Genus: Exostinus COPE, 1873
Etymology:
= Harpaosaurus GILMORE, 1928
Etymology:
= Prionosaurus GILMORE, 1928
Etymology:

Species: lancensis GILMORE, 1928
Etymology: In reference to Lance Creek, Wyoming, where the specimen was found.

Referred material:

Locality: Lance Creek, Niobrara County, Wyoming.

Horizon: Lance Formation.

Biostratigraphy:

Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.

Material:

USNM 10689: Right maxilla lacking anterior end.

 

Locality: UCMP V-5003, S 1/2, Sec. 20, T37N, R64W, Lance Creek, Powder River Basin, Niobrara County, Wyoming.

Horizon: Lance Formation.

Biostratigraphy:

Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.

Material:

Number: Not given: A few fragments of jaws.

 

Locality: UCMP V-5620, Lull Quarry, near the head of a tributary of Buck Creek, Powder River Basin, Niobrara County (Formerly Converse County), Wyoming.

Horizon: Lance Formation.

Biostratigraphy:

Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.

Material:

UCMP 46053: Left dentary broken posteriorly.

UCMP 46090: Anterior end of right dentary.

UCMP 46606: Anterior end of left dentary.

UCMP 46076: Anterior end of left dentary.

UCMP 47772: Posterior end of right dentary.

UCMP 47829: Almost complete right dentary, slightly broken anteriorly and posteriorly.

UCMP 50015: Complete left dentary of young individual.

UCMP 50016: Median fragment of left maxilla.

UCMP 50024: Posterior end of left maxilla.

UCMP 56091: Fragmentary parietal.

UCMP 56092: Fragmentary parietal.

 

Locality: UCMP V-5711, Bushy Tailed Blowout, Quarry area includes the floor of a small blowout approximately 10 yards northwest of the highest point on the ridge forming the northeastern side of the blowouts of loc. V-5003, S 1/2, Sec. 20, T37N, R64W, Lance Creek, Powder River Basin, Niobrara County, Wyoming.

Horizon: Lance Formation.

Biostratigraphy:

Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.

Material:

UCMP 49847: Right maxillary fragment.

UCMP 49848: Anterior end of left dentary slightly broken anteriorly.

UCMP 49849: Median fragment of left maxilla showing sculpture.

UCMP 49850: Posterior end of left maxilla.

UCMP 49864: Fragment of left dentary.

UCMP 56093: Almost complete parietal.

A few jaw fragments.

GAO & FOX, 1996

UALVP 29847: Parietal.

UALVP 29849-29850: Incomplete maxillaries.

UALVP 29838: Dentary.

UALVP 29920: Jugal.

 

BREITHAUPT, 1982

Locality: East flank of Rock Springs Uplift, Sweetwater County, Wyoming.

Horizon: Lance Formation.

Biostratigraphy:

Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.

Material:

UW 14136-14139: Right dentary fragments.

UW 14140: Left dentary fragment.

UW 14141: Left dentary fragment.

 

BRYANT, 1990

Locality: Eastern Montana.

Horizon: Hell Creek Formation.

Biostratigraphy:

Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.

Material:

UCMP 130734: Dentary fragment.

UCMP 123321: Maxilla.

UCMP 130731: Parietal.

 

ESTES, BERBERIAN & MESZOELY, 1969

Locality: Bug Creek Anthills locality, McCone County, Montana.

Horizon: Hell Creek Formation.

Biostratigraphy:

Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.

Material:

MCZ 3662a: Dentary and maxillary fragments.

MCZ 3662b: Frontal.

GAO & FOX, 1996

UALVP 29929: Frontal.

 

GAO & FOX, 1996

Localities: UC V-5616, Altman Blowout, this large blowout, site of the UCMP field-party camps from 1956 to 1960, is just east of the crest of the divide separating Lance Creek and Hancock Creek and 100 yards east of a dirt road. Fossils were found in anthills, talus, and sandstone exposed in the blowout, which is in the NE 1/4, Sec. 9, T37N, R64W, Eastern Wyoming.

Horizon: Lance Formation.

Biostratigraphy:

Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.

Material:

UALVP 29848: A parietal.

 

SAHNI, 1972

Locality: Clambank Hollow, just north of the Judith River on the Missouri River in north-central Montana.

Horizon: Judith River Formation.

Biostratigraphy:

Age: Judithian age, Campanian Stage, Senonian subepoch, Upper Gulf epoch, Late Cretaceous.

Material:

AMNH 8497: Right maxillary fragment.

 

= Harpaosaurus parvus GILMORE, 1928
Etymology:
= Marpagosaurus parvus (GILMORE, 1928) ESTES, 1964
Etymology:

Holotype: USNM 10803

Locality: "Peterson's Quarry," Lance Creek, Powder River Basin, Niobrara County, Wyoming.

Horizon: Lance Formation.

Biostratigraphy:

Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.

Material: Right maxilla with 14 teeth.

 

= Prionosaurus regularis GILMORE, 1928
Etymology:

Holotype: USNM 10750

Locality: Lance Creek, Niobrara County, Wyoming.

Horizon: Lance Formation.

Biostratigraphy:

Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.

Material: Incomplete maxilla.