Genus: Mosasaurus CONYBEARE, 1822
Etymology: Latin, mosa, "meuse river" and Greek, sauros, "lizard."
= Amphekepubis MEHL, 1930
Etymology:
= Batrachiosaurus HARLAN, 1839
Etymology:
= Batrachotherium HARLAN, 1839
Etymology:
= Pterycollosaurus DOLLO, 1882
Etymology:

Species: hoffmanni MANTELL, 1829
Etymology:

Holotype: MNHN AC. 9648

Locality: Subterraneous Quarry at St. Peter's Mount, near St. Petersburg, Maastricht, Limburg Province, The Netherlands.

Horizon: Maastricht Formation.

Biostratigraphy:

Age: Upper Maastrichtian Stage, uppermost Senonian subepoch, Gulf Epoch, Late Cretaceous.

Material: A disarticulated partial skull, 2 cervical vertebrae and a femur.

Mosasaurus hoffmannii after LINGHAM-SOLIAR, 1995: A) dorsal view based on IRSNB R26 and NHMUK 42929; B) lateral view based on the holotype and IRSNB R26; C) ventral view based on IRSNB R12 and NHMUK 42929.

Mosasaurus hoffmanni MNHN AC 9648 after STREET & CLADWELL, 2016.

OWEN, 1854

ROYAL COLLEGE OF SURGEONS 316: Cast of type.

Referred material:

IRSNB R 25 (R1503): Fragmentary premaxilla and left dentary.
Note: An immature individual.
Note: Paleopathologic with fracture and bony callus with infection.

IRSNB R26 (R1483): Fragmentary skull and lower jaw.

IRSNB 1621:

IRSNB IG 8444:

NHMUK (old BMNH)  42928/9:

NHMUK (old BMNH) R1223/4/5:

NHMM 009002:

NHMM 006696-48: Premaxillae.

NHMM 1989127:

NHMM 603092:

TSMNH 11201/2:

TSMNH 11243/4:

TSMNH 11208:

TSMNH 11376:

TSMNH 17281:

BARDET, 2012

Gg2004/5527, /5528: 2 teeth.

Gg2004/5536: A cervical vertebra.

Gg2004/34880: A caudal vertebra.

LINGHAM-SOLIAR, 2004

IRSNB R 27: Right dentary and associated postcranial material consisting a major part of the vertebrale column, some ribs and limb bones.
Note: An immature individual.
Note: Paleopathology, the dentary shows a well formed bony callus surrounding a fracture non-union.

SPAMER, DAESCHLER & VOSTREYS-SHAPIRO, 1995

ANSP 15731: Cast of type.

MULDER, 1999

MND 20.01601: Tooth.

MNHNP AC 9648: Left dentary.

JAGT, 2003

NHMM 003915: Left coracoid.
Note: Has impressions of the ichnofossil Radulichnus inopinatus, gastropod or chiton, probably chiton (JAMISON-TODD, UPCHURCH & MANNION, 2023).

JAGT, MULDER, GALLAGHER & SCHULP, 1999

Dentary and maxilla with teeth, articulated and associated vertebrae.
Note: Found in the Gulpen Formation, lanaye Member, overlying Maastricht Formation.

After GIBBS, 1849.

 

BARDET, 2012

Locality: Meudon, Ile-de-France region, Hauts-de-Seine Department, France.

Horizon: Meudon Chalk.

Biostraigraphy: B. mucronata zone.

Age: Upper Campanian Stage, Senonian subepoch, Gulf Epoch, Late Cretaceous.

Material:

MNHN 877: Tooth.

 

= cf. Mosasaurus hoffmannii BOLES, ULLMANN, PUTNAM, FORD & DECKHUT, 2024

Locality: Edelman Fossil Park, Mantua Township, Gloucester Country, New Jersey.

Horizon: Navesink Formation, Hornerstown Formation.

Biostratigraphy:

Age; Danian Stage, Early Paleocene Epoch, Early Paleogene Period, Tertiary Sub-era, Early Cenozoic Era.

Material:

RU-EFP-03592: Nearly complete right pterygoid (MFL).

RU-EFP-00109, 01826, 01901, 02703, 02916, 00120-1-00120-4: 9 isolated teeth (MFL).

RU-EFP-00033, 02645, 03640, 00043-1-00043-4: 7 isolated vertebral centra.

 

FARKE & PHILLIPS, 2017

Locality: MMNS Locality MS.73.001b, Union County, Mississippi.

Horizon: Owl Creek Formation.

Biostratigaphy:

Age: Late Maastrichtian Stage, Senonian Subepoch, Gulf Epoch, Late Cretaceous.

Material:

Number: Not given.

 

= Mosasaurus sp aff. M. hoffmanni MATEUS, POLCYN, JACOBS, ARAUJO, SCHULP, MARINHEIRO, PEREIRA & VINEYARD, 2011

Locality: Baba, Bentiaba, Fazenda Dos Cavaleiros, Namibe Province, Angolia.

Horizon:

Biostratigraphy:

Age: Middle ti yooer Maastrichtian Stage, Late Senonian Stage, Late Gulf Epoch, Late Cretaceous.

Material:

MGUAN-PA35: Isolated shed teeth, complete and partial humeri.

 

 

= Mosasaurus cf. hoffmanni MUSTAFA & ZALMOUT, 2001

Locality: At-Tayyba Area, west of Irbid City, North west Jordan.

Horizon: Upper Part of Amman Formation.

Biostratigraphy:

Age: Early Maastrichtian Stage, Late Senonian Subepoch, Late Gulf Epoch, Late Cretaceous.

Material:

YUPRT1: Isolated tooth.

 

REMPERT, MARTENS & VINKELES MELCHERS, 2024a

Locality: Union County, Mississippi.

Horizon: Owl Creek Formation, Prairie Bluff Chalk.

Biostratigraphy:

Age: Upper Maastrichtian Stage, Late Senonian Subepoch, Late Gulf Epoch, Late Cretaceous.

Material:

REMPC M-190: Marginal tooth crown.

 

= Mosasaurus cf. hoffmannii REMPERT, MARTENS & MELCHERS, 2024b

Locality: Holden Beach, Brunswick County, North Carolina.

Horizon: Peedee Formation.

Biostratigraphy:

Age: Upper Maastrichtian Stage, Late Senonian Subepoch, Late Gulf Epoch, Latest Cretaceous.

Material:

VANPS 13.0180: A large marginal tooth crown.

 

REMPERT, MELCHRS, REMPERT, HAQUE & ARMSTRON, 2022

Locality: Sidi Daoui Phosphate Quarry, Oulad Abdoun Basin, Morocco.

Horizon: Upper Couche III Layer.

Biostratigraphy:

Age: Upper Maastrichtian Stage, Late Senonian Subepoch, Late Gulf Epoch, Late Cretaceous.

Material:

REMPC M0001: A marginal tooth crown from the median of the jaw.

 

Locality: Sidi Chennane Phosphate Quarry, Oulad Abdoun Basin, Morocco.

Horizon: Upper Couche III Layer.

Biostratigraphy:

Age: Upper Maastrichtian Stage, Late Senonian Subepoch, Late Gulf Epoch, Late Cretaceous.

Material:

REMPC M0002, M0003: 2 isoalted marginal tooth crowns.

 

Locality:Unknown, Oulad Abdoun Basin, Morocco.

Horizon: Upper Couche III Layer.

Biostratigraphy:

Age: Upper Maastrichtian Stage, Late Senonian Subepoch, Late Gulf Epoch, Late Cretaceous.

Material:

AVM 01: A premaxillary tooth crown.

AVM 02: A marginal maxillary tooth crown.

CORN 01: A marginal tooth crown.

 

SCHULP, WALENKAMP, HOFMAN, STUIP & ROTHSCHILD, 2006

NHMM 006696: Fragmentary skull and dentary.

Brackdown as follows.

NHMM 006696-36: Left quadrate.

NHMM 006696-38: Left surangular.

NHMM 006696-39: Left articular.

NHMM 006696-40: Left angular.

Note: All pathologic.

NHMM 006696-41: Right surangular.

NHMM 006696-42: Right articular.

NHMM 006696-43: Rgith angular.

NHMM 006699: Partial right dentary (same individual as above).

 

Locality: Zichem, Calcarenite, Maastricht, Limburg Province, The Netherlands.

Horizon: Maastricht Formation.

Biostratigraphy:

Age: Upper Maastrichtian Stage, uppermost Senonian subepoch, Gulf Epoch, Late Cretaceous.

Material:

IRSNB R12: Skull and lower jaw of a subadult.
Note: The brain-cast shows severing and displacement of the brain stem do to a ‘blow’ to the skull, possibly by a tylosaurinae. Also the coronoid and surangular of the upper postmandibular unit also on the right side are dislocated.

IRSNB IG8427:

 


Locality: Kanne, Maastricht Calcerenite, The Netherlands.

Horizon: Maastricht Formation.

Biostratigraphy:

Age: Upper Maastrichtian Stage, uppermost Senonian subepoch, Gulf Epoch, Late Cretaceous.

Material:

IRSNB EFR27 (R1559): Left quadrate.

 


Locality: Kunrade, Maastricht Calcerenite, The Netherlands.

Horizon: Maastricht Formation.

Biostratigraphy:

Age: Upper Maastrichtian Stage, uppermost Senonian subepoch, Gulf Epoch, Late Cretaceous.

Material:

IRSNB 1626:

 


Locality: Near Maastricht, Limburg Province, The Netherlands.

Horizon: Upper Gulpen Formation, Lanaye Chalk.

Biostratigraphy:

Age: Maastrichtian Stage, uppermost Senonian subepoch, Gulf Epoch, Late Cretaceous.

Material:

IRSNB R24 (R1506):

Mr. Grarcet’s specimen.

 


Locality: St. Symphorien (Carriere Hardenport), The Netherlands.

Horizon: Ciply Phosphatic Chalk.

Biostratigraphy:

Age: Upper Maastrichtian Stage, uppermost Senonian subepoch, Gulf Epoch, Late Cretaceous.

Material:

IRSNB IG8697:

 

Locality: Maestricht, Limburg Province, The Netherlands.

Horizon:

Biostratigraphy:

Age: Upper Maastrichtian Stage, uppermost Senonian subepoch, Gulf Epoch, Late Cretaceous.

Referred material:

IRSNB 1559, 'R12': Fragmentary skull and skeleton.

 

BARDET, MARTIN, CORRAL, PEREDA SUBERBIOLA, & ASTIBIA, 2013

Locality: Disused sand quarry near the village of Lano, Lano quarry, Trevino County, about 20 km SE of the City of Vitoria, Basque-Cantabric Basin, Basque Province, Iberian Peninsula, Spain.

Horizon: Sobrepena Formation.

Biostratigraphy:

Age: Uppermost Campanian-Lowermost Maastrichtian Stage, Upper Senonian subepoch, Upper Upper Gulf Epoch, Late Cretaceous.

Material:

LU-GMS/LAN004: A marginal tooth from the anterior part of teh jaws.


BARDET & TUNOGLU, 2002

Locality: Beyler Dam, Devrekani Town, Kastamonu City area, Northern Turkey.

Horizon: Davutlar Formation.

Biostratigraphy:

Age: Late Maastrichtian Stage, uppermost Senonian subepoch, Gulf Epoch, Late Cretaceous.

Material:

HU. JMB-0057-99: Remains of a large skull including 2 maxillary fragments with teeth, a partial left jugal, several isolated teeth and indeterminate bones, probably from the palate, preserved in several blocks as part and counterpart.


DORTANGS, JAGT, MULDER & SCHULP, 1999

Locality: ENCI-Maastricht BV Quarry, south of Masstricht, Limburg Province, The Netherlands.

Horizon: Upper Lanaye Member, Gulpen Formation.

Biostratigraphy:

Age: Late Maastrichtian Stage, uppermost Senonian subepoch, Gulf Epoch, Late Cretaceous.

Material:

NHMM 1988141: Skull and fragmentary skeleton.

 

CALLAHAN, JOHNSON & MEHLING, 2010

Locality: Spheno Run, Maryland.

Horizon: New Egypt Formation, Monmouth Group.

Biostratigraphy:

Age: Late Maastrichtian Stage, uppermost Senonian subepoch, Gulf Epoch, Late Cretaceous.

Material:

Number: Not given:

 

= Mosasaurus sp. aff. M. hoffmanni FERNANDEZ, MARTIN & CASADIO, 2008

Locality: Locality A., S39°46'20", W66°40'42", Trapalco and Santa Rosa depression, Rio Negro Province, Northern Patagonia, Argentina.

Horizon: Upper part of the Jagüel Formation.

Biostratigraphy:

Age: Upper Maastrichtian Stage, Uppermost Senonian Subepoch, Uppermost Gulf Epoch, Late Cretaceous.

Material:

MML-PV1: Fragments of both dentaries, left parital premaxilla, anterior portion of both left and right maxillae, lateral fragment of the frontal, proimal portion of the humerus with the head ane postglenoid process, anterior fragment of the left angular, metacarpal I, 5 fragmentary teeth, atlas intercentrum and 1 trunk vertebrae.

 

GALLAGHER, 2014

Locality: West Jersey Marl Company works, Barnsboro, Gloucester County, New Jersey.

Horizon: Hornerstown Formation, Rancocas Group.

Biostratigraphy:

Age: Late Maastrichtian Stage, uppermost Senonian subepoch, Gulf Epoch, Late Cretaceous.

Material:

YPM 773: Skull and vertebrae.

 

GRIGORIEV, 2014

Locality: Outcrop in the Prolom ravine, within the city of Penza, near the Mironositskoe cemetery, European Russia.

Horizon:

Biostratigraphy:

Age: Maastrichtian Stage, Uppermost Senonian Subepoch, Uppermost Gulf Epoch, Late Cretaceous.

Material:

CCMG 10/2469, PRM 2546: Fragmentary skull including two dentaries, with one preserved replacement tooth in the alveolar maring on the left ramus (?), right and left splenials, the right, angular in articulation with the right surangular, coronoid, articular and prearticular, left surangular, articular and prearticular in the articulation as well.

Plaster copy: Prm 1-5/2546: Right dentary consistion of five parts with eight teeth sitting in the alveolar margins.

PRM 6-10/2546: Left dentary consisting of six parts with three teeth sitting in the alveoalr margins and one speerate tooth with the root.

PRM 13, 14/2546: Right and left splenials.

PRM 16/2546: Right angular in articulations with the right surangular, coronoid, articular and prearticular.

PRM 17/2546; Left angular.

PRM 19/2546: Left postorbitofrontal.

PRM 22/2546: Left squamosal.

PRM 21/2546: Parietals.

PRM 23/2546: elements of the scapulo-coracoid.

Numbers: Not given: Frontals, and ribs, however, they are not preserved to the present day.

 

HARRELL & MARTIN, 2014

Locality: SDSMT locality number V9325, as float in the channel of the Morequ River, approximately 2.0 km west of the bridge crossing State Highway 64, Zeibach County, South Dakota.

Horizon: Trail City Member, Fox Hills Formation.

Biostratigraphy:

Age: Middle Maastrichtian Stage, Uppermost Senonian Subepoch, Uppermost Gulf Epoch, Late Cretaceous.

Material:

TLAM NH.HR.2009.032.0001: Fragmentary skull.

 

HARRELL JR, & PEREZ-HUERTA, 2015

Locality: ALn-3, Sandy Ridge, Lowndes County,  Alabama.

Horizon: Ripley Formation.

Biostratigraphy:

Age: Early Maastrichtian Stage, Senonian subepoch, Gulf Epoch, Late Cretaceous.

Material:

ALMNH PV 1990.0003: Skeleton.

 

Locality: ALn-7, Braggs, Lowndes County,  Alabama.

Horizon: Prairie Bluff Formation.

Biostratigraphy:

Age: Maastrichtian Stage, Senonian subepoch, Gulf Epoch, Late Cretaceous.

Material:

ALMNH PV 1988.0018: Skeleton.

ALMNH PV 1991.0013.0002: Maxilla fragment.

ALMNH PV 1998.0018: Associated ribs.

 

Locality: Orion, Pike County,  Alabama.

Horizon: Ripley Formation.

Biostratigraphy:

Age: Early Maastrichtian Stage, Senonian subepoch, Gulf Epoch, Late Cretaceous.

Material:

ALMNH PV 1994.0001.0033: Isolated vertebra.

 

JAGT, BARTEN, BARTEN, CORNELISSEN, DORTAGS, EYSERMANS, JAGT-YAZYKOVA, RENKENS, & VERDING, 2015

Locality: ENCI-Maastricht BV Quarry, south of Masstricht, Limburg Province, The Netherlands.

Horizon: Low Nekum Member/high in the Emael Member, Mascht Formation.

Biostratigraphy: Belemnitella junior zone.

Age: Late Maastrichtian Stage, uppermost Senonian subepoch, Gulf Epoch, Late Cretaceous.

Material:

Number: Not given (nicknamed 'Lars'): Fragmentary skeleton.

 

JAGT, DECKERS, LEEBEECK, DONOVAN & NIEUWENHUIS, 2020

Locality: ENCI-Heidelbertg Cement Group Quarry, south of Masstricht, Limburg Province, The Netherlands.

Horizon: Base of the Valkenburg Member, Mascht Formation.

Biostratigraphy:

Age: Late Maastrichtian Stage, uppermost Senonian subepoch, Gulf Epoch, Late Cretaceous.

Material:

NHMM MD 5256.01: Fragmentary dorsal vertebra.

Note: With Episkeletozoans and bioerosial traces of Gnathicnhus pentax from regular echnoid on a vertebra, JAMISON-TODD, UPCHURCH & MANNION, 2023.

NHMM 2019 003: Fragmenary dorsal vertebra.

Note: With bioerosial traces of Gnathicnhus pentax from regular echnoid on a vertebra, JAMISON-TODD, UPCHURCH & MANNION, 2023.

 

= Mosasaurus cf. hoffmanni JAGT, MOTCHUROVA-DEKOVA, IVANOV, CAPPETTA, & SCHULP, 2007

Locality: Labirinta Cave, situated between the villages of Drashan and Breste, east of Cherven Briag, Vratsa district, northwest Bulgaria.

Horizon: Kajlaka Formation.

Biostratigraphy:

Age: Maastrichtian Stage, Uppermost Senonian Subepoch, Uppermost Gulf Epoch, Late Cretaceous.

Material:

NMNHS 11897: A partial lower jaw with 2 teeth associated.

 

JAGT, SCHULP, SEVERIJNS, CORNELISSEN, VERDING, & MULDER, 2007, JAGT, CORNELISSEN, MULDER, SCHULP, SEVERIJNS & VERDING, 2008

Locality: Ankerpoort-Curfs quarry, Geulheim, Valkenburg aan de Geul, Liege Province, The Netherlands.

Horizon: Uppermost Meerssen Member, Maastricht Formation.

Biostratigraphy:

Age: Late Maastrichtian Stage, uppermost Senonian subepoch, Gulf Epoch, Late Cretaceous.

Material:

NHMM 2007 093: Left and right dentaries and maxillae, both coronoids incomplete, left spleninal, isolated teeth and crowns and numerous small, unidentifed bone fragments.

 

= Mosasaurus cf. hoffmanni LINDGREN & JAGT, 2005

Locality: Near the town of Aalborg, northern Jyllard, Denmark.

Horizon: Mons Klint Formation.

Biostratigraphy:

Age: Lower? Maastrichtian Stage, Upper Senonian Subepoch, Upper Gulf Epoch, Late Cretaceous.

Material:

MGUH 27472: Shed fragmentary tooth.

NHMD 226499: Tooth.

Locality: Stevns Klint, Stevns Peninsula, Denmark.

Horizon: Hojerup Member or Sigerslev Member.

Biostratigraphy:

Age: Maastrichtian Stage, uppermost Senonian subepoch, Gulf Epoch, Late Cretaceous.

Material:

OESM 8783, NHMD 227349: Teeth.


= Mosasaurus cf. hoffmanni LINGHAM-SOLIAR, 1991

Locality: Mont. Igdama, Illemmeden Basin of Niger, Central West Africa.

Horizon: Dukamaje Formation, ("Mosasaurus" Shales).

Biostratigraphy:

Age: Late Cretaceous.

Material:

BMNH R11913: Fragment of a large tooth crown.

 

LINGHAM-SOLIAR, 1994

Locality: Manzadi, Bas Congo, Democratic Republic of the Congo (Formerly the Belgian Congo, Zaire).

Horizon:

Biostratigraphy:

Age: Maastrichtian Stage, uppermost Senonian subepoch, Gulf Epoch, Late Cretaceous.

Material:

Number: Not given: 2 teeth.

 

= Mosasaurus cf. hoffmanni MACHALSKI, JAGT, DORTANGS, MULCER & RADWANSKI, 2003

Locality: Nasilow, Poland.

Horizon: Danian Greensand.

Biostratigraphy:

Age: Danian Stage, Lower Paleocene Epoch, Paleogene Peroid, Lower Tertiary subear, Cenozoic era.
Note: Reworked from the uppermost Maastrichtian Stage, uppermost Senonian subepoch, Gulf Epoch, Late Cretaceous.

Material:

IGPUW AR-1, IGPUW AR-2: 2 teeth.
Note: Found together and possibly belonging to the same single individual.

ITPUW AR-3: Tooth crown.


Locality: Opoka at Nasilow, Poland.

Horizon:

Biostratigraphy:

Age: Upper Maastrichtian Stage, uppermost Senonian subepoch, Gulf Epoch, Late Cretaceous.

Material:

IGPUW AR-4, OGP 1254: Tooth crowns.


OWEN, 1854

Locality: Near Maestricht, Limburg Province, The Netherlands.

Horizon:

Biostratigraphy:

Age: Upper Maastrichtian Stage, uppermost Senonian subepoch, Gulf Epoch, Late Cretaceous.

Material:

ROYAL COLLEGE OF SURGEONS 317: Dorsal centrum.

ROYAL COLLEGE OF SURGEONS 318: Tooth.

 


Locality: St. Peter’s Mount, Maestricht, Limburg Province, The Netherlands.

Horizon:

Biostratigraphy:

Age: Upper Maastrichtian Stage, uppermost Senonian subepoch, Gulf Epoch, Late Cretaceous.

Material:

ROYAL COLLEGE OF SURGEONS 319: Small teeth from the pterygoid.

 

PERSSON, 1959

Localities: Blaksudden, Ivo, Skane (Scania) Province, Sweden.

Horizon:

Biostratigraphy:

Age: Lower Campanian Stage, Senonian subepoch, Gulf Epoch, Late Cretaceous.

Material:

Number: Not given: 29 teeth, Balsberg 1 tooth and Ignaberga 1 tooth, 31 tooth crowns belonging LM 3, RM 23, SM 2 and UM 3.

 

= cf Mosasaurus hoffmanni (or new genus) PALCI, CALDWELL, PAPAZZONI & FORNACIARI, 2014

Locality: San Valentino, Province of Salerno, Italy.

Horizon: Either Argille di Viano Formation (early Paleocene-Middle Eocene, but containing reworked boulders coming from the Flysch di Monte Cassio Formation, Late Campanian-Late Maastrichtian) or from the Argille Varicolori di Cassio Formation (late Cenomanian-Late Campanian).

Material:

IPU 30200: Fragmentary snout.

 

Locality: Marzana, Province of Pavia, Italy.

Horizon: Scaglia Rossa Formation.

Biostratigraphy:

Age: Turonian Stage, Uppermost Gallic Subepoch, Lower Gulf Epoch, Late Cretaceous-Maastrichtian Stage, uppermost Senonian subepoch, Gulf Epoch, Late Cretaceous.

Material:

NHMV V1001: Fragmentary skull.


SULIMSKI, 1968
Note: = Mosasaurus cf. hoffmanni MACHALSKI, JAGT, DORTANGS, MULCER & RADWANSKI, 2003

Locality: Maruszow, Poland.

Horizon:

Biostratigraphy:

Age: Upper Campanian Stage, Upper Senonian Subepoch, Upper Gulf Epoch, Late Cretaceous.

Material:

ZPAL R. I/1-5: A fragmentary right upper jaw with 3 teeth (mostly bone bases.)

 

= Mosasaurus cf. hoffmanni TANIMOTO, SATO & TAKATA, 2004, TANIMOTO & YOKOI, 2005
Note: Orginally refered to Mosasaurus cf. conodon.

Locality: Sobura, Kaizuka City, Osaka Prefecture, Honshu Island, Western Japan.

Horizon: Azenotani Formation, Izumi Group.

Biostratigraphy:

Age: Maastrichtian Stage, Upper Senonian Subepoch, Upper Gulf Epoch, Late Cretaceous.

Material:

Number: Not given: Fragmentary lower jaw and teeth.

 


= Mosasaurus belgicus HOLL, 1829
Etymology:

Holotype:

Locality:

Horizon:

Biostratigraphy:

Age:

Material:

 

= Mosasaurus camperi MEYER, 1832
Etymology:

Holotype:

Locality: Sweden.

Horizon:

Biostratigraphy:

Age: Upper Senonian subepoch, Upper Upper Gulf Epoch, Late Cretaceous.

Material: Teeth.

Referred material:

LYDEKKER, 1888

Locality: St. Peter's Mount, near Maastricht, Limburg Province, The Netherlands.

Horizon: Late Cretaceous.

Material:

NHMUK (old BMNH) 11589: Cast of imperfect skull.

NHMUK (old BMNH) 42939: Anterior extremity of the united premaxillae of a juvenile.

NHMUK (old BMNH) 42928: Part of the roof of a cranium.

NHMUK (old BMNH) 42930: Under surface of part of the superior process of the premaxillae, and the anterior extremities of the prefrontals.

NHMUK (old BMNH) 42935: The conjoint basioccipital and basisphenoid.

NHMUK (old BMNH) 42949: Imperfect left quadrate.

NHMUK (old BMNH) 42975: Imperfect left quadrate.

NHMUK (old BMNH) R 1223: Fragmentary left maxilla with 3 teeth.

NHMUK (old BMNH) 42940: Fragment of maxilla.

NHMUK (old BMNH) 42936: Part of the left maxilla.

NHMUK (old BMNH) 42932: Pterygoid with 2 teeth.

NHMUK (old BMNH) R 1224: Dentary portion of the right ramus of the mandible.

NHMUK (old BMNH) 42931: Anterior portion of the right ramus of the mandible of a juvenile.

NHMUK (old BMNH) 42938: Part of a small dentary with imperfect teeth.

NHMUK (old BMNH) 42950-1: Undetermined bones of the skull.

NHMUK (old BMNH) 42997: Right splenial.

NHMUK (old BMNH) 42939: Jaw.

NHMUK (old BMNH) 4291: A number of crowns of small teeth.

NHMUK (old BMNH) 42941: Crown a of a very large tooth.

NHMUK (old BMNH) 42929: Cervical vertebrae and imperfect bone of the skull.

NHMUK (old BMNH) R 1225: Tooth, imperfect humerus, centrum of a vertebra, portions of ribs and metapodial bones.

NHMUK (old BMNH) 42933: Several trunk vertebrae and a small imperfect quadrate and the root of a tooth.

NHMUK (old BMNH) 42946: Imperfect cervical vertebra.

NHMUK (old BMNH) 42946a: Imperfect vertebra.

NHMUK (old BMNH) 42946b: Imperfect centrum of a small cervical vertebra.

NHMUK (old BMNH) 42946c: Imperfect cervical vertebra.

NHMUK (old BMNH) 42943: Imperfect neural arch of a cervical or dorsal vertebra.

NHMUK (old BMNH) (old BMNH) NH 42946d: Imperfect posterior trunk vertebra.

NHMUK (old BMNH) 42946e: Imperfect centrum of posterior trunk vertebra.

NHMUK (old BMNH) 42946f: Centrum of a posterior trunk vertebra.

NHMUK (old BMNH) 42946g: Imperfect centrum of a posterior trunk vertebra.

NHMUK (old BMNH) 42946H: Centrum of a posterior trunk vertebra.

NHMUK (old BMNH) 42946i: Imperfect centrum of a smaller trunk vertebra.

NHMUK (old BMNH) 42946j, k: 2 similar but smaller centra.

NHMUK (old BMNH) 42946l: 3 imperfect posterior trunk vertebrae.

NHMUK (old BMNH) 42934: Anterior or middle caudal vertebra with the interior chevron attached.

NHMUK (old BMNH) 42946m: Centrum of a smaller vertebra.

NHMUK (old BMNH) 42946n: The centra of 2 posterior caudal vertebra.

NHMUK (old BMNH) 42946o: 6 imperfect posterior caudal vertebrae.

NHMUK (old BMNH) 42944: Imperfect posterior caudal vertebra.

NHMUK (old BMNH) 11589a: Cast of a bone witch is probably an ulna.

NHMUK (old BMNH) 42961: Carpus?

NHMUK (old BMNH) 42943, 42945: 2 imperfect undetermined bones.

 

SCHRODER, 1885

Locality: Old Prussia.

Horizon:

Biostratigraphy:

Age: Upper Senonian subepoch, Upper Upper Gulf Epoch, Late Cretaceous.

Material:

Number: Not given: Schwanzwirbel.

 

Locality: Sweden.

Horizon:

Biostratigraphy:

Age: Upper Senonian subepoch, Upper Upper Gulf Epoch, Late Cretaceous.

Material:

Number: Not given: Sp 1, and sp 2: Teeth.

 

= Mosasaurus maximus COPE, 1869
Etymology:

Holotype: AMNH 1389

Locality: Middletown, Monmouth County, New Jersey.

Horizon: Navesink Formation, Monmouth Group.

Biostratigraphy:

Age: Middle Maastrichtian Stage, uppermost Senonian subepoch, Gulf Epoch, Late Cretaceous.

Material: Quadrate, jaw fragments and 7 vertebrae.

Referred material:

BAIRD, 1986

Locality: Loc. A, Prince George’s County, Maryland.

Horizon: Severn Formation.

Biostratigraphy:

Age: Early Late Maastrichtian Stage, uppermost Senonian subepoch, Gulf Epoch, Late Cretaceous.

Material:

MAPS A1218a: Thoracic Vertebra.

 


Locality: Loc. B, Prince George’s County, Maryland.

Horizon: Severn Formation.

Biostratigraphy:

Age: Early Late Maastrichtian Stage, uppermost Senonian subepoch, Gulf Epoch, Late Cretaceous.

Material:

Hartsein Coll: Tooth.

 


Locality: Loc. C, Prince George’s County, Maryland.

Horizon: Severn Formation.

Biostratigraphy:

Age: Early Late Maastrichtian Stage, uppermost Senonian subepoch, Gulf Epoch, Late Cretaceous.

Material:

MAPS A1202c: Tooth.

 

GALLAGHER, CAMPBELL, JAGT, & MULDER, 2005

Locality: From a pit located on Crowleys Ridge, approximately 10 km northeast of Bloomfield, Stoddard County, near the extreme southeastern "boot hill" of Missouri.

Horizon: Owl Creek Formation.

Biostratigraphy:

Age: Latest Late Maastrichtian Stage, uppermost Senonian subepoch, Gulf Epoch, Late Cretaceous. Near the K/T boundary.

Material:

NJSM 20676: Tooth.

NJSM 20677: Vertebra.

 

GALLAGHER, PARRIS & SPAMER, 1986

Locality: Inversand marl pit in Sewell, Mantua Township, Gloucester County, New Jersey.

Horizon: Navesink (= New Egypt) Formation, Monmouth Group.

Biostratigraphy:

Age: Middle Maastrichtian Stage, uppermost Senonian subepoch, Gulf Epoch, Late Cretaceous.

Material:

NJSM 11053, 11052: 2 complete skulls.

 


Locality: Big Brook, New Jersey.

Horizon: Mount Laurel ("Wenonah") Formation, Monmouth Group.

Biostratigraphy:

Age: Lower Maastrichtian Stage, Upper Senonian Subepoch, Upper Gulf Epoch, Late Cretaceous.

Material:

Number: Not given:

 

GALLEGHER, 1993

Locality: Inversand Pit, Sewell, Gloucester County, New Jersey.

Horizon: MFL, Basal Hornerstown Formation, Greensand, Rancocas Group.

Biostratigraphy:

Age: Late Maastrichtian Stage, uppermost Senonian subepoch, Gulf Epoch, Late Cretaceous.

Material:

NJSM 11332: Teeth.

 

HAWKINS, 1990

Locality: Near Fort Braggs, Alabama.

Horizon: Prairie Bluff Chalk.

Biostratigraphy:

Age: Late Maastrichtian Stage, uppermost Senonian subepoch, Gulf Epoch, Late Cretaceous.

Material:

Number: Not given: Fragmentary skull and fragmentary skeleton.

 

= Mosasaurus maximus (?) MULDER, 1999

Locality: ?Sewell, Gloucester County, New Jersey.

Horizon:

Biostratigraphy:

Age: Maastrichtian Stage, uppermost Senonian subepoch, Gulf Epoch, Late Cretaceous.

Material:

AMNH 14815: ‘Metapodial”, metacarpal 1.

Referred material:

RUSSELL, 1967

Locality: Monmouth County, New Jersey.

Horizon:

Biostratigraphy:

Age: Maastrichtian Stage, uppermost Senonian subepoch, Gulf Epoch, Late Cretaceous.

Material:

ANSP 8481: Tooth.

 


Locality: Woodbury, Gloucester County, New Jersey.

Horizon:

Biostratigraphy:

Age: Maastrichtian Stage, uppermost Senonian subepoch, Gulf Epoch, Late Cretaceous.

Mateiral:

ANSP 8507: Tooth.

 


Locality: Lumberton, New Jersey.

Horizon:

Biostratigraphy:

Age: Maastrichtian Stage, uppermost Senonian subepoch, Gulf Epoch, Late Cretaceous.

Material:

ANSP 8515: Tooth.

 


Locality: Burlington County, New Jersey.

Horizon:

Biostratigraphy:

Age: Maastrichtian Stage, uppermost Senonian subepoch, Gulf Epoch, Late Cretaceous.

Material:

ANSP 8529 (MULDER, 1999): Pygal vertebra.


Locality:

Horizon:

Biostratigraphy:

Age:

Material:

AMNH ?1391:

AMNH ?1393:

AMNH ?1397:

AMNH 1398:

AMNH 1404:

AMNH 1406:

YPM 305:

YPM 306:

YPM 307:

YPM 311:

YPM ?414:

YPM 470:

YPM 508:

YPM 509:

YPM 510:

YPM 690:

YPM 1504: Fragmentary parietal.

YPM 1527:

YPM 1529:

YPM 1530:

YPM 1574:

YPM 1575:

YPM 1582: Fragmentary parietal.

YPM 24933: Fragmentary skull.

ANSP 8462:

ANSP 8472:

ANSP ?8473:

ANSP ?8474:

ANSP 8476:

ANSP 8478:

ANSP 8479:

ANSP 8482:

ANSP 8503:

ANSP 8585:

ANSP 8508:

ANSP 8513:

ANSP 8514:

ANSP 8516:

ANSP 8517:

ANSP 8519:

ANSP 8520:

ANSP 8524:

ANSP 8531:

ANSP 8540:

ANSP 8546:

ANSP 8547:

ANSP 9641:

CNHM PR 491:

TMM: Specimens.

BELL, 1997

TMM 313-1:

YPM 430:

 

GILMORE, 1926

Locality: McNairy County, Tennessee.

Horizon: Coon Creek Formation.

Biostratigraphy:

Age: Late Cretaceous.

Material:

USNM 10540: Cornoid.

 

= Mosasaurus MORTON, 1834 (partim)

Locality:

Horizon:

Biostratigraphy:

Age:

Material:

Number: Not given:

 

= Mosasaurus mitchilli LEIDY, 1865 (partim)
Etymology:

Locality:

Horizon:

Biostratigraphy:

Age:

Material:

Number: Not given:

 

= Mosasaurus princeps MARSH, 1869
Etymology:

Holotype: YPM 430

Locality: Hornerstown, Monmouth County, New Jersey.

Horizon: Hornerstown Formation, Rancocas Group.

Biostratigraphy:

Age: Late Maastrichtian Stage, uppermost Senonian subepoch, Gulf Epoch, Late Cretaceous.

Material: Fragmentary skull and fragmentary skeleton.

 

= Mosasaurus fulciatus COPE, 1869-1870
Etymology:

Holotype: AMNH 1398

Locality: Monmouth County, New Jersey.

Horizon:

Biostratigraphy:

Age: Late Cretaceous.

Material: Quadrate, 3 dorsal vertebrae and an imperfect humerus.

 

= Mosasaurus oarthrus COPE, 1869-1870
Etymology:

Holotype: AMNH 1392

Locality: Barnsboro, Gloucester County, New Jersey.

Horizon: Hornerstown Formation, Rancocas Group.

Biostratigraphy:

Age: Late Maastrichtian Stage, uppermost Senonian subepoch, Gulf Epoch, Late Cretaceous.

Material: Fragmentary left quadrate.

= Mosasaurus dekayi BRONN, 1838
Etymology:

Holotype:

Locality: From the foot of Nevesink Hills, Sandy Hook, Monmouth County, New Jersey.

Horizon: Navesink Formation, Monmouth Group.

Biostratigraphy:

Age: Middle Maastrichtian Stage, uppermost Senonian subepoch, Gulf Epoch, Late Cretaceous.

Material: Tooth Crown.

After GIBBS, 1849.

Referred material:

BAIRD, 1986

MAPS A129a, Johnson coll.

 

Locality: Loc. C, Prince George’s County, Maryland.

Horizon: Severn Formation.

Biostratigraphy:

Age: Early Late Maastrichtian Stage, uppermost Senonian subepoch, Gulf Epoch, Late Cretaceous.

Material: Tooth crown.

 

BELL, MARTIN, PARRIS & GRANDSTAFF, 1994

Locality: Missouri River, South Dakota.

Horizon: upper DeGrey Member of the Pierre Shale.

Biostratigraphy: Baculies compressus zone.

Age: Upper Campanian Stage, Upper Senonian Subepoch, Upper Gulf Epoch, Late Cretaceous.

Material:

Number: Not given: Partial skull.

 

HOGANSON, ERICKSON & GITMAN, 1994, HOGANSON & ERICKSON, 2004

Locality: Missouri River valley, North Dokata.

Horizon: Fox Hills Formation.
Note: Interfingering of Western Interior Seaway with the Hell Creek Formation.

Biostratigraphy:

Age: Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.

Material:

Number: Not given: Teeth.

 

HOGANSON, ERICKSON & HOLLAND, 2007

Locality: UND locality RF132-79-1, Emmons County, North Dakota.

Horizon: Fox Hills Formation.
Note: Interfingering of Western Interior Seaway with the Hell Creek Formation.

Biostratigraphy:

Age: Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.

Material:

UND-PC 15895: Tooth.


LYDEKKER, 1888

Locality: New Jersey.

Horizon:

Biostratigraphy:

Age: Late Cretaceous.

Material:

NHMUK (old BMNH) 11590: Cast of a tooth.
Note: Illustrated by Leidy, 1865, pl. 10, fig. 3.

 

MULDER, 1999

Locality: Greensand, Holmdel, Monmouth County, New Jersey.

Horizon:

Biostratigraphy:

Age: Maastrichtian Stage, uppermost Senonian subepoch, Gulf Epoch, Late Cretaceous.

Material:

AMNH 1397: Left and right quadrate.

AMNH 1388: Several dorsal and caudal vertebrae.

AMNH 2393: Several dorsal and caudal vertebrae.

 

RUSSELL, 1967

Locality:

Horizon:

Biostratigraphy:

Age:

Material:

AMNH 1402:

YPM 410:

YPM 930: Tooth and a short series of anterior caudal vertebrae.

YPM 1582:

ANSP 8468:

ANSP 8471:

ANSP 8495:

ANSP 8511:

 

= Ichthyosaurus de BLAINVILLE, 1827

Locality:

Horizon:

Biostratigraphy:

Age:

Material:

Number: Not given:

 

= Mosasaurus DEKAY, 1830

Locality:

Horizon:

Biostratigraphy:

Age:

Material:

Number: Not given:

 


= Mosasaurus major DEKAY, 1842
Etymology:

Holotype:

Locality:

Horizon:

Biostratigraphy:

Age:

Material: Tooth.

 

= Mosasaurus maximiliani PICTET, 1853
Etymology: In honor of Prince Maximilian zu Wied.

Holotype:

Locality:

Horizon:

Biostratigraphy:

Age:

Material:

After GIBBS, 1849.

 

= Mosasaurus meirssii MARSH, 1869
Etymology:

Holotype: YPM 443

Locality: Meirs Farm, Near Hornerstown, Monmouth County, New Jersey.

Horizon: Late New Egypt Formation, (Hornerstown Formation?, Rancocas Group.)

Biostratigraphy:

Age: Late Maastrichtian Stage, uppermost Senonian subepoch, Gulf Epoch, Late Cretaceous.

Material: Tooth.

_____________________________________________________________________________________
Species: missouriensis (HARLAN, 1834) LEIDY, 1858
Etymology:
= Ichthyosaurus missouriensis HARLAN, 1834
= Ictiosaurus missuriensis (HARLAN, 1834) HARLAN, 1834
= Batrachiosaurus missouriensis (HARLAN, 1834) HARLAN, 1839
= Batrachotherium missouriensis (HARLAN, 1834) HARLAN, 1839
= Mosasaurus maximiliani GOLDFUSS, 1845
Etymology: In honor of Prince Maximilian zu Wied.
= Mosasaurus nevidii MEYER, 1845
= Mosasaurus missuriensis COPE, 1869 (sic)
= Pterycollosaurus maximiliani DOLLO, 1882
(All based on the holotype)

Holotype: RFWUIP 1327, MNHN 9587

Locality: Big Bend along the Missouri River, Between Fort Lookout and Fort Pierre, southeast of Pierre, South Dakota.

Horizon: DeGrey Formation (Upper Pierre Formation), Pierre Shale Group.

Biostratigraphy:

Age: Upper Campanian Stage, Upper Senonian Subepoch, Upper Gulf Epoch, Late Cretaceous.

Material: Skull, most of the vertebral column and fragments of the pectoral girdle.

Note: MNHN 9587: Is the missing snout tip (CALDWELL & BELL, 2005)

Dorsal view of Mosasaurus missouriensis (after GOLDFUSS, 1845 and HARLAN, 1835) from CAMP, 1942.

 

Referred material:

USNM 4910:

USNM 8096:

BELL, 1997

FWU GLODFUSS 1327:

 


Locality: NW 1/4, NW 1/4 Sec. 33, T19N, R21E, Feigns County, 35 miles N of Lewistown, Montana.

Horizon: Bearpaw Formation.

Biostratigraphy:

Age: Middle Campanian Stage, Upper Senonian Subepoch, Upper Gulf Epoch, Late Cretaceous.

Material:

USNM 8086:

 

Locality: South Dakota.

Horizon: Upper Pierre Formation.

Biostratigraphy:

Age: Late Cretaceous.

Material:

USNM 4910:

 

Locality: South Dakota.

Horizon: Sharon Springs Member, Pierre Shale.

Biostratigraphy:

Age: Campanian Stage, Senonian subepoch, Upper Gulf epoch, Late Cretaceous.

Material:

As per NICHOLLS, 1988

AMNH 5815 (mixed in with plesiosaur of same number)

SDSM PC 8021-B (field number), SDSM J.A. N. 7772 (field number), SDSM uncatalogued, UCM 20565, 38087, 48353, UCMP V5407/61221, V76181/45208:

 

KONISHI, NEWBREY & CALDWELL, 2014

Locality: Near Welling, Cardston County, southern Alberta Province, Canada.

Horizon: Bearpaw Formation.

Biostratigraphy:

Age: Campanian Stage, Senonian subepoch, Upper Gulf epoch, Late Cretaceous.

Material:

RTMP 2008.036.0001, 2003.400.0001: Skull and nearly complete skeleton. Note with stomach contents (Aulopiformes (RTMP 2008.036.0006), Lamniformes (RTMP 2008.36.0003), Squalicorax sp (RTMP 2008.036.005)).

 

MARTIN, 2016

Locality: Big Bend along the Missouri River, southeast of Pierre, South Dakota.

Horizon: Lower DeGrey Formation (Upper Pierre Formation), Pierre Shale Group.

Biostratigraphy:

Age: Upper Campanian Stage, Upper Senonian Subepoch, Upper Gulf Epoch, Late Cretaceous.

Material:

Number: Not given: Skull.

Note: A tooth of Notidanodon sp was found in it.

 

THURMOND, 1969

Locality: 1 1/2 miles NE of Riesel, McLennan County, Texas.

Horizon: Probably Wolfe City Member.

Biostratigraphy:

Age: Campanian Stage, Senonian subepoch, Upper Gulf epoch, Late Cretaceous.

Material:

UTA (TAM) uncatalogued: Skull and mandible fragments and vertebrae.

 

= Mosasaurus sp., cf. M. missouriensis YAMASHITA, KONISHI, & SATO, 2015

Locality: South of Lethbridge, southern Alberta Province, Canada.

Horizon: Muddy Unit 1, Bearpaw Formation.

Biostratigraphy:

Age: Campanian Stage, Senonian subepoch, Upper Gulf epoch, Late Cretaceous.

Material:

RTMP 2012.010.0001:

= Mosasaurus maximiliani Referred material:

OWEN, 1854

Locality: New Jersey.

Horizon:

Biostratigraphy:

Age: Late Cretaceous.

Material:

ROYAL COLLEGE OF SURGEONS 320: Cast of the basioccipital.

ROYAL COLLEGE OF SURGEONS 321: Anterior caudal vertebra.

ROYAL COLLEGE OF SURGEONS 322: Anterior caudal vertebra.

ROYAL COLLEGE OF SURGEONS 323: A caudal vertebra.

ROYAL COLLEGE OF SURGEONS 324: Posterior caudal vertebra.

ROYAL COLLEGE OF SURGEONS 325: Tooth from the left dentary.

ROYAL COLLEGE OF SURGEONS 326: Crown of a tooth.

 


Locality: Chesapeake Canal, Delaware.

Horizon: Greensand Formation.

Biostratigraphy:

Age: Late Cretaceous.

Material:

ROYAL COLLEGE OF SURGEONS 327: Cast of a tooth.

 

= Mosasaurus horridus WILLISTON, 1895
Etymology:

Holotype: KUVP 1034

Locality: Cheyenne River, Custer County, South Dakota.

Horizon: Bearpaw Formation.

Biostratigraphy:

Age: Middle Campanian Stage, Upper Senonian Subepoch, Upper Gulf Epoch, Late Cretaceous.

Material: Nearly complete skull, 8 vertebrae, portions of the left front paddle.


Mosasaurus horridus skull after WILLISTON (1898); A) dorsal; B) lateral and C) ventral views.

_____________________________________________________________________________________

Species: iguanavus (COPE, 1868) (nomen dubium)
Etymology:
= Clidastes iguanavus COPE, 1868 (nomen dubium)

Holotype: YPM 1601

Locality: A marl pit near Swedesboro, Gloucester County, New Jersey.

Horizon: Marshalltown Formation, Matawan Group.

Biostratigraphy:

Age: Upper Campanian Stage, Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.

Material: A vertebrae.

Referred material:

= Clidastes cf. C. iguanavus ROBB, 2004

Locality: Maple Shade, Burlington County, New Jersey.

Horizon: Merchantville Formation, Matawan Group.

Biostratigraphy:

Age: Coniacian Stage, Lowermost Senonian subepoch, Gulf Epoch, Late Cretaceous.

Material: ANSP 21539: Tooth crown.


_____________________________________________________________________________________

Species: conodon (COPE, 1881) BAIRD & CASE, 1966
Etymology:
= Clidastes conodon COPE, 1881

Holotype: AMNH 1380

Locality: Near Freehold, Monmouth County, New Jersey.

Horizon: Navesink Formation, Monmouth Group.

Biostratigraphy:

Age: Middle Maastrichtian Stage, uppermost Senonian subepoch, Gulf Epoch, Late Cretaceous.

Material: Skull and anterior cervical vertebrae.

Referred material:

BAIRD 1986

Locality: Loc. B, Prince George’s County, Maryland.

Horizon: Severn Formation.

Biostratigraphy:

Age: Early Late Maastrichtian Stage, uppermost Senonian subepoch, Gulf Epoch, Late Cretaceous.

Material:

USNM 33648: Marginal tooth crown.

Hartstein coll: Marginal tooth crown.

USNM 336473: Palatal tooth.

Bohaska coll:

 

BELL & MARTIN, 1995

Locality:

Horizon: DeGrey Member, Pierre Shale.

Biostratigraphy:

Age: Upper Campanian Stage, Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.

Material:

Number: Not given: A tooth perforating the base of the left quadrate.

 

CICIMURRI, BELL & STOFFER, 1999

Locality: Approximately 1/22 mile south of the Sage Creek Primitive Campground, Badlands National Park, Shannon County, South Dakota.

Horizon: Verendrye Member, Pierre Shale.

Biostratigraphy: Baculites cuneatus biozone.

Age: Upper Campanian Stage, Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.

Material:

BADL 99831: Partial left maxilla and 7 dorsal vertebrae.

 

GALLAGHER, 2005

Locality: From the foot of Nevesink Hills, Sandy Hook, Monmouth County, New Jersey.

Horizon: Navesink Formation, Monmouth Group.

Biostratigraphy:

Age: Middle Maastrichtian Stage, uppermost Senonian subepoch, Gulf Epoch, Late Cretaceous.

Material:

Number: Not given: Lower jaw.

IRWIN, 2008

Locality: SE 1/4, Sec. 17, T11S, R26W, near Columbus, Hempstead County, Arkansas.

Horizon: Upper Marlbrook Marl Formation.

Biostratigraphy:

Age: Late Campanian Stage, upper Senonian subepoch, Gulf Epoch, Late Cretaceous.

Material:

USNM 11396: 35 posterior vertebrae and a nearly complete pelvis.

 

IKEJIRI & LUCAS, 2004

Locality: Trinidad, Las Animias County, Colorado.

Horizon: Upper part of Pierre Shale.

Biostratigraphy: Baculites cuneatus or B. compressus ammonite zone.

Age: Upper Campanian-Lower Maastrichtian Stage, Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.

Material:

TSJC (Trinidad Sate Junior College) 1998.2: Partial skeleton.

 

Locality: U.S. Fish and Wildlife Lands, Phillips County, Northern Montana.

Horizon: Bearpaw Shale.

Biostratigraphy:

Age: Upper Campanian Stage, Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.

Material:

MOR 006: Skull and dentary.

Mosasaurus conodon after IKEJIRI & LUCAS, 2004; A) dorsal and B) lateral views.

Mosasaurus conodon modifed from IKEJIRI & LUCAS, 2004; A) dorsal and B) lateral views.

 

RUSSELL, 1967

Locality:

Horizon:

Biostratigraphy:

Age:

Material:

AMNH 1387:

AMNH ?1395:

AMNH 1397:

YPM ?379:

YPM ?1500:

YPM 1510:

YPM 1573:

ANSP 8469:

ANSP 8480:

ANSP 8501:

ANSP 8502:

ANSP ?8504:

ANSP ?8509:

USNM 11904:

?Taylor Marl, Texas; Marlbrook Marl, Arkansas; Verendrye Member, Lower Pierre Formation, Virgin Creek Member, Upper Pierre Formation, South Dakota; Navesink and younger Cretaceous New Jersey.

 

Locality: E center section 28, T6N, R80W, Huges County, South Dakota.

Horizon: Middle Pierre Formation.

Biostratigraphy:

Age: Late Cretaceous.

Material:

USNM 18255:

 

RUSSELL, 1967

Locality: Near Columbus, Arkansas.

Horizon: Malbrook Marl.

Biostratigraphy:

Age: Late Cretaceous.

Material:

Number: Not given:

_____________________________________________________________________________________

Species: ? lundgreni (SCHRODER, [1884] 1885) KUHN, 1939
Etymology:
= Leiodon lundgreni SCHRODER, (1884) 1885
= Liodon lundgreni (SCHRODER, [1884] 1885) LYDEKKER, 1888

Holotype:

Locality: Sweden.

Horizon:

Biostratigraphy:

Age: Upper Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.

Material: Teeth.
_____________________________________________________________________________________

Species: scanicus (NILSSON, 1835) SCHRODER, (1884) 1885
Etymology:
= Plesiosaurus scanicus NILSSON, 1835

Holotype:

Locality: Sweden.

Horizon:

Biostratigraphy:

Age: Upper Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.

Material: Skull fragment.
_____________________________________________________________________________________

Species: lemonnieri DOLLO, 1889
Etymology:

Holotype: IRSNB R 28 (formerly 1470)

Locality: Mesvin, 4 kil. from Mons, Belgium.

Horizon: Craie Burne phosphate de Ciply.

Biostratigraphy:

Age: Middle Maastrichtian Stage, uppermost Senonian subepoch, Gulf Epoch, Late Cretaceous.

Material: Fragmentary skull and fragmentary skeleton.

Skull after LINGHAM-SOLIAR, 2000. A) Dorsal and B) Lateral views.

Referred material:

IRSNB 3119: 45 prepygals, 13 pygals, 25 intercaudals and 49-50 terminals.

IRSNB 3210: 38 prepygals, 22 pygals, 15 intercaudals and 32 terminals.

IRSNB 1610, 1629, 3078, 3105, 3110, 3112, 3113, 3114, 3116, 3117, 3118, 3120, 3125, 3127, 3128, 3129, 3131, 3131, 3132, 3133, 3150, 3151, 3153, 3156, 3157, 3169, 3186, 3187, 3189, 3193, 3857, 3858, 3860, 3861, 3902, 3911, 4583, 4698:

IRScNB 3211: Skull and dentary.

BAIRD & CASE, 1966

Locality: Spoil heaps piled up by a bulldozer excavating for an irrigation pond in north-central Burlington County, this pond lies on the Emil Novak farm on the east side of the Eayrestown Road, 1.2 miles southeast of Lumberton and 2.2 miles northwest of Vincetown, Burlington County, New Jersey.

Horizon: Navesink Formation, Monmouth Group.

Biostratigraphy:

Age: Middle Maastrichtian Stage, uppermost Senonian subepoch, Gulf Epoch, Late Cretaceous.

Material:

PU 18826: Thoracic vertebra.

 

BARDET, 2012

Locality: Ciply, Mons Basin, Belgium.

Horizon: Phosphatic Chalk.

Biostratigraphy:

Age: Maastrichtian Stage, Upper Senonian subepoch, Upper Upper Gulf Epoch, Late Cretaceous.

Material:

MNHN 1898-5: Skull bones.

 

BARDET, CAPPETTA, PEREDA SUBERBIOLA, MOUTY, AL MALEH, AHMAD, KHRATA & GANNOUM, 2000

Locality: Bardeh mine, Palmyra area, Palmyrides China of Syria.

Horizon: Phosphatic deposit.

Biostratigraphy:

Age: Lower Maastrichtian Stage, Upper Senonian subepoch, Upper Upper Gulf Epoch, Late Cretaceous.

Material:

Number: Not given: Teeth.

BARDET, 2012

MNHN 1938-2: Broken dentary.

Locality: Mine A, Charquieh, Palmyra area, Palmyrides China of Syria.

Horizon: Phosphatic deposit.

Biostratigraphy:

Age: Late Cretaceous.

Material:

Number: Not given: Teeth.

BARDET, 2012

MNHN-BAR1: Teeth.

 

Locality: Khneifiss mine, Palmyra area, Palmyrides China of Syria.

Horizon: Phosphatic deposit.

Biostratigraphy:

Age: Late Cretaceous.

Material:

Number: Not given: Teeth.

 

LINGHAM-SOLIAR, 1991

Locality: Belgium.

Horizon:

Biostratigraphy:

Age: Late Cretaceous.

Material:

IRSNB 3201: Skull and fragmentary skeleton of an immature specimen.

 

= Mosasaurus cf. lemonnieri LINGHAM-SOLIAR, 1994

Locality: Manzadi, Bas Congo, Democratic Republic of the Congo (Formerly the Belgian Congo, Zaire).

Horizon:

Biostratigraphy:

Age: Maastrichtian Stage, uppermost Senonian subepoch, Gulf Epoch, Late Cretaceous.

Material:

Number: Not given: 4 teeth probably from different individuals and different parts of the dental ramus.

 

= Mosasaurus cf. lemonnieri MACHALSKI, JAGT, DORTANGS, MULCER & RADWANSKI, 2003

Locality: Opoka at Nasilow, Poland.

Horizon:

Biostratigraphy:

Age: Upper Upper Maastrichtian Stage, uppermost Senonian subepoch, Gulf Epoch, Late Cretaceous.

Material:

ZPAL R. 9/1, R. 9/2, R. 9/5: 3 tooth crowns.

 

MEIJER, 1984

Locality: Limburg, Ciply, Hainault Province, Belgium.

Horizon:

Biostratigraphy:

Age: Maastrichtian Stage, Senonian Subepoch, Gulf Epoch, Late Cretaceous.

Material:

Numbers: Not given (Haarlem, Leiden and Maastricht Museum): Teeth.

 

OWOCKI & MADZIA, 2020

Locality: 'Craie (brune) phosphate de Ciply' of the Mons Basin, Southern Belgium.

Horizon:

Biostratigraphy: Belemnella obtusa Zone.

Age: Early Maastrichtian Stage, Senonian Subepoch, Gulf Epoch, Late Cretaceous.

Material:

IRSNB R 400: Tooth crown.

_____________________________________________________________________________________

Species: hardenponti DOLLO,
Etymology:

Holotype:

Locality:

Horizon:

Biostratigraphy:

Age:

Material:
_____________________________________________________________________________________

Species: lonzeensis DOLLO, 1904
Etymology:

Holotype:

Locality: Lonzee, near Gembloux, Namur Province, Belgium.

Horizon:

Biostratigraphy:

Age: Lower Senonian Subepoch, Gulf Epoch, Late Cretaceous.

Material: Quadrate, and caudal vertebrae.
_____________________________________________________________________________________

Species: johnsoni (MEHL, 1930)
Etymology:
= Amphekepubis johnsoni MEHL, 1930

Holotype: University of Missouri No. 509VP

Locality: Approximately 40 miles east and a little north of Monterey, Nuevo Leon State, Mexico.

Horizon: San Felipe Formation.

Biostratigraphy:

Age: Late Cretaceous.

Material: Sacral vertebra, pubis, ischium, ilium, left femur, left tibia, left first metatarsal, and 4th metatarsal.
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Species: beaugei ARMBOURG, 1952
Etymology:

Holotype: MNHN PMC 7

Locality: Unknown, Oulad Abdoun Basin, Morocco.

Horizon: Couche III

Biostratigraphy:

Age: Late Maastrichtian Stage, uppermost Senonian subepoch, Gulf Epoch, Late Cretaceous.

Material: Isolated tooth.

Referred material:

MNHN PMC 8-9: 2 isolated teeth.

BARDET, SUBERBIOLA, IAROCHENE, BOUYAHYAOUI, BOUYA & AMAGHZAZ, 2004

Locality: Andre Delpit Locality, Morocco.

Horizon: Couche III

Biostratigraphy:

Age: Late Maastrichtian Stage, uppermost Senonian subepoch, Gulf Epoch, Late Cretaceous.

Material:

MNHN PMC 10: Isolated tooth.

 

Locality: Bou Jniba Locality, Morocco.

Horizon: Couche III

Biostratigraphy:

Age: Late Maastrichtian Stage, uppermost Senonian subepoch, Gulf Epoch, Late Cretaceous.

Material:

MNHN PMC 11-13: 3 isolated teeth.

 

Locality: Grand Daoui area, Morocco.

Horizon: Sidi Daoui Zone.

Biostratigraphy:

Age: Late Maastrichtian Stage, uppermost Senonian subepoch, Gulf Epoch, Late Cretaceous.

Material:

OCP-DEK/GE 83: Incomplete skull and lower jaw.

OCP-DEK/GE 303: Incomplete skull and lower jaw.

 

 

CHURCHER, 1988

Locality: Dakhleh Oasis, Western Desert of Egypt.

Horizon: Duwi Formation.

Biostratigraphy:

Age: Campanian Stage, Senonian subepoch, Upper Gulf epoch, Late Cretaceous.

Material:

Number: Not given.:

 

= cf beaugei PRICE 1957
= Mosasaurus cf. beugei BONAPARTE, 1978 (sic)

Locality: South America.

Horizon:

Biostratigraphy:

Age: Late Cretaceous.

Material:

Number: Not given:Teeth.

 

= Mosasaurus cf. beaugei REMPERT, MARTENS & MELCHERS, 2024b

Locality: Holden Beach, Brunswick County, North Carolina.

Horizon: Peedee Formation.

Biostratigraphy:

Age: Upper Maastrichtian Stage, Late Senonian Subepoch, Late Gulf Epoch, Latest Cretaceous.

Material:

CW.001: A rooted marginal tooth.

SS.001, VANPS 13.0185: Marginal tooth crowns.

 


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Species: poultneyi MARTIN, 1953 (nomen nudum)
Etymology:

Holotype: SDSM 452

Locality: At the spillway of Elm Lake, 10 miles west of Fredrick, Me 1/4, Sec. 6, T 127 N, R 64 W, Brown County, South Dakota.

Horizon: Lower Virgin Creek member, Pierre Formation.

Biostratigraphy:

Age: Lower Maastrichtian Stage, uppermost Senonian subepoch, Gulf Epoch, Late Cretaceous.

Material: Fragmentary skull, jaws and skeleton.
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Species: mokoroa WELLES & GREGG, 1971
Etymology:

Holotype: CM zfr 1

Locality: Exact locality unknown, Haumuri Bluff, Cheviot and Waipara District, South Island, New Zealand.

Horizon: Lower I. korojonense Zone.

Biostratigraphy:

Age: Haumurian, Maastrichtian Stage, uppermost Senonian subepoch, Gulf Epoch, Late Cretaceous.

Material: Skull and vertebrae.

Mosasaurus mokoroa skull in dorsal and lateral view modified from Welles & Gregg, 1971.

Referred material:

= Taniwhasaurus oweni HECTOR, 1874

DM R1535: Paddle.

 

= Mosasaurus minor GIBBS, 1881
Etymology:

Holotype:

Locality: Alabama.

Horizon:

Biostratigraphy:

Age:

Late Cretaceous.

Material: Fragmentary skeleton.

After GIBBS, 1849.

 

= Leiodon haumuriensis HECTOR; HUTTON, 1895
Etymology:

Number:

Locality:

Horizon:

Biostratigraphy:

Age:

Material:

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Species: hobetsuensis SUZUKI, 1985
Etymology:

Holotype: HMG 12

Locality: Pankerusanosawa Creek, Hobetsu-cho, central Hokkaido Island, Japan.

Horizon: Hakobuchi Group.

Biostratigraphy:

Age: Lower Maastrichtian Stage, uppermost Senonian subepoch, Gulf Epoch, Late Cretaceous.

Material:

HMG 12-1: Marginal tooth.

HMG 12-2 thru 12-11: Dorsal vertebrae.

HMG 12-12 thru 12-21: Thoracic dorsal ribs.

HMG 12-22 thru 12-25: Right lumbar ribs.

HMG 12-26 thru 12-33: Left thoracic dorsal ribs.

HMG 12-34: Interclavicle.

HMG 12-35: Right clavicle?

HMG 12-36: Right scapula.

HMG 12-37: Right coracoid.

HMG 12-38: Right humerus.

HMG 12-39: Right radius.

HMG 12-40: Right ulna.

HMG 12-41: Right radiale.

HMG 12-42: Right intermedium.

HMG 12-43: Right ulnae.

HMG 12-44: Left ulnae.

HMG 12-45: Right distal carpal 2.

HMG 12-46: Right distal carpal 3.

HMG 12-47: Right distal carpal 4.

HMG 12-48: Right metacarpal 1.

HMG 12-49: Left metacarpal 1.

HMG 12-50: Right metacarpal 2.

HMG 12-51: Right metacarpal 3.

HMG 12-52: Right metacarpal 4.

HMG 12-53: Right metacarpal 5.

HMG 12-54: Right fore first phalange of first digit.

HMG 12-55: Left fore first phalange of first digit.

HMG 12-56: Right fore second phalange of first digit.

HMG 12-57: Left fore second phalange of first digit.

HMG 12-58: Right fore third phalange of first digit.

HMG 12-59: Left fore third phalange of first digit.

HMG 12-60: Right fore first phalange of second digit.

HMG 12-61: Right fore second phalange of second digit.

HMG 12-62: Left fore third phalange of second digit.

HMG 12-63: Right fore first phalange of third digit.

HMG 12-64: Left fore third phalange of third digit.

HMG 12-65: Right fore first phalange of fourth digit.

HMG 12-66: Right fore second phalange of fourth digit.

HMG 12-67: Right fore first phalange of fifth digit.
_____________________________________________________________________________________

Species: prismaticus SAKUAI, CHITOKU & SHIBUYA, 1999
Etymology:

Holotype: HMB-1056

Locality: Hobetsu River, 42°49’50”N, 142°9’31”E, Hobetsu, near Penkeobiraruka-sawa, Hobetus Disctrict of Mukawa Town, Hokkaido Island, Japan.

Horizon: Hakobuchi Group.

Biostratigraphy:

Age: Upper Campanian-Early Maastrachtian Stage, uppermost Senonian subepoch, Gulf Epoch, Late Cretaceous.

Material: Parietal, occipital unit, epipterygoid? and a marginal tooth.

Referred material:

= Mosasaurus cf. prismaticus TANIMOTO, SHINZYO & FUJIMOTO, 2001

Locality: Takinoike, Izumisano City, Osaka Prefecture, Honshu Island, Japan.

Horizon: Izumi Group.

Biostratigraphy:

Age: Maastrichtian Stage, Uppermost Senonian Subepoch, Gulf Epoch, Late Cretaceous.

Material:

Number: Not given: Tooth.

= Kourisodon sp TANIMOTO & KANAZAWA 2001, TANIMOTO, KANAZAWA & SATO, 2004, TANIMOTO, 2008 (as per KONISHI, TANIMOTO, CALDWELL & KANZAWA, 2010)

Locality: Kanewari, Sanuki City, Kagawa Prefecture, Shikoku Island, Southwest Japan.

Horizon: Hiketa Formation, Izumi Group.

Biostratigraphy:

Age: Late Campanian Stage, Senonian Subepoch, Gulf Epoch, Late Cretaceous.

Material:

KSNHM-F6-2: Fragmentary skull and fragmentary dentary.