Species: marshi DOLLO, 1882
Etymology: In honor of Othneil Marsh.

Holotype: IRSNB R38

Locality: Near Zichen, Limburg Province, The Netherlands.

Horizon: Maastricht Formation.

Biostratigraphy:

Age: Upper Maastrichtian Stage, uppermost Senonian subepoch, Gulf Epoch, Late Cretaceous.

Material: Fragmentary skull, and postcranial remains including vertebrae, a portion of an interclavicle, ribs, left scapula, abraded humerus, radius and ulna and a single phalanx.

Skull based on IRSNB R38, IRSNB 1622 and NHMUK R5868 (Modified from Lingham-Soliar, 1992).

Skull after LINGHAM-SOLIAR, 1994.

Referred material:

(Listed in Lingham-Solair, 1992)

IRNSB R40, 1622, 1675, NHMM 198349, 1989104, 1989126, 1423, 003900:

IRNSB 3673: A series of the posterior portion of the vertebral column

NHMM 1989124: Well preserved radius.

TSMNH 11298: Scapula.

TSMNH 11291: Corocoid.

TSMNH 11276: Corocoid.

TSMNH 11209: Articulated series of 6 posterior caudal vertebrae.

TSMNH 3582, TSMNH 11227: Fused “sacra”.

MULDER, 2001

IRScNB 1497: Co-ossified vertebrae, pathology.

MULDER, 2003

TM 11226, 11318, 17246: Anterior dorsal vertebrae.

TM 11331: Fragmentary posterior cervical vertebra.

 

Locality: Near EbenEmael, Liege Province, Belgium.

Horizon: Upper Gulpen Formation.

Biostratigraphy:

Age: Upper Maastrichtian Stage, uppermost Senonian subepoch, Gulf Epoch, Late Cretaceous.

Material:

IRNSB R39 (old 1496): Well preserved scapula and moderately well preserved coracoid.

MULDER, 2001

Co-ossified vertebrae, pathology.

BARDET, JAGT, KUYPERS & DORTANGS, 1998

Locality: Romontbos quarry, Eben Emael, Liege Province, Belgium.

Horizon:

Biostratigraphy:

Age: Upper Maastrichtian Stage, uppermost Senonian subepoch, Gulf Epoch, Late Cretaceous.

Material:

Dortangs Collection, no. RD 172: Caudal vertebra.
Note: Showing scavenging by the common dogfish (Squalid) shark, Centrophoroides appendiculatus.

 

JAGT, DECKERS, LEEBEECK, DONOVAN & NIEUWENHUIS, 2020

Locality: ENCI-Heidelbertg Cement Group Quarry, south of Masstricht, Limburg Province, The Netherlands.

Horizon: Base of the Gronsveld Member, Maaastricht Formation.

Biostratigraphy:

Age: Late Maastrichtian Stage, uppermost Senonian subepoch, Gulf Epoch, Late Cretaceous.

Material:

NHMM JJ 11964: Fragmentary cervical vertebra.

Note: With bioerosian by 2 indeterminate osytsers, 2 cheilostome bryozon colonies.

 

Locality: ENCI-Heidelbertg Cement Group Quarry, south of Masstricht, Limburg Province, The Netherlands.

Horizon: Base of the Emael Member, Maastricht Formation.

Biostratigraphy:

Age: Late Maastrichtian Stage, uppermost Senonian subepoch, Gulf Epoch, Late Cretaceous.

Material:

NHMM 2012 073: A set of associted bones of teh skull adn post-cranial skeleton.

Note: WIth traces of bioerosian.

 

LYDEKKER, 1888

Locality: Maastricht, Limburg Province, The Netherlands.

Horizon:

Biostratigraphy:

Age: Maastrichtian Stage, Upper Senonian Subepoch, Upper Gulf Epoch, Late Cretaceous.

Material:

BMNH 44822: Imperfect cervical vertebrae.

MULDER, 2003

NMH 42900: Imperfect cervical vertebrae.
Note: These vertebrae were refered to Halisaurus platyspondylus by LINGHAM-SOLIAR, 1996 but refuted by MULDER, 2003.

 

MULDER, 2001

Locality: Ankerpoort-‘t Rooth Quarry, Bemelen, The Netherlands.

Horizon: ?Nekum Member, Maastricht Formation.

Biostratigraphy:

Age: Upper Maastrichtian Stage, uppermost Senonian subepoch, Gulf Epoch, Late Cretaceous.

Material:

MND 20.01.826: Dorsal vertebra.
Note: with infectious reactive new bone formation.

MND20.01.842: Pygal vertebrae.
Note: fused by ligamentous ossification.

 

Locality: Maastricht, Limburg Province, The Netherlands.

Horizon:

Biostratigraphy:

Age: Upper Maastrichtian Stage, uppermost Senonian subepoch, Gulf Epoch, Late Cretaceous.

Material:

MNHNP AC 9649-9775: Substantial portion of a vertebral column.
Note: fused by infectious spondylitis.

_____________________________________________________________________________________
Species: depressus (COPE, 1869)
Etymology:
= Mosasaurus gracilis COPE, 1869 non OWEN, 1951
= Mosasaurus depressus COPE, 1869 (nomen nudum)
= Mosasaurus depressus COPE, 1869

Holotype:

Locality: Birmingham (Pemberton), Burlington County, New Jersey.

Horizon: Navesink Formation, Monmouth Group.

Biostratigraphy:

Age: Middle Maastrichtian Stage, uppermost Senonian subepoch, Gulf Epoch, Late Cretaceous.

Material: Quadrate cranial fragments and dorsal vertebra.
Note: Missing.

Referred material:

AMNH 1382:

AMNH 1430:

AMNH 2304:

 

MULDER, 1999

Locality: Monmouth County, New Jersey.

Horizon:

Biostratigraphy:

Age: Maastrichtian Stage, uppermost Senonian subepoch, Gulf Epoch, Late Cretaceous.

Material:

ANSP 8535: Cervical vertebra.
Note: Figured by Leidy, 1865, pl 7. figs, 2, 3.

Locality: Inversand Company Pit, Sewell, Gloucester County, New Jersey.

Horizon: Navesink (= New Egypt) Formation, Monmouth Group.

Biostratigraphy:

Age: Middle Maastrichtian Stage, uppermost Senonian subepoch, Gulf Epoch, Late Cretaceous.

Material:

ANSP 20184: Two casts of tooth.

ANSP 20815: Cast of tooth.

 

= Mosasaurus copeanus MARSH, 1869
Etymology:

Holotype: YPM 312

Locality: Marlborough, Monmouth County, New Jersey.

Horizon: Navesink Formation, Monmouth Group.

Biostratigraphy:

Age: Middle Maastrichtian Stage, uppermost Senonian subepoch, Gulf Epoch, Late Cretaceous.

Material: Fragmentary skull.

 

= Halisaurus fraternus MARSH, 1869
Etymology:

Holotype: YPM 445

Locality:

Horizon:

Biostratigraphy:

Age:

Material:

 

= Liodon laticaudus MARSH, 1870
Etymology:
= Tylosaurus laticaudus (MARSH, 1870) HAY, 1902

Holotype: YPM ?411

Locality: Hornerstown, Monmouth County, New Jersey.

Horizon: Hornerstown Formation, Rancocas Group.

Biostratigraphy:

Age: Late Maastrichtian Stage, uppermost Senonian subepoch, Gulf Epoch, Late Cretaceous.

Material:

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Species: primaevus RUSSEL, 1967 (DUNKLE, in CRANDELL, 1958)
Etymology:
= Platecarpus cf. brachycephalus DUNKLE, in CRANDELL, 1958
Etymology:

Holotype: USNM 18254

Locality: NE Corner Sec. 35, T112N, R80W, near Pierre, Hughes County, South Dakota.

Horizon: DeGrey Member, Lower Pierre Shale.

Biostratigraphy:

Age: Late Cretaceous.

Material: Quadrate, 3 anterior dorsal vertebrae and both scapulae-coracoid.

Referred material:

BELL, SHELDON, LAMB & MARTIN, 1996

Locality: South Dakota.

Horizon: DeGrey Member, Pierre Shale.

Biostratigraphy:

Age: Late Cretaceous.

Material:

Number: Not given: Fragmentary skeleton with 4 fragmentary skeletons of babies in it’s body cavity, approximately 1.2-1.5 m.

 

HOLMES, 1996

Locality: South Saskatchewan locality, Alberta Province, Canada.

Horizon: Bearpaw Formation.

Biostratigraphy:

Age: Middle Campanian Stage, middle Senonian subepoch, upper Gulf Epoch, Late Cretaceous.

NMC 11835: Associated remains comprising most of skull including partial braincase, quadrates and mandible fragments, complete presaral column and pygal series, many rib fragments, but few complete elements, endochondral pectoral girdle, humeri, epipodials, and Incomplete, scattered carpals, metacarpals, and phalanges, both pelvic girdles, femora, eppipodials, and incomplete, scattered tarsals, metatarsals, and phalanges.

NMC 11840: Well preserved braincase and much of posterior portion of dematocranium and quadrates, cervical vertebral column, and right scapulocoroid.

P. 628.1: Articulated, complete posterior pygal series and most of tail, rear limbs and girdles.

Plioplatecarpus primaevus frontals from HOLMES, 1996. A) NMC 11835, B) NMC 11840.

Plioplatecarpus primaevus frontals after HOLMES, 1996. A) dorsal and B) lateral views.

 

IRWIN, 2008

Locality: NW 1/4, Sec. 16, T11S, R26W, Hempstead County, Arkansas.

Horizon: Upper Malbrook Marl Formation.

Biostratigraphy:

Age: Upper Campanian Stage, Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.

Material:

SMPSMU 76393: Isolated tooth crown.

 

LINGHAM-SOLIAR, 1992

Locality: Approximately 1.6 km northeast of the intersection of Arkansas Highways 26 and 51, Clark County, Arkansas.

Horizon: Upper Malbrook Marl Formation.

Biostratigraphy:

Age: Upper Campanian Stage, Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.

Material:

BMNH 5868: Quadrate, ribs, vertebrae, a well preserved humerus, an intermedium, and several phalanges.

IRWIN, 2008

FMNH P27457: Axis vertebra.

 

TOKARYK, 1993

Locality: SMNH locality 72J11-0041 is situated between the Snakebite and Cruikshank Coulee localities, approximately 5 kilometers northeast of the village of Main Centre, Saskatchewan Province, Canada.

Horizon: Bearpaw Shale, Late Cretaceous.

Biostratigraphy:

Age: Upper Campanian-Lower Maastrichtian Stage, middle Senonian subepoch, upper Gulf Epoch, Late Cretaceous.

Material:

SMNH P1756.1: Anterior portion of skeleton. Lower jaws, partial premaxilla and maxilla, left humerus, partial left coracoid, left scapula, right coracoid, partial right scapula, axis, cervical vertebrae 3-7, and dorsal vertebrae 8-19.

_____________________________________________________________________________________

Species: houzeaui (DOLLO, 1889) DOLLO, 1904
Etymology:
= Oterognathus houzeaui DOLLO, 1889

Holotype: IRSNB R35 (Old IRSNB 3111)

Locality: Ciply, Hainaut Province, Belgium.

Horizon: Brown Phosphatic Chalk.

Biostratigraphy:

Age: Maastrichtian Stage, uppermost Senonian subepoch, Gulf Epoch, Late Cretaceous.

Material: Fragmentary skull and fragmentary skeleton.

Referred material:

IRNSB 3099, 3103, R 36 (old 3100),3107, 3108, 3155, 3160, 3859, 3154, 3152, 3130, 3122, R37:

 

Skull after LINGHAM-SOLIAR, 1994 (IRSNB 31010, 3130 & R35).

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Species: crassartus (COPE, 1872) LINGHAM-SOLIAR & NOLF, 1990
Etymology:
= Platecarpus crassartus (COPE, 1872)
= Liodon near L. proriger COPE, 1872
= Liodon crassartus COPE, 1872
= Prognathodon crassartus (COPE, 1872) RUSSELL, 1967

Holotype: AMNH 1562

Locality: Near Eagle Tail, Kansas.

Horizon: Lower Pierre Formation.

Biostratigraphy:

Age: Late Cretaceous.

Material: A series of dorsal, lumbar, and caudal vertebrae, with some bones of the limbs.

Referred material:

LINGHAM-SOLIAR & NOLF, 1989

Locality: Arkansas.

Horizon: Marlbrook Marl.

Biostratigraphy:

Age: Upper Campanian Stage, Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.

Material:

BMNH R5868: Humerus, fragment of basioccipital, articular, quadrate, phalanges, tooth, and numerous vertebrae.

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Species: peckensis CUTHBERTSON & HOLMES, 2015
Etymology: In reference to Fort Peck Reservoir, along the shores of which the specimen was discovered.

Holotype: MOR 1062

Locality: MOR locality BS104, along a beach on Fort Peck Reservoir within the Charles M. Russel National Wildlife Refuge, Valley County, northeast Montana.

Horizon: Bearpaw Shale.

Biostratigraphy:

Age: Campanian Stage, Senonian subepoch, Upper Gulf epoch, Late Cretaceous.

Material: Nearly complete, disartculated skull, 26 vertebrae, most of which from an articulated series, and rib fragments.

Frontal from CUTHBERTSON & HOLMES, 2015.

Skull modified from CUTHBERTSON & HOLMES, 2015, A) Dorsal, and B) Lateral views.