Species: propython COPE, 1869
Etymology:

Holotype: ANSP 10193

Locality: Uniontown, Perry County, Alabama.

Horizon: Selma Formation.

Biostratigraphy:

Age: Campanian Stage, Senonian subepoch, Upper Gulf epoch, Late Cretaceous.

Material: Skull and nearly complete skeleton.

Frontal from JIMENEZ-HUIDOBRO, SIMOES & CALDWELL, 2016.

Clidastes skull after Carroll & Debraga, 1992.

Referred material:

BEHLKE & BOYD, 2003

Locality: Logan County, Kansas.

Horizon: Smoky Hill Chalk Member, Niobrara Formation.

Biostratigraphy:

Age: Santonian or Campanian Stage, Middle Senonian Subepoch, Uppermiddle Gulf Epoch, Late Cretaceous.

Material:

Number: Not given: Articulated cervical and dorsal series, several caudals, ribs and a nearly complete left front paddle.

 

= Clidastes cf. propython GERMANN, 2018

Locality: Alsen near Lägerdorf, Schleswig-Holstein State, Germany.

Horizon:

Biostratigraphy: Papillosa-Zone, Conica/gracilis zone.

Age: Early Campanian Stage, Senonian Subepoch, Gulf Epoch, Late Cretaceous.

Material:

Number: Not given: Tooth with fragment of jaw, a fragmentary tooth in a jaw fragment and a fragment of a sacral vertebra.

GRIGORIEV, ARKHANGELSKY, & MERKULOV, 2015

Locality: Beloe Ozero locality, Lysogorskii District, Saratov Region, European Russia.

Horizon: Rybushka Formation.

Biostratigraphy:

Age:

Campanian Stage, Middle Senonian Subepoch, Uppermiddle Gulf Epoch, Late Cretaceous.

Material:

ZIN PH, no. 5/48: Fused frontals.

 

JIMENEZ-HUIDOBRO, SIMOES & CALDWELL, 2016.

Locality:

Horizon:

Bisotratigraphy:

Age:

Material:

FHSM VP 985.0012: Frontal.

FHSM VP 985.0012 frontal from JIMENEZ-HUIDOBRO, SIMOES & CALDWELL, 2016.

 

KUEHNE & KUEHNE, 2018

Locality: Stone Bidge Site, Camden County, New Jersey.

Horizon: Englishtown Formation.

Biostratigraphy:

Age: Late Campanian Stage, Senonian Subepoch, Late Gulf Epoch, Late Cretaceous.

Material:

NJSM GP 21913: Caudal vertebral centrum.

NJSM GP 21609: Tooth.

 

 

LINDGREN & SIVERSON, 2004

Locality: Asen, Kristianstad Basin, N 56°08’56.1”, E 14°29’56.0”, Skane (Scania) Province, Sweden.

Horizon:

Biostratigraphy: Belemnellocamax mammillatus zone.

Age: Early Campanian Stage, Senonian subepoch, Upper Gulf epoch, Late Cretaceous.

Material:

RM PZ R1778: The 3rd? cervical vertebra from a juvenile.

RM PZ R1779: 6th or 7th cervical vertebra.

RM PZ R1797: Second premaxillary or second dentary tooth crown.

RM PZ R1798: A large tooth-crown from the mid portion of jaw.

RM PZ R1799: A large tooth-crown from the anterior part of a mid-portion of the jaw.

RM PZ R1800: Crown from the anterior part of the mid-portion of the jaw of a juvenile.

RM PZ R1801: Crown from the posterior portion of the jaw.

RM PZ R1803: Crown from the posterior portion of the jaw.

RM PZ R1804, R1805: Pterygoid teeth crowns.

LO 82312: Tooth-crown of a juvenile.

LO 8232t: Tooth-crown from mid-portion of the jaw.

Number: Not given: 12 pterygoid teeth, 93 marginal teeth, 2 cervical vertebrae, 2 dorsal vertebrae, 6 cervical/dorsal vertebrae and 10 caudal vertebrae.

 

Locality: Axeltorp Kristianstad Basin, Skane (Scania) Province, Sweden.

Horizon:

Biostratigraphy: Belemnellocamax mammillatus zone.

Age: Early Campanian Stage, Senonian subepoch, Upper Gulf epoch, Late Cretaceous.

Material:

RM PZ R1752a, b: 2 fragments of an axis.

 

 

Locality: Ignaberga, Kristianstad Basin, Skane (Scania) Province, Sweden.

Horizon:

Biostratigraphy: Belemnellocamax mammillatus zone.

Age: Early Campanian Stage, Senonian subepoch, Upper Gulf epoch, Late Cretaceous.

Material:

Number: Not given: 3 marginal teeth.

 

Locality: Ivo Klack, Kristianstad Basin, Skane (Scania) Province, Sweden.

Horizon:

Biostratigraphy: Belemnellocamax mammillatus zone.

Age: Early Campanian Stage, Senonian subepoch, Upper Gulf epoch, Late Cretaceous.

Material:

RM PZ R1027: Terminal caudal vertebra.

Number: Not given: 1 marginal tooth.

 

 

Locality: Ugnsmunnarna, Kristianstad Basin, Skane (Scania) Province, Sweden.

Horizon:

Biostratigraphy: Belemnellocamax mammillatus zone.

Age: Early Campanian Stage, Senonian subepoch, Upper Gulf epoch, Late Cretaceous.

Material:

Number: Not given: 3 marginal teeth.

 

MARTIN, SCHUMACHER PARRIS & GRANDSTAFF, 1998

Locality: Marindahl Dam Spillway, Yankton county, South Dakota.

Horizon: 4.6 meters below the Niobrara/Pierre Contact.

Biostratigraphy:

Age: Late Cretaceous.

Material:

WHO unnumbered: A block of chalk containing frontal, prefrontal, pterygoid, 2 dentaries, jugal, 4 cervical vertebrae, trunk vertebrae and 10 ribs.

 

OM, MANNING, BADGER & WHITLEY, 2016

Locality: Stone Bridge Locality, Cherry Hill, Camden County, southern New Jersey.

Horizon: Upper Woodbury Formation.

Biostratigraphy:

Age: Early-Middle Campanian Stage, Senonian Subepoch, Gulf Epoch, Late Cretaceous.

Material:

Number: Not given: Teeth and caudal vertebra.

RUSSELL, 1970

Locality: Harrell Station Area, Southeast of Marion junction, Dallas County, Alabama.

Horizon: Selma Formation.

Biostratigraphy:

Age: Late Cretaceous.

Material:

P 27324: Disarticulated skull, presacral vertebra.

P 27828: Pterygoid and 6 vertebrae.

P 27439: Parts of skull, atlas and cervical vertebrae.

P 27440: Scapula and coarcoid fragments.

P 27449, 27545: Humerii.

P 27478: Tooth and metacarpal.

P 27484: Fragmentary disarticulated skull and presacral vertebrae.

P 27546: Quadrate.

PR 37: Cranial elements and humerus.

PR 38: Disarticulated skull and vertebrae.

PR 39: Cranial elements and presacral vertebrae.

PR 40: Disarticulated skull, most of vertebral column and appendicular elements.

PR 41: Cranial elements, vertebrae and hind limbs.

PR 42, 43, 46: Vertebrae.

PR 45: Cranial elements, presacral vertebrae and pectoral elements.

PR 47: Presacral vertebrae.

PR 48: Cranial elements, presacral vertebrae and humerus.

PR 49: Pygal vertebrae.

PR 50: Dorsal vertebrae.

PR 52: Disarticulated skull and atlas.

PR 53: Caudal vertebrae.

PR 138: Quadrate, marginal tooth and caudal vertebra.

PR 142: Cranial elements and vertebrae.

PR 192: Pectoral elements.

PR 200: Pygal vertebra and phalanges.

PR 211: Postorbitofrontal.

PR 238: Dentary.

 

Locality: Cedarville, Hale County, Alabama.

Horizon: Selma Formation.

Biostratigraphy:

Age: Late Cretaceous.

Material:

PR 51: Vertebrae.

 

Locality: Boligee, Greene County, Alabama.

Horizon: Selma Formation.

Biostratigraphy:

Age: Late Cretaceous.

Material:

PR 63: Humerus.

 

Locality: Mount Hebron, Greene County, Alabama.

Horizon: Selma Formation.

Biostratigraphy:

Age: Late Cretaceous.

Material:

PR 150: Cranial elements and cervical vertebrae.

 

Locality: West Greene, Greene County, Alabama.

Horizon: Selma Formation.

Biostratigraphy:

Age: Late Cretaceous.

Material:

PR 148: Cranial elements and vertebrae.

PR 156: Cranial elements, vertebrae and humerus.

PR 162: Quadrate and vertebrae.

PR 164: Disarticulated skull, most of vertebral column, and appendicular elements.

PR 175: Dorsal vertebrae.

PR 176: Caudal vertebrae.

PR 183: Radius.

 

Locality: Unknown.

Horizon: Selma Formation?

Biostratigraphy:

Age: Late Cretaceous.

Material:

P 27355 : Dentary and splenial.

 

RUSSELL, 1967

Locality:

Horizon:

Biostratigraphy:

Age:

Material:

AMNH 182:

AMNH R173:

AMNH 1507:

AMNH 1525:

AMNH 1546:

AMNH 1593:

AMNH ND2: Fragmentary skull.

YPM 1308:

YPM 1315:

YPM 1328:

and 13 unnumbered specimens.

USNM 3765:

USNM 3778:

MCZ 1625:

KU 1000:

KU 1005:

KU 1058:

KU 1108:

KU 1124:

KU 1134:

CNHM UR901:

CNHM P 12856:

CM 1190:

BELL, 1997

ANSP 10193:

UCMP 34535:

YPM 1105:

YPM 1310:

SHELDON, 1997

RMM 5830, 1788:

 

Locality: From bluff near A. Finney's ranch, Cottonwood Creek, SSW of Edgemont, Fall River County, South Dakota.

Horizon: Lower Pierre Formation.

Biostratigraphy:

Age: Late Cretaceous.

Material:

AMNH 5812:

 

NICHOLLS, 1988

Locality: South Dakota.

Horizon: Sharon Springs Member, Pierre Shale.

Biostratigraphy:

Age: Campanian Stage, Senonian subepoch, Upper Gulf epoch, Late Cretaceous.

Material:

As per NICHOLLS, 1988

AMNH 5812, FMNH PR 560, 671, KUVP uncatalogued, SDSM PC 8026 (field number), RTMP P.83.19.2, UCM 20568, UCM uncatalogued, USNM 2778-66, 2859-66:

SHELDON, 1996

Locality: Logan County, Kansas.

Horizon: Smoky Hill Chalk Member, Niobrara Formation.

Biostratigraphy:

Age: Santonian or Campanian Stage, Middle Senonian Subepoch, Uppermiddle Gulf Epoch, Late Cretaceous.

Material:

YPM 1318, 1319, 1324, 1310: Adult.

YPM 1316, 1321: Subadult.

YPM 1371,

YPM 1368: Skull of a late juvenile.

YPM 1314: Early juvenile.

LINDGREN & SIVERSON, 2004

YPM 40354: 6th or 7th cervical vertebra.

 

Locality: Wallace County, Kansas.

Horizon: Niobrara County.

Biostratigraphy:

Age: Late Cretaceous.

Material:

YPM 1316: Adult.

YPM 1317: Subadult.

 

Locality: Gove County, Kansas.

Horizon: Niobrara Formation.

Biostratigraphy:

Age: Late Cretaceous.

Material:

YPM 40368: Early juvenile.

 

YAMASHITA, KONISHI, & SATO, 2015

Locality: Southern Manitoba Province, Canada.

Horizon: Pembina Member, Pierre Shale.

Biostratigraphy: Baculites obtusus ammonite zone.

Age: Earliest Middle Campanian Stage, Senonian subepoch, Upper Gulf epoch, Late Cretaceous

Material:

CFDC M74.10.06:

 

= ?Clidastes cineriarum COPE, 1871
Etymology:

Holotype: AMNH ?1579

Locality: 6 miles south of Sheridan, Kansas.

Horizon:

Biostratigraphy:

Age: Late Cretaceous.

Material: Pterygoid teeth and vertebrae.

 

= Edestosaurus dispar MARSH, 1871
Etymology:

Holotype: YPM 1100

Locality:

Horizon:

Biostratigraphy:

Age:

Material:

 

= Edesteosaurus velox MARSH, 1871
Etymology:

Holotype: YPM 1105

Locality: Kansas.

Horizon:

Biostratigraphy:

Age: Late Cretaceous.

Material:

Referred material:

WILLISTON, 1898

Small mandible.

Mandibles.

Fragmentary skulls and fragmentary skeletons.

WILLISTON & CASE, 1892

Locality: Butte Creek, Logan County, Kansas.

Horizon: Smoky Hill Chalk Member, Niobrara Formation.

Biostratigraphy:

Age: Santonian or Campanian Stage, Middle Senonian Subepoch, Uppermiddle Gulf Epoch, Late Cretaceous.

Material:

KUVP 1022: Nearly complete skeleton.

 

Locality: Unknown, Kansas.

Horizon: Smoky Hill Chalk Member, Niobrara Formation.

Biostratigraphy:

Age: Santonian or Campanian Stage, Middle Senonian Subepoch, Uppermiddle Gulf Epoch, Late Cretaceous.

Material:

KUVP 1108: Right quadrate.

 

= Clidastes wymani MARSH, 1871
Etymology:

Holotype: YPM 1120

Locality: North Fork of Smoky Hill, Kansas.

Horizon:

Biostratigraphy:

Age: Late Cretaceous.

Material: Chevron caudal vertebrae.

Referred material:

Number: Not given: Fragmentary skull and anterior vertebrae.

 

= Clidastes pumilus MARSH, 1871
Etymology:

Holotype: YPM 1123

Locality: Logan County, Kansas.

Horizon: Smoky Hill Chalk Member, Niobrara Formation.

Biostratigraphy:

Age: Santonian or Campanian Stage, Middle Senonian Subepoch, Uppermiddle Gulf Epoch, Late Cretaceous.

Material: Quadrate of an early juvenile.

Referred material:

ABEL, 1919

Locality: Western Kansas.

Horizon:

Biostratigraphy:

Age: Late Cretaceous.

Material:

KUVP 69552: Front paddle.

 

= Edestosaurus tortor COPE, 1871
Etymology:
= Clidastes tortor (COPE, 18721) COPE, 1874

Holotype: AMNH 1513

Locality: Fossil Spring canon, Kansas.

Horizon:

Biostratigraphy:

Age: Late Cretaceous.

Material: Fragmentary skull and fragmentary skeleton.

Dentary after WILLISTON, 1898.

 

Referred material:

WILLISTON, 1898

Locality: Smoky Hill River, Kansas.

Horizon: Smoky Hill Chalk Member, Niobrara Formation.

Biostratigraphy:

Age: Santonian or Campanian Stage, Middle Senonian Subepoch, Uppermiddle Gulf Epoch, Late Cretaceous.

Material:

KUVP 1000: Skull, lower jaws, quadrate, vertebral column, left scapula and coracoid, pelvic girdle, parts of front paddle and ribs.

Clidastes velox skull after WILLISTON (1898). A) dorsal, B) lateral and C) ventral views.

Locality: Unknown, Kansas.

Horizon: Smoky Hill Chalk Member, Niobrara Formation.

Biostratigraphy:

Age: Santonian or Campanian Stage, Middle Senonian Subepoch, Uppermiddle Gulf Epoch, Late Cretaceous.

Material:

KUVP 14278: Left quadrate.

KUVP?: Posterior cervical vertebrae.

KUVP?: Frontal bone.

KUVP?: Left quadrate.

 

= Edestosaurus stenops COPE, 1871
Etymology:

Holotype: AMNH 1575

Locality: Near Fossil spring, Kansas.

Horizon:

Biostratigraphy:

Age: Late Cretaceous.

Material: Fragmentary skull.

Referred material:

Locality: On a bluff on Butte Creek, Kansas.

Horizon:

Biostratigraphy:

Age: Late Cretaceous.

Material:

Number: Not given: Fragmentary skeletons.

 

= Edestosaurus rex MARSH, 1872
Etymology:

Holotype: YPM 1310

Locality: Smoky Hill River, Kansas.

Horizon: Niobrara Formation.

Biostratigraphy:

Age: Late Cretaceous.

Material: Fragmentary skeleton.

 

= Clidastes medius MERRIAM, 1894
Etymology:

Holotype:

Locality:

Horizon:

Biostratigraphy:

Age:

Material:

 

= Clidastes westi WILLISTON & CASE, 1892
Etymology:

Holotype: KU 1026

Locality: McAllister, Wallace County, Kansas.

Horizon: Lower Pierre Formation.

Biostratigraphy:

Age: Campanian Stage, Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.

Material: Nearly complete skull, lower jaw, 84 vertebrae, ribs, front and hind paddles.

Clidastes westi dentary modified from WILLISTON, 1898.

_____________________________________________________________________________________

Species: liodontus MERRIAM, 1894
Etymology:

Holotype:

Locality: Logan County, Kansas.

Horizon: Smoky Hill Chalk Member, Niobrara Formation.

Biostratigraphy:

Age: Santonian or Campanian Stage, Middle Senonian Subepoch, Uppermiddle Gulf Epoch, Late Cretaceous.

Material: Maxillae, premaxilla and dentaries.

Top view of skull after RUSELL, 1967.

Referred material:

AMNH 192:

AMNH 1548:

YPM 1333:

YPM 1334:

YPM 1335: Skull.

YPM 3982:

YPM 3996: Fragmentary skull.

YPM 24914:

3 unnumbered specimens at YPM:

USNM 6546:

USNM 11647:

USNM 11719:

MCZ 278:

MCZ 1618:

MCZ 1621:

KU 1039:

KU 1121:

FHM 10668:

PU 17249:

BELL, 1997

KU 1022:

FHM VP2071:

 

Locality: South Dakota.

Horizon: Sharon Springs Member, Pierre Shale.

Biostratigraphy:

Age: Campanian Stage, Senonian subepoch, Upper Gulf epoch, Late Cretaceous.

Material:

KUVP 50104:

As per NICHOLLS, 1988

 

BELL, BARNES & POLCYN, 2007

Localty: Western part of the Big Bend Region, Brewster County, Texas.

Horizon: Lower Pen Formation.

Biostratigraphy:

Age: Upper Coniacian Stage, Lower most Senonian Subepoch-Santonian Stage, Lower Senonian Subepoch, Lower Gulf Epoch, Early Late Cretaceous.

Material:

Number: Not given:

 

SHELDON, 1996

Locality: Logan County, Kansas.

Horizon: Niobrara Formation.

Biostratigraphy:

Age: Late Cretaceous.

Material:

YPM 1367: Late juvenile.

_____________________________________________________________________________________

Species: moorevillensis SHANNON, 1975 (nomen ex dissertatione, nomen nudum)
Etymology: In reference to Mooreville Chalk, Selma Group.

Holotype:

Locality:

Horizon: Mooreville Chalk, Selma Group.

Biostratigraphy: Middle Tylosaurus Acme-zone – Clidastes Acme zone.

Age: Niobraran age, Upper Santonian-Lower Campanian Stage, Senoniain Subepoch, Gulf Epoch, Late Cretaceous.

Material:

Referred material:

BEHLKE, OTT, KELLY & SPEARING, 2002

Loclaity: Logan County, Kansas.

Horizon: Niobrara Formation.

Biostratigraphy:

Age: Early Campanian Stage, Gulf Epoch, Late Cretaceous.

Material:

Number: Not given: Nealry complete specimen.
Note: 7 meters long.

 

BELL, 1997

Locality:

Horizon:

Biostratigraphy:

Age:

Material:

RMM 070:

GSA TC218:

RMNH PR495: