YPM 1318, 1319, 1324, 1310: Adult.

YPM 1316, 1321: Subadult.

YPM 1371,

YPM 1368: Skull of a late juvenile.

YPM 1314: Early juvenile.

LINDGREN & SIVERSON, 2004

YPM 40354: 6th or 7th cervical vertebra.

 


Locality: Wallace County, Kansas.

Horizon: Niobrara County.

Biostratigraphy:

Age: Late Cretaceous.

Material:

YPM 1316: Adult.

YPM 1317: Subadult.

 


Locality: Gove County, Kansas.

Horizon: Niobrara Formation.

Biostratigraphy:

Age: Late Cretaceous.

Material:

YPM 40368: Early juvenile.

 

YAMASHITA, KONISHI, & SATO, 2015

Locality: Southern Manitoba Province, Canada.

Horizon: Pembina Member, Pierre Shale.

Biostratigraphy: Baculites obtusus ammonite zone.

Age: Earliest Middle Campanian Stage, Senonian subepoch, Upper Gulf epoch, Late Cretaceous

Material:

CFDC M74.10.06:

 

= ?Clidastes cineriarum COPE, 1871
Etymology:

Holotype: AMNH ?1579

Locality: 6 miles south of Sheridan, Kansas.

Horizon:

Biostratigraphy:

Age: Late Cretaceous.

Material: Pterygoid teeth and vertebrae.

 

= Edestosaurus dispar MARSH, 1871
Etymology:

Holotype: YPM 1100

Locality:

Horizon:

Biostratigraphy:

Age:

Material:

 

= Edesteosaurus velox MARSH, 1871
Etymology:

Holotype: YPM 1105

Locality: Kansas.

Horizon:

Biostratigraphy:

Age: Late Cretaceous.

Material:

Referred material:

WILLISTON, 1898

Small mandible.

Mandibles.

Fragmentary skulls and fragmentary skeletons.

WILLISTON & CASE, 1892

Locality: Butte Creek, Logan County, Kansas.

Horizon: Smoky Hill Chalk Member, Niobrara Formation.

Biostratigraphy:

Age: Santonian or Campanian Stage, Middle Senonian Subepoch, Uppermiddle Gulf Epoch, Late Cretaceous.

Material:

KUVP 1022: Nearly complete skeleton.

 

Locality: Unknown, Kansas.

Horizon: Smoky Hill Chalk Member, Niobrara Formation.

Biostratigraphy:

Age: Santonian or Campanian Stage, Middle Senonian Subepoch, Uppermiddle Gulf Epoch, Late Cretaceous.

Material:

KUVP 1108: Right quadrate.

 

= Clidastes wymani MARSH, 1871
Etymology:

Holotype: YPM 1120

Locality: North Fork of Smoky Hill, Kansas.

Horizon:

Biostratigraphy:

Age: Late Cretaceous.

Material: Chevron caudal vertebrae.

Referred material:

Number: Not given: Fragmentary skull and anterior vertebrae.

 

= Clidastes pumilus MARSH, 1871
Etymology:

Holotype: YPM 1123

Locality: Logan County, Kansas.

Horizon: Smoky Hill Chalk Member, Niobrara Formation.

Biostratigraphy:

Age: Santonian or Campanian Stage, Middle Senonian Subepoch, Uppermiddle Gulf Epoch, Late Cretaceous.

Material: Quadrate of an early juvenile.

Referred material:

ABEL, 1919

Locality: Western Kansas.

Horizon:

Biostratigraphy:

Age: Late Cretaceous.

Material:

KUVP 69552: Front paddle.

 

= Edestosaurus tortor COPE, 1871
Etymology:
= Clidastes tortor (COPE, 18721) COPE, 1874

Holotype: AMNH 1513

Locality: Fossil Spring canon, Kansas.

Horizon:

Biostratigraphy:

Age: Late Cretaceous.

Material: Fragmentary skull and fragmentary skeleton.

Referred material:

WILLISTON, 1898

Locality: Smoky Hill River, Kansas.

Horizon: Smoky Hill Chalk Member, Niobrara Formation.

Biostratigraphy:

Age: Santonian or Campanian Stage, Middle Senonian Subepoch, Uppermiddle Gulf Epoch, Late Cretaceous.

Material:

KUVP 1000: Skull, lower jaws, quadrate, vertebral column, left scapula and coracoid, pelvic girdle, parts of front paddle and ribs.

 

Locality: Unknown, Kansas.

Horizon: Smoky Hill Chalk Member, Niobrara Formation.

Biostratigraphy:

Age: Santonian or Campanian Stage, Middle Senonian Subepoch, Uppermiddle Gulf Epoch, Late Cretaceous.

Material:

KUVP 14278: Left quadrate.

KUVP?: Posterior cervical vertebrae.

KUVP?: Frontal bone.

KUVP?: Left quadrate.

 

= Edestosaurus stenops COPE, 1871
Etymology:

Holotype: AMNH 1575

Locality: Near Fossil spring, Kansas.

Horizon:

Biostratigraphy:

Age: Late Cretaceous.

Material: Fragmentary skull.

Referred material:

Locality: On a bluff on Butte Creek, Kansas.

Horizon:

Biostratigraphy:

Age: Late Cretaceous.

Material:

Number: Not given: Fragmentary skeletons.

 

= Edestosaurus rex MARSH, 1872
Etymology:

Holotype: YPM 1310

Locality: Smoky Hill River, Kansas.

Horizon: Niobrara Formation.

Biostratigraphy:

Age: Late Cretaceous.

Material: Fragmentary skeleton.

 

= Clidastes medius MERRIAM, 1894
Etymology:

Holotype:

Locality:

Horizon:

Biostratigraphy:

Age:

Material:

 

= Clidastes westi WILLISTON & CASE, 1892
Etymology:

Holotype: KU 1026

Locality: McAllister, Wallace County, Kansas.

Horizon: Lower Pierre Formation.

Biostratigraphy:

Age: Campanian Stage, Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.

Material: Nearly complete skull, lower jaw, 84 vertebrae, ribs, front and hind paddles.

_____________________________________________________________________________________

Species: liodontus MERRIAM, 1894
Etymology:

Holotype:

Locality: Logan County, Kansas.

Horizon: Smoky Hill Chalk Member, Niobrara Formation.

Biostratigraphy:

Age: Santonian or Campanian Stage, Middle Senonian Subepoch, Uppermiddle Gulf Epoch, Late Cretaceous.

Material: Maxillae, premaxilla and dentaries.

Referred material:

AMNH 192:

AMNH 1548:

YPM 1333:

YPM 1334:

YPM 1335: Skull.

YPM 3982:

YPM 3996: Fragmentary skull.

YPM 24914:

3 unnumbered specimens at YPM:

USNM 6546:

USNM 11647:

USNM 11719:

MCZ 278:

MCZ 1618:

MCZ 1621:

KU 1039:

KU 1121:

FHM 10668:

PU 17249:

BELL, 1997

KU 1022:

FHM VP2071:

 


Locality: South Dakota.

Horizon: Sharon Springs Member, Pierre Shale.

Biostratigraphy:

Age: Campanian Stage, Senonian subepoch, Upper Gulf epoch, Late Cretaceous.

Material:

KUVP 50104:

As per NICHOLLS, 1988

 

BELL, BARNES & POLCYN, 2007

Localty: Western part of the Big Bend Region, Brewster County, Texas.

Horizon: Lower Pen Formation.

Biostratigraphy:

Age: Upper Coniacian Stage, Lower most Senonian Subepoch-Santonian Stage, Lower Senonian Subepoch, Lower Gulf Epoch, Early Late Cretaceous.

Material:

Number: Not given:

 

SHELDON, 1996

Locality: Logan County, Kansas.

Horizon: Niobrara Formation.

Biostratigraphy:

Age: Late Cretaceous.

Material:

YPM 1367: Late juvenile.


_____________________________________________________________________________________

Species: moorevillensis SHANNON, 1975 (nomen ex dissertatione, nomen nudum)
Etymology: In reference to Mooreville Chalk, Selma Group.

Holotype:

Locality:

Horizon: Mooreville Chalk, Selma Group.

Biostratigraphy: Middle Tylosaurus Acme-zone – Clidastes Acme zone.

Age: Niobraran age, Upper Santonian-Lower Campanian Stage, Senoniain Subepoch, Gulf Epoch, Late Cretaceous.

Material:

Referred material:

BEHLKE, OTT, KELLY & SPEARING, 2002

Loclaity: Logan County, Kansas.

Horizon: Niobrara Formation.

Biostratigraphy:

Age: Early Campanian Stage, Gulf Epoch, Late Cretaceous.

Material:

Number: Not given: Nealry complete specimen.
Note: 7 meters long.

 

BELL, 1997

Locality:

Horizon:

Biostratigraphy:

Age:

Material:

RMM 070:

GSA TC218:

RMNH PR495: