Genus: Tylosaurus MARSH, 1872
Etymology Greek, tylos, “knob” and Greek, sauros, “lizard.”
= Hainosaurus DOLLO, 1885
Etymology:
= Liodon COPE, 1869-1870 (partim)
Etymology:
= Rhamphosaurus COPE, 1872 non FITZINGER, 1843 (Ramphosaurus [without the h] bellii, Scincidae, Acontinae)
Etymology: Greek, rhamphos, "beak" and Greek, sauros, "lizard," beak lizard.
= Rhinosaurus MARSH, 1872 non FISCHER von WALDHEIM, 1847 (Rhinosaurus jasikovi, Seymouriamorpha, Seymouriidae)
Etymology: Greek, rhinos, "nose" and Greek, sauros, "lizard," nose lizard.
Species: proriger (COPE, 1869) LEIDY, 1873
Etymology:
= Macrosaurus proriger COPE, 1869
= Macrosaurus pririger COPE, 1869 (sic)
= Liodon proriger (COPE, 1869) COPE, 1869-1870
= Rhamphosaurus proriger (COPE, 1869) COPE, 1872
= Rhinosaurus proriger (COPE, 1869) MARSH, 1872
= Tylosaurus (proriger) proriger (COPE, 1869) MARSH, 1872
Holotype: MCZ
Locality: Vicinity of Monument Rocks, the old overland stage station, Gove
County, Kansas.
Horizon: Smoky Hill Chalk Member, Niobrara Formation.
Biostratigraphy:
Age: Santonian or Campanian Stage, Middle Senonian Subepoch, Uppermiddle Gulf
Epoch, Late Cretaceous.
Material: Anterior portion of skull, cranial fragments, and 13 vertebrae.
Referred material:
Tylosaurus proriger after Williston, 1858; A) dorsal; B) left laterial, and C) ventral views.
AMNH 221:
AMNH 1493:
AMNH 1529:
AMNH 1535:
AMNH 1555:
AMNH 1560:
AMNH 1585:
AMNH 1592: Fragmentary skull.
AMNH 2160:
AMNH 4909 frontal from JIMENEZ-HUIDOBRO, SIMOES & CALDWELL, 2016.
AMNH 4909: Skull.
YPM 1288:
YPM 1302:
YPM 1305:
YPM 3873:
YPM 3977:
YPM 3978:
YPM 3981:
YPM 3984:
YPM 3987:
YPM 3990: Fragmentary skull.
YPM 3993:
YPM 3999:
YPM 4002: Fragmentary skeleton.
24 unnumbered specimens.
ANSP 5:
USNM 3776:
USNM 3794: Extremity of snout.
USNM 3798:
USNM 3897:
USNM 6086:
USNM 17909:
USNM 43: Caudal vertebrae.
USNM 3885: Tooth.
USNM 3891: Fragments of jaw and splenial bone.
MCZ 1030:
KU 1020: Cranial fragments, quadrates, and paddle elements.
KU 1023:
KU 1032:
KU 1033:
KU 1084:
KU 1115:
KU 1135:
KU 1194:
KUVP 1075: Fragmentary skeleton with impressions of dermal scales.
KUVP?: Fragmentary skull and fragmentary skeleton.
KUVP 6634: Mandibular tooth.
KUVP 1195: Fragment of skull and fragmentary skeleton.
KUVP?: Ilium, radius and metacarpal.
KUVP 1017: Pelvic girdles.
KUVP 1029: Ulna and radius.
KUVP 1016: Radius, humerus coracoid and cartilage.
KUVP 1189: Left quadrate, anterior caudal vertebrae.
KUVP 1013: 28 vertebrae.
FHM 4:Ür
CNHM P15144:
CNHM UR902:
CMN (old NMC) 8162: Skull and nearly complete skeleton (wall mount).
PU 12000:
STEWART & MALLON, 2018
CMN 51268: Fused parietals.
CMN 51260: Left prootic.
CMN 51261: Right prootic.
CMNH 51262: Right quadrte.
CMNH 51263: A supraoccipital.
From the Smoky Hill member, Niobrara Formation, Kansas and Colorado; Lower
Pierre Formation, Kansas and South Dakota; Telegraph Creek Formation, Montana.
BELL, 1997
DMNH 8100:
FHM VP3(4):
YPM 1268:
SHELDON, 1997
RMM 1913:
JIMENEZ-HUIDOBRO, SIMOES & CALDWELL, 2016.
RMM 5610: Skull.
RMM 5610 frontal from JIMENEZ-HUIDOBRO, SIMOES & CALDWELL, 2016.
Locality: 2 1/2 miles S of Wallace on Pinnell Ranch, Wallace County, Kansas.
Horizon:
Biostratigraphy:
Age: Late Cretaceous.
Material:
KU 5033:
Locality: SW 1/4, SE 1/4, sec. 36, T29S, R65W near Walsenberg, 1/3 mile NW
boundary between Huerfano and Las Animas Counties, Huerfano County, Colorado.
Horizon: Niobrara Formation.
Biostratigraphy:
Age: Late Cretaceous.
Material:
AMNH 7220:
Locality: South Dakota.
Horizon: Pierre Formation.
Biostratigraphy:
Age: Late Cretaceous.
Material:
CNHM UR820:
ANONYMOUS, 1997
Locality: Kansas.
Horizon: Niobrara Formation.
Biostratigraphy:
Age: Late Cretaceous.
Material:
Fred Nuss and Candace Williams, (fossil collectors): Skull and fragmentary
skeleton of a very large specimen.
BELL & BARNES, 2007
Locality:?
Horizon?
Biostratigraphy:?
Age?
Material:
GSM 1: Fragmentary specimen.
Note: Has fragmentary remains of Clidastes sp in its stomach region.
STERNBERG, 1918, EVERHART, 2004
Locality: South side of (Twin) Butte Creek, 15 miles southeast of Russell
Springs, Logan County, Kansas.
Horizon: Smoky Hill Chalk Member, Niobrara Chalk Formation.
Biostratigraphy:
Age: Late Santonian to Early Campanian Stage, Middle Senonian Subepoch,
Uppermiddle Gulf Epoch, Late Cretaceous.
Material:
USNM 8898: Nearly complete specimen.
Note: With stomach contents of partially
digested polycotylid plesiosaur bones.
Note: Gastroliths were found.
EVERHART & EVERHART,
1997
Locality: Western Kansas.
Horizon: Smoky Hill Chalk Member, Niobrara Formation.
Biostratigraphy:
Age: Santonian or Campanian Stage, Middle Senonian Subepoch, Uppermiddle Gulf
Epoch, Late Cretaceous.
Material:
Number: Not given: Skull and 7 cervical vertebrae.
FINSLEY, 1996
Locality: Rockwall County, Texas.
Horizon: Taylor Group.
Biostratigraphy:
Age: Late Cretaceous.
Material:
Dallas Museum of Natural History: Fragmentary skeleton.
HARRELL JR, & PEREZ-HUERTA, 2015
Locality: West Greene, Greene County, Alabama.
Horizon: Lower unnamed Member, Mooreville Formation.
Biostratigraphy:
Age: Campanian Stage, Senonian subepoch, Gulf Epoch, Late Cretaceous.
Material:
ALMNH PV 1985.0022: Parital skeleton.
LAMBE, 1914
Locality: Gove County, Kansas.
Horizon: Niobrara Formation.
Biostratigraphy:
Age: Late Cretaceous.
Material:
Acc. No. 57: Skull and skeleton.
Locality: Hackberry Creek, near Gove County, Kansas.
Horizon: Niobrara Formation.
Biostratigraphy:
Age: Late Cretaceous.
Material:
NMC 8162: Skull.
LAMBE, 1915
Locality: Beaver Creek, Logan county, Kansas.
Horizon: Niobrara Formation.
Biostratigraphy:
Age: Late Cretaceous.
Material:
NMC 346, 346a: Scapula and coracoid.
MARTIN & BJORK, 1987
Locality: Soske Ranch Locality, SDSM V8120, Fall River County, South Dakota.
Horizon: Bammon Ferruginous Member of the Pierre Shale.
Biostratigraphy:
Age: Campanian Stage, Senonian subepoch, Gulf Epoch, Late Cretaceous.
Material:
SDSM & T (SDSM) 10439: Fragmentary skull and skeleton.
Note: Has stomach contents of Platecarpus tympaniticus, Hesperornis sp,
shark and fish.
MARTIN, SCHUMACHER PARRIS & GRANDSTAFF, 1998
Locality: Along the Missouri River, Yankton county, South Dakota.
Horizon: 4.6 meters below the Niobrara/Pierre Contact.
Biostratigraphy:
Age: Late Cretaceous.
Material:
SDSM 32772: Cranium and lower jaws, rigs, phalanges.
Locality: SDSM V925, South Dakota.
Horizon: 2 meters bellows the Niobrara/Pierre contact.
Biostratigraphy:
Age: Late Cretaceous.
Material:
SDSM 25796: Weahtered cranium, lower jaw, articulated distal half of tail,
rib and phalanges fragments.
MATTER & MILLER, 1972
Locality: Kansas.
Horizon: Niobrara Formation.
Biostratigraphy:
Age: Late Cretaceous.
Material:
V63734: Vertebra.
NICHOLLS, 1988
Locality: Manitoba Province, Canada.
Horizon: Pembina Member, Pierre Shale.
Biostratigraphy:
Age: Campanian Stage, Senonian subepoch, Upper Gulf epoch, Late Cretaceous.
Material:
M77.17.07, FMNH PR 591: Fragmentary specimens.
Locality: South Dakota.
Horizon: Sharon Springs Member, Pierre Shale.
Biostratigraphy:
Age: Campanian Stage, Senonian subepoch, Upper Gulf epoch, Late Cretaceous.
Material:
As per NICHOLLS, 1988
FMNH UR 820, VP 642, VP 2496, KUVP 5033, 27842, 50090, SDSM 10439, USNM 2859-66:
OSBORN
Locality: Kansas.
Horizon: Niobrara Formation.
Biostratigraphy:
Age: Late Cretaceous.
Material:
Number: Not given: Almost complete skull and skeleton.
RUSSELL, 1967
Locality: Texas.
Horizon: Taylor Formation.
Biostratigraphy:
Age: Late Cretaceous.
Material:
TMM 40601: Quadrate.
Locality: NE 1/4, SE 1/4 sec. 36, T2S, R33E, Bighorn County, Montana.
Horizon: Telegraph Creek Formation.
Biostratigraphy:
Age: Late Cretaceous.
Material:
USNM 17909: Skull.
SCHWIMMER, STEWART & WILLIAMS, 1997
Locality: Logan County, Kansas.
Horizon: Niobrara Formation.
Biostratigraphy:
Age: Senonian subepoch, Upper Gulf epoch, Late Cretaceous.
Material:
FHSM VP-3: Associated skeleton.
Note: Left mandible with bite marks showing serrations from a shark.
SHELDON, 1996
Locality: Logan County, Kansas.
Horizon: Niobrara Formation..
Biostratigraphy:
Age: Late Cretaceous.
Material:
YPM 3987, 3975, 127, 12621: Adult.
FMNH-UR-902, YPM 1268: Neonate.
YPM 4002, 1250: Subadult.
Locality: Granham County, Kansas.
Horizon: Niobrara Formation.
Biostratigraphy:
Age: Late Cretaceous.
Material:
YPM 40777, 40757: Subadults.
STEWART & CARPENTER, 1990
Locality: Red Bird, Wyoming.
Horizon: Sharon Springs Member, Pierre Shale.
Biostratigraphy:
Age: Campanian Stage, Senonian subepoch, Upper Gulf epoch, Late Cretaceous.
Material:
UCM 29668: Nearly complete gladius of Tusoteuthis longa that shows puncture
wounds possibly from Tylosaurus proriger.
THEIS & JURGEN, 1995
Locality: Unknown, Kansas.
Horizon: Niobrara Formation.
Biostratigraphy:
Age: Late Cretaceous.
Material:
Collection of Roemer-Museum, Hildesheim: Skull and 4 cervical vertebrae.
THURMOND, 1969
Locality: Cryer Creek 7.5' Quadrangle, 32'14"51N, 96'30'54W, Ellis
County, Texas.
Horizon: Lower marl Member of the Taylor Formation, near base of Wolfe City
Member.
Biostratigraphy:
Age: Late Cretaceous.
Material:
STRECKER MUSEUM 3025: Fragmentary skull.
= Rhinosaurus micromus MARSH, 1872
Etymology:
= Liodon micromus (MARSH, 1872)COPE, 1874
= Tylosaurus micromus (MARSH, 1872) MARSH, 1880
Holotype: YPM 1268
Locality: Smoky Hill River, Kansas.
Horizon: Niobrara Formation.
Biostratigraphy:
Age: Late Cretaceous.
Material: Skull and skeleton.
_____________________________________________________________________________________
Species: nepaeolicus (COPE, 1874) MERRIAM, 1894
Etymology:
= Liodon nepaeolicus COPE, 1874
Holotype: AMNH 1565
Locality: 1/2 mile south of the Solomon River, Kansas.
Horizon: Smoky Hill Chalk Member, Niobrara Formation.
Biostratigraphy:
Age: Santonian or Campanian Stage, Middle Senonian Subepoch, Uppermiddle Gulf
Epoch, Late Cretaceous.
Material: Partial skull and 1 vertebra.
Referred material:
AMNH 124: Fragmentary skull.
AMNH 131:
AMNH 134:
AMNH 1524:
AMNH 1561:
AMNH 2167:
AMNH 2319:
YPM 3969:
YPM 3970:
YPM 3976:
YPM 3979:
YPM 3980: Fragmentary skull.
YPM 3992:
YPM 4000:
20 unnumbered specimens.
USNM 3894:
MCZ 1592:
MCZ 1604:
MCZ 1626:
FHM 1654:
BELL, 1997
FHM VP2209, VP2292:
YPM 40761:
BELL & BARNES, 2007
Locality: Near Terlingua, Big Bend Region, Brewster County,
Texas.
Horizon: Upper San Vicente Member, Boquillas Formation.
Biostratigraphy:
Age: Upper Coniacian Stage, Lowermost Senonian subepoch, Gulf
Epoch, Late Cretaceous.
Material:
MRM KB-M-3: Skull and skeleton.
Note: Has stomach contents of two size classes of Platecarpus af. P.
plainfrons and the shark Ptychodus mortoni.
EVERHART, 2002
Locality: Near Glade, Phillips County, Kansas.
Horizon: Lower Smoky Hill Chalk Member, Niobrara Formation.
Biostratigraphy: Protosphyraena pernicosa zone.
Age: Upper Coniacian Stage, Lowermost Senonian subepoch, Gulf Epoch, Late
Cretaceous.
Material:
FHSM VP-2209: Both maxillae, frontal, anterior portion of the parietal, both
squamosals, the quadrates, both mandibles, cervical and dorsal vertebrae, and
some limb elements.
Locality: Gove County, Kansas.
Horizon: near Hattin’s (1982) Marker Unit 5, Lower
Smoky Hill Chalk Member, Niobrara Formation.
Biostratigraphy:
Age: Upper Coniacian Stage, Lowermost Senonian subepoch, Gulf Epoch, Late
Cretaceous.
Material:
FHSM VP-7262: Premaxilla, complete right and left maxillae, complete frontal,
a large fragment of the posterior parietal, left quadrate, both pterygoids,
portions of both lower jaws including complete left dentary, and numerous vertebrae.
SHELDON,1996
Locality: Logan County, Kansas.
Horizon: Niobrara Formation.
Biostratigraphy:
Age: Late Cretaceous.
Material:
YPM 1304: Subadult.
Locality: Rooks County, Kansas.
Horizon: Niobrara Formation.
Biostratigraphy:
Age: Late Cretaceous.
Material:
YPM 3970: Subadult.
YPM 40752: Adult.
Locality: Trego County, Kansas.
Horizon: Niobrara Formation.
Biostratigraphy:
Age: Late Cretaceous.
Material:
YPM 3974 frontal frontal from JIMENEZ-HUIDOBRO, SIMOES & CALDWELL, 2016.
YPM 3974: Subadult.
YPM 3976: Neonate.
YPM 40445: Late Juvenile.
YPM 40452: Adult.
Locality: Ellis County, Kansas.
Horizon: Niobrara Formation.
Biostratigraphy:
Age: Late Cretaceous.
Material:
YPM 40766, 40764: Early juvenile.
= Tylosaurus kansasensis EVERHART, 2005
Etymology: Named for the Kaw Indians, and American Indian tribe also known
as the Kanza, from which the name of the state (Kansas) is derived and where
allof the known specimens have been collected.
= Species: Nova BELL, 1997
, EVERHART, 2004
Holotype: FHSM VP-2295
Locality: 8 miles northeast of the town of Ellis, Ellis County, Kansas.
Horizon: Lower Smoky Hill Chalk Member, Niobrara Formation.
Biostratigraphy:
Age: Late Coniacian Stage, lowest Senonian subepoch, lower Gulf Epoch, Early
Late Cretaceous.
Material: A complete articulated skull wiht 7 cervical vertebrae.
Note: The frontal and right dentary have several long, deep gouges and punctures
that see to be unhealed bit marks from an fatal encounter with a larger mosasaur
(EVERHART, 2005, 2008).
Paratype:
LACM 127815
Locality: SW of Plainville, Rooks County, Kansas.
Horizon: Lower Smoky Hill Chalk Member, Niobrara Formation.
Biostratigraphy:
Age: Late Coniacian Stage, lowest Senonian subepoch, lower Gulf Epoch, Early
Late Cretaceous.
Material: A complete skull, vertebrae and limb elements.
Locality: Trego County, Kansas.
Horizon: Lower Smoky Hill Chalk Member, Niobrara Formation.
Biostratigraphy:
Age: Late Coniacian Stage, lowest Senonian subepoch, lower Gulf Epoch, Early
Late Cretaceous.
Material:
FHSM VP-2495: A disarticulated, near-complete skull missing the premaxilla and
anteriormost portion of the vertebral column (83 vertebrae) of a juvenile.
FHSM VP-2495frontal from JIMENEZ-HUIDOBRO, SIMOES & CALDWELL, 2016.
Referred material:
FGM V-43: A complete, articulated skull including the right quadrate, premaxilla,
right sclerotic ring and both lower jaws, and 2 cervical vertebrae.
Locality: Ellis County, Kansas.
Horizon: Lower Smoky Hill Chalk Member, Niobrara Formation.
Biostratigraphy:
Age: Late Coniacian Stage, lowest Senonian subepoch, lower Gulf Epoch, Early
Late Cretaceous.
Material:
FHSM VP-15631: Complete skull, pectoral girle and most of both front limbs,
several ribs and a series of 42 consecutive cervical, dorsal and caudal vertebrae.
Locality: Gove County, Kansas.
Horizon: Lower Smoky Hill Chalk Member, Niobrara Formation.
Biostratigraphy:
Age: Late Coniacian Stage, lowest Senonian subepoch, lower Gulf Epoch, Early
Late Cretaceous.
Material:
FHSM VP-78: A nearly complete skull and 4 vertebrae.
FHSM VP-3366: Partial skull, including premaxilla and a right quadrate, 24
cervical and dorsal vertebrae, rib fragments and preserved thoracic cartilage.
Note: Pathologic, 1 rib has a healed fracture.
Note: Serrated bite marks are on the skeletal remains from Squalicorax
falcatus.
FHSM VP-14848: Fragments of the skull, incluiding the parietal, braincase
and left quadrate, 48 cervical, dorsal and caudal vertebrae of a small (2m)
long individual.
FHSM VP-15632: A fragmentary skull, including the premaxilla, frontal/parietal,
both quadrates and both lower jaws, and 5 cervical vertebrae.
Locality: Graham County, Kansas.
Horizon: Lower Smoky Hill Chalk Member, Niobrara Formation.
Biostratigraphy:
Age: Late Coniacian Stage, lowest Senonian subepoch, lower Gulf Epoch, Early
Late Cretaceous.
Material:
FHSM VP-9350: Partial skull including the premaxilla, left maxilla, left
quadrate and both lower jaws of a juvenile.
Locality: Unknown, Logan County, Kansas.
Horizon: Smoky Hill Chalk Member, Niobrara Formation.
Biostratigraphy:
Age: Late Coniacian Stage, lowest Senonian subepoch, lower Gulf Epoch, Early
Late Cretaceous.
Material:
MCZ 1589: Fragments of an adult skull including the premaxilla, left maxilla,
right dentary and left quadrate.
Locality: Graham County, Kansas.
Horizon: Lower Smoky Hill Chalk Member, Niobrara Formation.
Biostratigraphy:
Age: Late Coniacian Stage, lowest Senonian subepoch, lower Gulf Epoch, Early
Late Cretaceous.
Material:
YPM 40796: Fragmentary remains, including the parietal and a quadrate,but
no premaxilla.
Locality: 0.5 km of the locality in Gove County that produced FHSM VP-7262, Kansas.
Horizon: Near Hattin’s (1982) Marker Unit 5, Lower Smoky Hill Chalk
Member, Niobrara Formation.
Biostratigraphy:
Age: Upper Coniacian Stage, Lowermost Senonian subepoch, Gulf Epoch, Late
Cretaceous.
Material:
FHSM VP-13742: A large but badly weathered skull, premaxilla, portions of
the maxillae, a complete frontal and both lower jaws (lacking both quadrates)
and limb elements.
Note: 3 tips of shart teeth, Cretoxyrhina mantelli, were embedded
in the top of the premaxilla and lateral side of both dentaries. The anterior
end of the
premaxilla had apparently been sheard off by the bite of a shark and the
damaged bone shows on evidence of healing.
_____________________________________________________________________________________
Species: bernardi (DOLLO, 1885) JIMENEZ-HUIDOBRO & CALDWELL, 2019
= Hainosaurus bernardi DOLLO, 1885
Etymology:
Holotype: IRSNB R23 (old no. R1564)
Locality: 'La Malogne", near the town of Mesvin, Belgium.
Horizon: Ciply Phosphatic Chalk.
Biostratigraphy:
Age: Upper Maastrichtian Stage, uppermost Senonian subepoch, Gulf Epoch, Late
Cretaceous.
Material: Fragmentary skull and fragmentary skeleton.
Referred material:
Locality: Baudoir, Belgium.
Horizon: Phosphatic Chalk.
Biostratigraphy:
Age: Maastrichtian Stage, uppermost Senonian subepoch, Gulf Epoch, Late Cretaceous.
Material:
IRSNB 3672: Nearly complete skull and large number of vertebrae.
= ?Leiodon anceps DEPERET & RUSSO, 1925
Locality:
Horizon:
Biostratigraphy:
Age:
Material:
Number: Not given:
= ?Leiodon anceps PERSSON, 1959
Locality:
Horizon:
Biostratigraphy:
Age:
Material:
Number: Not given:
= Hainosaurus cf. bernardi MULDER & MAI, 1999
Locality: Folx-les-caves, Eastern Brabant Massif, Belgium.
Horizon: Tuffeau jaunatre.
Biostratigraphy: Zone CC 22 of Sissigh 1977.
Age: Early Late Campanian Stage, Senonian subepoch, Upper Gulf epoch, Late
Cretaceous.
Material:
MB.R. 890: Parietal.
_____________________________________________________________________________________
Species: pembinensis (NICHOLLS, 1988)
BULLARD & CALDWELL, 2007
= Hainosaurus pembinensis NICHOLLS, 1988
Etymology:
Holotype: MT 2
Locality: MDM quarry No. 18, North Cox pit, NW 1/4, tp. 4, rge, 7, West Principal,
West Principal Meridian, Manitoba Province, Canada.
Horizon: Lower part of the Pembina Member, Pierre shale.
Biostratigraphy:
Age: Lower Campanian Stage, Senonian subepoch, Upper Gulf epoch, Late Cretaceous.
Material: Incomplete skull and mandible, left humerus, femur, articulated
vertebral column lacking first 2, possibly the 3rd cervical vertebrae.
Skull after BULLARD & CALDWELL, 2010. A) dorsal and B) lateral views.
Referred material:
MDM M73.17.02:
MDM M74.01.02:
MDM M74.05.06:
MDM M74.06.06: Fragmentary skull and fragmentary skeleton.
MDM M74.08.06:
MDM M74.09.06:
MDM M77.03.09:
MDM M77.05.07:
MDM M83.13.18:
MMMN V95:
NMC 40320:
_____________________________________________________________________________________
Species: neumilleri (MARTIN, 2007) JIMENEZ-HUIDOBRO & CALDWELL, 2019
= Hainosaurus newmilleri MARTIN, 2007b
Etymology: In honor of Paul Neumiller, who discovered the specimen and reported
its occurrence to U.S. Army Corps of Engineers personallel on whose holding
the specimen was found. The scientific community commends Neumiller for his
natural curiosity, kindness, and proper action to preserve the specimen for
the public good.
Holotype: SDSM 75705
Locality: SDSM Locality V2005-20, along the bank of the Missouri River, Gregory
County, South Dakota.
Horizon: DeGrey Formation, Pierre Shale Group.
Biostratigraphy:
Age: Late Campanian Stage, Senonian subepoch, Upper Gulf epoch, Late Cretaceous.
Material: Partial skull, paddle bones, and associated vertebrae.
_____________________________________________________________________________________
Species: saskatchewanensis JIMENZ-HUIDOBRO, CALDWELL, PARAELLA & BULLARD, 2018
Etymology: In reference to the Canadian province of Saskatchewan, where the only specimen was found.
Holotype: RSM P2588.1
Locality: Near Herbert Ferry, southern shore of Lake Diefenbaker, Saskatchewan Province, Canada.
Horizon: Snakebite Member, Bearpaw Formation.
Biostratigraphy:
Age: Upper Campanian Stage, Senonian subepoch, Upper Gulf epoch, Late Cretaceous.
Material: A moderately complete and articulated skull associated with a quite complete disarticulated postcranium.
Tylosaurus saskatchewanensis after JIMENZ-HUIDOBRO, CALDWELL, PARAELLA & BULLARD, 2018, dorsal view of skull.
Tylosaurus saskatchewanensis after JIMENZ-HUIDOBRO, CALDWELL, PARAELLA & BULLARD, 2018: A) right side of skull; B) left side of skull.