96MADAGASCAR

MAHAJANGA BASIN

FROM THE NEIGHBOURING OF AMBONDROMAMY

BATHONIAN STAGE

MIDDLE DOGGER EPOCH

MIDDLE JURASSIC

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1904

SUPERLEGION: NEODIAPSIDA BENTON, 1985

LEGION: ARCHOSAUROMORPHA HUENE, 1946

SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985

INFRALEGION: DINOSAURIFORMES NOVAS, 1992

SUPERCOHORT: DINOSAURIA OWEN, 1842

COHORT: THEROPODA EMEND MARSH, 1881

Incertae sedis

Morphotype 2

Gen. sp indet.

_____________________________________________________________________________________

MADAGASCAR

MAHAJANGA BASIN

HILLS W OF AMBONDROMAMY

BATHONIAN STAGE

MIDDLE DOGGER EPOCH

MIDDLE JURASSIC

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1903

INFRACLASS: NEODIAPSIDA BENTON, 1985

LEGION: ARCHOSAUROMORPHA HUENE, 1946

SUPERCOHORT: CRUROTARSI SERENO & ARCUCCI, 1990

COHORT: CROCODYLOTARSI BENTON & CLARK, 1988

MAGNORDER: SUCHIA KREBS, 1974

SUPERORDER: CROCODYLOMORPHA WALKER, 1968

GRANDORDER: CROCODILIFORMES HAY, 1930

MICRORDER: MESOEUROCROCODYLIA WHETSTONE & WHYBROW, 1983

ORDER: MESOSUCHIA EMEND HUXLEY, 1875

PARVORDER: METASUCHIA BENTON & CLARK, 1988

GONDWANASUCHIA DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004

NOTOSUCHIMORPHA DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004

NOTOSUCHIFORMES DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004

TERRIASUCHIA DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004

INFRAORDER: NOTOSUCHIA GASPARINI, 1971

Incertae sedis

*Razanandrongobe sakalavae

_____________________________________________________________________________________

MADAGASCAR

MAHAJANGA BASIN

HILLS N-NW OF AMBONDROMAMY

BATHONIAN STAGE

MIDDLE DOGGER EPOCH

MIDDLE JURASSIC

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1903

INFRACLASS: NEODIAPSIDA BENTON, 1985

LEGION: ARCHOSAUROMORPHA HUENE, 1946

SUPERCOHORT: CRUROTARSI SERENO & ARCUCCI, 1990

COHORT: CROCODYLOTARSI BENTON & CLARK, 1988

MAGNORDER: SUCHIA KREBS, 1974

SUPERORDER: CROCODYLOMORPHA WALKER, 1968

GRANDORDER: CROCODILIFORMES HAY, 1930

MICRORDER: MESOEUROCROCODYLIA WHETSTONE & WHYBROW, 1983

ORDER: MESOSUCHIA EMEND HUXLEY, 1875

PARVORDER: METASUCHIA BENTON & CLARK, 1988

GONDWANASUCHIA DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004

NOTOSUCHIMORPHA DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004

NOTOSUCHIFORMES DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004

TERRIASUCHIA DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004

INFRAORDER: NOTOSUCHIA GASPARINI, 1971

Incertae sedis

Razanandrongobe sakalavae

_____________________________________________________________________________________

MADAGASCAR

MAHAJANGA BASIN

UNDETERMINED LOCALITIES

BATHONIAN STAGE

MIDDLE DOGGER EPOCH

MIDDLE JURASSIC

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1903

INFRACLASS: NEODIAPSIDA BENTON, 1985

LEGION: ARCHOSAUROMORPHA HUENE, 1946

SUPERCOHORT: CRUROTARSI SERENO & ARCUCCI, 1990

COHORT: CROCODYLOTARSI BENTON & CLARK, 1988

MAGNORDER: SUCHIA KREBS, 1974

SUPERORDER: CROCODYLOMORPHA WALKER, 1968

GRANDORDER: CROCODILIFORMES HAY, 1930

MICRORDER: MESOEUROCROCODYLIA WHETSTONE & WHYBROW, 1983

ORDER: MESOSUCHIA EMEND HUXLEY, 1875

PARVORDER: METASUCHIA BENTON & CLARK, 1988

GONDWANASUCHIA DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004

NOTOSUCHIMORPHA DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004

NOTOSUCHIFORMES DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004

TERRIASUCHIA DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004

INFRAORDER: NOTOSUCHIA GASPARINI, 1971

Incertae sedis

Razanandrongobe sakalavae

_____________________________________________________________________________________

MADAGASCAR

AMBILOBE

MAHAJANGA

MORONDAVA

AMBONDROMAMY

AMBONDROMAMY FAUNA

ISALO IIIB

BATHONIAN STAGE

UPPER MIDDLE DOGGER EPOCH

MIDDLE JURASSIC

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: AMPHIBIA LINNAEUS, 1758

SUBCLASS: LISSAMPHIBIA, HAECKEL, 1866?

Incertae sedis

Gen. sp indet.

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: CHELONOMORPHA KUHN, 1960

SUPERCOHORT: TESTUDINES LINNAEUS, 1758

Incertae sedis

Gen. sp indet.

SUBCLASS: DIAPSIDA OSBORN, 1903

INFRACLASS: NEODIAPSIDA BENTON, 1985

DIVISION: LEPIDOSAUROMORPHA BENTON, 1983

SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)

MAGNORDER: SPHENODONTIFORMES Hay, 1930

ORDER: SPHENODONTIA WILLISTON, 1925

Family: SPHENODONTIDAE Cope, 1869

Gen. sp indet.

LEGION: ARCHOSAUROMORPHA HUENE, 1946

SUPERCOHORT: CRUROTARSI SERENO & ARCUCCI, 1990

COHORT: CROCODYLOTARSI BENTON & CLARK, 1988

MAGNORDER: SUCHIA KREBS, 1974

SUPERORDER: CROCODYLOMORPHA WALKER, 1968

GRANDORDER: CROCODILIFORMES HAY, 1930

MICRORDER: MESOEUROCROCODYLIA WHETSTONE & WHYBROW, 1983

ORDER: MESOSUCHIA EMEND HUXLEY, 1875

PARVORDER: METASUCHIA BENTON & CLARK, 1988

INFRAORDER: METAMESOSUCHIA HULKE, 1878

NEOSUCHIA CLARK, IN BENTON & CLARK, 1988

Family: ATOPOSAURIDAE Gervais, 1871

Gen. sp indet.

SUPERCOHORT: AVEMETATARSALIA BENTON, 1999

COHORT: PROLACERTIFORMES CAMP, 1945 NOVA

SUPERORDER: TAPINOPLATIA PETERS, 2000

GRANDORDER: CHARACIOPODA PETERS, 2000

MICRORDER: FENESTRASAURIA PETERS, 2000

ORDER: PTEROSAURIA KAUP, 1834

SUBORDER: RHAMPHORHYNCHOIDEA PLIENINGER, 1901

Incertae sedis

Gen. sp indet.

SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985

INFRALEGION: DINOSAURIFORMES NOVAS, 1992

SUPERCOHORT: DINOSAURIA OWEN, 1842

COHORT: THEROPODA EMEND MARSH, 1881

ORDER: Incertae sedis

Gen. sp indet.

COHORT: ORNITHISCHIFORMES COOPER, 1985

SUPERORDER: SAUROPODOMORPHA HUENE, 1932

GRANDORDER: BRONTOSAURIA OLSHEVSKY, 1991 (emend)

MICRORDER: SAUROPODA MARSH, 1878

ORDER: EUSAUROPODA

SUBORDER: NEOSAUROPODA BONAPARTE, 1986

Incertae sedis

Gen. sp indet.

MAGNORDER: ORNITHISCHIA SEELEY, 1888

Incertae sedis

Gen. sp indet.

_____________________________________________________________________________________

NORTHERN MADAGASCAR

MAJUNGA BASIN

NEAR ANDRANOMANY

MANARY-ABO

BATHONIAN STAGE

MIDDLE DOGGER EPOCH

MIDDLE JURASSIC

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1904

SUPERLEGION: NEODIAPSIDA BENTON, 1985

LEGION: ARCHOSAUROMORPHA HUENE, 1946

SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985

INFRALEGION: DINOSAURIFORMES NOVAS, 1992

SUPERCOHORT: DINOSAURIA OWEN, 1842

COHORT: ORNITHISCHIFORMES COOPER, 1985

SUPERORDER: SAUROPODOMORPHA HUENE, 1932

GRANDORDER: BRONTOSAURIA OLSHEVSKY, 1991 (emend)

MICRORDER: SAUROPODA MARSH, 1878

ORDER: EUSAUROPODA

Incertae sedis

non Bothriospondylus, B?madagascariensis

_____________________________________________________________________________________


MADAGASCAR

MAHAJANGA BASIN

FROM AN AREA BETWEEN AMBONDROMAMY AND PORT BERGE

BATHONIAN STAGE

MIDDLE DOGGER EPOCH

MIDDLE JURASSIC

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1904

SUPERLEGION: NEODIAPSIDA BENTON, 1985

LEGION: ARCHOSAUROMORPHA HUENE, 1946

SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985

INFRALEGION: DINOSAURIFORMES NOVAS, 1992

SUPERCOHORT: DINOSAURIA OWEN, 1842

COHORT: THEROPODA EMEND MARSH, 1881

Incertae sedis

Morphotype 1

Gen. sp indet.

Morphotype 2

Gen. sp indet.

Morphotype 3

Gen. sp indet.

Morphotype 4

Gen. sp indet.

Morphotype 5

Gen. sp indet.

Morphotype 6

Gen. sp indet.

Morphotype 7

Gen. sp indet.

_____________________________________________________________________________________

MADAGASCAR

MAHAJANGA BASIN

NORTH OF PORT-BERGE

BATHONIAN STAGE

MIDDLE DOGGER EPOCH

MIDDLE JURASSIC

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1904

SUPERLEGION: NEODIAPSIDA BENTON, 1985

LEGION: ARCHOSAUROMORPHA HUENE, 1946

SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985

INFRALEGION: DINOSAURIFORMES NOVAS, 1992

SUPERCOHORT: DINOSAURIA OWEN, 1842

COHORT: THEROPODA EMEND MARSH, 1881

Incertae sedis

Morphotype 1

Gen. sp indet.

Morphotype 2

Gen. sp indet.

Morphotype 3

Gen. sp indet.

Morphotype 4

Gen. sp indet.

Morphotype 5

Gen. sp indet.

Morphotype 6

Gen. sp indet.

_____________________________________________________________________________________

MADAGASCAR

MAHAJANGA BASIN

BETWEEN TSINJORAN AND THE BRIDGE ON THE KAMORO RIVER

BATHONIAN STAGE

MIDDLE DOGGER EPOCH

MIDDLE JURASSIC

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1904

SUPERLEGION: NEODIAPSIDA BENTON, 1985

LEGION: ARCHOSAUROMORPHA HUENE, 1946

SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985

INFRALEGION: DINOSAURIFORMES NOVAS, 1992

SUPERCOHORT: DINOSAURIA OWEN, 1842

COHORT: THEROPODA EMEND MARSH, 1881

Incertae sedis

Morphotype 8

Gen. sp indet.

_____________________________________________________________________________________

MADAGASCAR

MAHAJANGA BASIN

NEAR THE BRIDGE ON THE KAMORO RIVER ALONG THE WEATHERED SIDES OF A LOW TANETY (LOCAL NAME FOR "HILL") ON THE RIGHT BANK OF THE KAMORO RIVER

BATHONIAN STAGE

MIDDLE DOGGER EPOCH

MIDDLE JURASSIC

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1904

SUPERLEGION: NEODIAPSIDA BENTON, 1985

LEGION: ARCHOSAUROMORPHA HUENE, 1946

SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985

INFRALEGION: DINOSAURIFORMES NOVAS, 1992

SUPERCOHORT: DINOSAURIA OWEN, 1842

COHORT: THEROPODA EMEND MARSH, 1881

Incertae sedis

Gen. sp indet.

_____________________________________________________________________________________

MADAGASCAR

MAHAJANGA BASIN

SOUTH-WEST TO AMBONDROMAMY

FROM AN AREA NEAR TSINJORANO

BATHONIAN STAGE

MIDDLE DOGGER EPOCH

MIDDLE JURASSIC

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1904

SUPERLEGION: NEODIAPSIDA BENTON, 1985

LEGION: ARCHOSAUROMORPHA HUENE, 1946

SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985

INFRALEGION: DINOSAURIFORMES NOVAS, 1992

SUPERCOHORT: DINOSAURIA OWEN, 1842

COHORT: THEROPODA EMEND MARSH, 1881

Incertae sedis

Morphotype 2

Gen. sp indet.

Morphotype 3

Gen. sp indet.

Morphotype 4

Gen. sp indet.

Morphotype 6

Morphotype 8

Gen. sp indet.

_____________________________________________________________________________________

NORTHWEST MADAGASCAR

MAHAJANGA PROVINCE

UNDETERMINED HILLS WEST OF AMBONDROMAMY

ISALO IIIB SUBUNIT

'FACIES MIXTE DINOSAURIENS’

BATHONIAN STAGE

MIDDLE DOGGER EPOCH

MIDDLE JURASSIC

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA

SUPERLEGION: NEODIAPSIDA BENTON, 1985

LEGION: ARCHOSAUROMORPHA HUENE, 1946

ARCHOSAURIA INCERTAE SEDIS

*Razanadrongobe sakalavae

_____________________________________________________________________________________

NORTHWEST MADAGASCAR

MAHAJANGA PROVINCE

HILLS NNW OF ANDRANOMAMY

ISALO IIIB SUBUNIT

'FACIES MIXTE DINOSAURIENS’

BATHONIAN STAGE

MIDDLE DOGGER EPOCH

MIDDLE JURASSIC

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA

SUPERLEGION: NEODIAPSIDA BENTON, 1985

LEGION: ARCHOSAUROMORPHA HUENE, 1946

ARCHOSAURIA INCERTAE SEDIS

Razanadrongobe sakalavae

_____________________________________________________________________________________

N. W. MADAGASCAR

MAHAJANGA PROVINCE

8 KM (5 MILES) NORTH OF ANABORANO, AND 75 MILES SSW OF DIEGO SUAREZ

AMBARAKARAKA

MIDDLE SAKAMENA FORMATION

INDUAN STAGE

UPPER SCYTHIAN EPOCH

EARLY TRIASSIC

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: AMPHIBIA LINNAEUS, 1758

SUBCLASS: LABYRINTHODONTIA OWEN, 1860

SUPERORDER: TEMNOSPONDYLIA ZITTEL, 1887

MICROORDER: LIMNARCHIA YATES & WARREN, 2000

ORDER: STEREOSPONDYLOMORPHA YATES & WARREN, 2000

Family: LYDEKKERINIDAE Watson, 1919

*Deltacephalus whitei

_____________________________________________________________________________________


MADAGASCAR

MAHAJANGA PROVINCE

JURASSIC?

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1903

SUPERLEGION: NEODIAPSIDA BENTON, 1985

LEGION: ARCHOSAUROMORPHA HUENE, 1946

SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985

INFRALEGION: DINOSAURIFORMES NOVAS, 1992

SUPERCOHORT: DINOSAURIA OWEN, 1842

COHORT: ORNITHISCHIFORMES COOPER, 1985

SUPERORDER: SAUROPODOMORPHA HUENE, 1932

Incertae sedis

Gen. sp indet.


_____________________________________________________________________________________

 

MADAGASCAR

MAHAJANGA PROVINCE

MAJUNGA REGION

SW OF THE BASIN OF KELIFELY

KANDREHO FORMATION

MIDDLE OR UPPER TOARCIAN STAGE

UPPER LIAS EPOCH

EARLY JURASSIC

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1903

SUPERLEGION: NEODIAPSIDA BENTON, 1985

LEGION: ARCHOSAUROMORPHA HUENE, 1946

SUPERCOHORT: CRUROTARSI SERENO & ARCUCCI, 1990

COHORT: CROCODYLOTARSI BENTON & CLARK, 1988

MAGNORDER: SUCHIA KREBS, 1974

SUPERORDER: CROCODYLOMORPHA WALKER, 1968

GRANDORDER: CROCODILIFORMES HAY, 1930

MICRORDER: MESOEUROCROCODYLIA WHETSTONE & WHYBROW, 1983

ORDER: MESOSUCHIA EMEND HUXLEY, 1875

ORDER: THALATTOSUCHIA EMEND FRAAS, 1901

Family: TELEOSAURIDAE Geoffroy, 1831

Incertae sedis

Steneosaurus sp

_____________________________________________________________________________________

MADAGASCAR

MAJUNGA PROVINCE

MAJUNGA BASIN

MIDDLE JURASSIC

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1903

SUPERLEGION: NEODIAPSIDA BENTON, 1985

LEGION: ARCHOSAUROMORPHA HUENE, 1946

SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985

INFRALEGION: DINOSAURIFORMES NOVAS, 1992

SUPERCOHORT: DINOSAURIA OWEN, 1842

COHORT: ORNITHISCHIFORMES COOPER, 1985

SUPERORDER: SAUROPODOMORPHA HUENE, 1932

GRANDORDER: BRONTOSAURIA OLSHEVSKY, 1991 (emend)

MICRORDER: SAUROPODA MARSH, 1878

Genus nova.

_____________________________________________________________________________________

NORTHWESTERN MADAGASAR

MAHAJANGA PROVINCE

ABOUT 150 KM SW OF THE COASTAL TOWN OF MAHAJANGA (MAJUNGA)

FROM THE VICINITY OF AMBONDROMAMY

ISALO IIIB FORMATION

BATHONIAN STAGE

MIDDLE DOGGER EPOCH

MIDDLE JURASSIC

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1903

SUPERLEGION: NEODIAPSIDA BENTON, 1985

LEGION: ARCHOSAUROMORPHA HUENE, 1946

SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985

INFRALEGION: DINOSAURIFORMES NOVAS, 1992

SUPERCOHORT: DINOSAURIA OWEN, 1842

COHORT: THEROPODA EMEND MARSH, 1881

AVEPODA PAUL, 2002

SUPERORDER: NEOTHEROPODA BAKKER, 1986

GRANDORDER: TETANURAE GAUTHIER, 1986

MICRORDER: COELUROSAURIA HUENE, 1914

ORDER: SEGNOSAURIA BARSBOLD & PERLE, 1980

Family: Nova?

*Archaeodontosaurus descouensi

_____________________________________________________________________________________

MADAGASCAR

MAHAJANGA PROVINCE

20 M EASTWARD OF THE BAY NARINDA

UNNAMED UNIT

BATHONIAN STAGE

MIDDLE DOGGER EPOCH

MIDDLE JURASSIC

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1904

SUPERLEGION: NEODIAPSIDA BENTON, 1985

LEGION: ARCHOSAUROMORPHA HUENE, 1946

SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985

INFRALEGION: DINOSAURIFORMES NOVAS, 1992

SUPERCOHORT: DINOSAURIA OWEN, 1842

COHORT: ORNITHISCHIFORMES COOPER, 1985

SUPERORDER: SAUROPODOMORPHA HUENE, 1932

GRANDORDER: BRONTOSAURIA OLSHEVSKY, 1991 (emend)

MICRORDER: SAUROPODA MARSH, 1878

ORDER: EUSAUROPODA

Incertae sedis

non Bothriospondylus, B?madagascariensis

_____________________________________________________________________________________

 

 

MADAGASCAR

MAHAJANGA PROVINCE

MAHAJANGA BASIN

AMBONDROMANY REGION

ISALO III

BATHONIAN STAGE

MIDDLE DOGGER EPOCH

MIDDLE JURASSIC

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: MAMMALIA LINNAEUS, 1758

SUBCLASS: PROTOTHERIA GILL, 1872

ORDER: INCERTAE SEDIS

Incertae sedis

*Ambondro mahabo

_____________________________________________________________________________________

MAHAJANA PROVINCE

MAHAJANA PROVINCE

ANDRANOSAMONTA

LATE JURASSIC

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1904

SUPERLEGION: NEODIAPSIDA BENTON, 1985

LEGION: ARCHOSAUROMORPHA HUENE, 1946

SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985

INFRALEGION: DINOSAURIFORMES NOVAS, 1992

SUPERCOHORT: DINOSAURIA OWEN, 1842

COHORT: ORNITHISCHIFORMES COOPER, 1985

SUPERORDER: SAUROPODOMORPHA HUENE, 1932

GRANDORDER: BRONTOSAURIA OLSHEVSKY, 1991 (emend)

MICRORDER: SAUROPODA MARSH, 1878

ORDER: EUSAUROPODA

Incertae sedis

non Bothriospondylus, B?madagascariensis

_____________________________________________________________________________________

 

MADAGASCAR

MAHAJANGA PROVINCE

ANKINGANIVALKA

LATE JURASSIC

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1904

SUPERLEGION: NEODIAPSIDA BENTON, 1985

LEGION: ARCHOSAUROMORPHA HUENE, 1946

SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985

INFRALEGION: DINOSAURIFORMES NOVAS, 1992

SUPERCOHORT: DINOSAURIA OWEN, 1842

COHORT: ORNITHISCHIFORMES COOPER, 1985

SUPERORDER: SAUROPODOMORPHA HUENE, 1932

GRANDORDER: BRONTOSAURIA OLSHEVSKY, 1991 (emend)

MICRORDER: SAUROPODA MARSH, 1878

ORDER: EUSAUROPODA

Incertae sedis

non Bothriospondylus, B?madagascariensis

_____________________________________________________________________________________

MADAGASCAR

MAHAJANGA PROVINCE

PORT-BERGE

LATE JURASSIC

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1904

SUPERLEGION: NEODIAPSIDA BENTON, 1985

LEGION: ARCHOSAUROMORPHA HUENE, 1946

SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985

INFRALEGION: DINOSAURIFORMES NOVAS, 1992

SUPERCOHORT: DINOSAURIA OWEN, 1842

COHORT: ORNITHISCHIFORMES COOPER, 1985

SUPERORDER: SAUROPODOMORPHA HUENE, 1932

GRANDORDER: BRONTOSAURIA OLSHEVSKY, 1991 (emend)

MICRORDER: SAUROPODA MARSH, 1878

ORDER: EUSAUROPODA

Incertae sedis

non Bothriospondylus, B?madagascariensis

_____________________________________________________________________________________

 

MADAGASCAR

MAHAJANGA PROVINCE

NE REPUBLICK MALGACHE

MAJUNGA

LATE JURASSIC

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1904

SUPERLEGION: NEODIAPSIDA BENTON, 1985

LEGION: ARCHOSAUROMORPHA HUENE, 1946

SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985

INFRALEGION: DINOSAURIFORMES NOVAS, 1992

SUPERCOHORT: DINOSAURIA OWEN, 1842

COHORT: ORNITHISCHIFORMES COOPER, 1985

SUPERORDER: SAUROPODOMORPHA HUENE, 1932

GRANDORDER: BRONTOSAURIA OLSHEVSKY, 1991 (emend)

MICRORDER: SAUROPODA MARSH, 1878

ORDER: EUSAUROPODA

Incertae sedis

non Bothriospondylus, B?madagascariensis

_____________________________________________________________________________________

 

 

MADAGASCAR

MAHAJANGA PROVINCE

RIGHT BANK OF THE RIVER BETSIBOKA

MAEVARANO

CENOMANIAN STAGE

UPPERMIDDLE GALLIC SUBEPOCH

LOWERMOST GULF EPOCH

EARLY CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1903

SUPERLEGION: NEODIAPSIDA BENTON, 1985

LEGION: ARCHOSAUROMORPHA HUENE, 1946

SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985

INFRALEGION: DINOSAURIFORMES NOVAS, 1992

SUPERCOHORT: DINOSAURIA OWEN, 1842

COHORT: THEROPODA EMEND MARSH, 1881

AVEPODA PAUL, 2002

SUPERORDER: NEOTHEROPODA BAKKER, 1986

GRANDORDER: CERATOSAURIA NOVAS, 1992

AVEROSTRA PAUL, 2001

ORDER: CERATOSAURIA MARSH, 1884

Superfamily: ABELISAUROIDEA Novas, 1989

Family: ABELISAURIDAE Bonaparte & Novas, 1985

Subfamily: CARNOTAURINAE Sereno, 1998

Majungasaurus crenatissimus

_____________________________________________________________________________________
MADASCAR

MAHAJANGA PROVINCE

MAJUNGA

LOWER CENOMANIAN STAGE

UPPERMIDDLE GALLIC SUBEPOCH

LOWERMOST GULF EPOCH

EARLY CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1903

SUPERLEGION: NEODIAPSIDA BENTON, 1985

LEGION: ARCHOSAUROMORPHA HUENE, 1946

SUPERCOHORT: CRUROTARSI SERENO & ARCUCCI, 1990

COHORT: CROCODYLOTARSI BENTON & CLARK, 1988

MAGNORDER: SUCHIA KREBS, 1974

SUPERORDER: CROCODYLOMORPHA WALKER, 1968

GRANDORDER: CROCODILIFORMES HAY, 1930

MICRORDER: MESOEUROCROCODYLIA WHETSTONE & WHYBROW, 1983

ORDER: MESOSUCHIA EMEND HUXLEY, 1875

PARVORDER: METASUCHIA BENTON & CLARK, 1988

INFRAORDER: METAMESOSUCHIA HULKE, 1878

NEOSUCHIA CLARK, IN BENTON & CLARK, 1988

Family: TREMATOCHAMPSIDAE Buffetaut, 1974

*Miadanasuchus oblita
= Trematochampsa oblita

_____________________________________________________________________________________

MADAGASCAR

MAHAJANGA PROVINCE

RIGHT BANKS OF THE RIVER BETSIOBKA

MAEVARANO

MEME LOCAILTY

CENOMANIAN STAGE

UPPERMIDDLE GALLIC SUBEPOCH

LOWERMOST GULF EPOCH

EARLY CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1903

SUPERLEGION: NEODIAPSIDA BENTON, 1985

LEGION: ARCHOSAUROMORPHA HUENE, 1946

SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985

INFRALEGION: DINOSAURIFORMES NOVAS, 1992

SUPERCOHORT: DINOSAURIA OWEN, 1842

COHORT: THEROPODA EMEND MARSH, 1881

AVEPODA PAUL, 2002

SUPERORDER: NEOTHEROPODA BAKKER, 1986

GRANDORDER: CERATOSAURIA NOVAS, 1992

AVEROSTRA PAUL, 2001

ORDER: CERATOSAURIA MARSH, 1884

Superfamily: ABELISAUROIDEA Novas, 1989

Family: ABELISAURIDAE Bonaparte & Novas, 1985

Subfamily: CARNOTAURINAE Sereno, 1998

Majungasaurus crenatissimus

 

COHORT: ORNITHISCHIFORMES COOPER, 1985

SUPERORDER: SAUROPODOMORPHA HUENE, 1932

Incertae sedis

Gen. sp indet.


_____________________________________________________________________________________

 

MADAGASCAR

MAHAJANGA PROVINCE

MAHAJANGA BASIN

ANKAZOMIHABOKA SANDSTONE

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: CHELONOMORPHA KUHN, 1960

SUPERCOHORT: TESTUDINES LINNAEUS, 1758

COHORT: CASICHELYDIA emend GAFFNEY & MEYLAN, 1988

MAGNORDER: PLEURODIRA GAFFNEY & MEYLAN, 1988

ORDER: PLEURODIRIDA COPE, 1864

SUBORDER: EUPLEURODIRA GAFFNEY & MEYLAN, 1988

Superfamily: PELOMEDUSOIDES Cope, 1868 Cope, 1868

Family: PELOMEDUSIDAE Cope, 1868

Gen. sp indet.

SUBCLASS: DIAPSIDA OSBORN, 1903

SUPERLEGION: NEODIAPSIDA BENTON, 1985

LEGION: ARCHOSAUROMORPHA HUENE, 1946

SUPERCOHORT: CRUROTARSI SERENO & ARCUCCI, 1990

COHORT: CROCODYLOTARSI BENTON & CLARK, 1988

MAGNORDER: SUCHIA KREBS, 1974

SUPERORDER: CROCODYLOMORPHA WALKER, 1968

GRANDORDER: CROCODILIFORMES HAY, 1930

MICRORDER: MESOEUROCROCODYLIA WHETSTONE & WHYBROW, 1983

ORDER: MESOSUCHIA EMEND HUXLEY, 1875

PARVORDER: METASUCHIA BENTON & CLARK, 1988

INFRAORDER: METAMESOSUCHIA HULKE, 1878

NEOSUCHIA CLARK, IN BENTON & CLARK, 1988

Family: TREMATOCHAMPSIDAE Buffetaut, 1974

Incertae sedis

Gen. sp indet.

_____________________________________________________________________________________

MADAGASCAR

MAJUNGA PROVINCE

IN A VILLAGE NEAR MAJUNGA

2/3 KM NORTH OF HOTEL BERIVOTRO

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1903

SUPERLEGION: NEODIAPSIDA BENTON, 1985

LEGION: ARCHOSAUROMORPHA HUENE, 1946

SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985

INFRALEGION: DINOSAURIFORMES NOVAS, 1992

SUPERCOHORT: DINOSAURIA OWEN, 1842

COHORT: THEROPODA EMEND MARSH, 1881

AVEPODA PAUL, 2002

SUPERORDER: NEOTHEROPODA BAKKER, 1986

GRANDORDER: CERATOSAURIA NOVAS, 1992

AVEROSTRA PAUL, 2001

ORDER: CERATOSAURIA MARSH, 1884

Superfamily: ABELISAUROIDEA Novas, 1989

Family: ABELISAURIDAE Bonaparte & Novas, 1985

Subfamily: CARNOTAURINAE Sereno, 1998

Majungasaurus crenatissimus
= Megalosaurus crenatissimus


_____________________________________________________________________________________

 

MADAGASCAR

MAHAJANGA PROVINCE

MAHAJANGA BASIN

MAEVARANO FORMATION

CAMPANIAN STAGE?

SENONIAN SUBEPOCH

GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: MAMMALIA LINNAEUS, 1758

SUBCLASS: PROTOTHERIA GILL, 1872

INFRACLASS: TRECHNOTHERIA McKENNA, 1975

LEGION: CLADOTHERIA McKENNA, 1975

SUBLEGION: ZATHERIA MCKENNA, 1975

INFRALEGION: INCERTAE SEDIS

Genus: Nova


_____________________________________________________________________________________

MADAGASCAR

MAHAJANGA PROVINCE

50 KM FROM MAJUNGA

REGION OF BERIVOTRA

UPPER SENONIAN SUBEPOCH

GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: CHELONOMORPHA KUHN, 1960

SUPERCOHORT: TESTUDINES LINNAEUS, 1758

COHORT: CASICHELYDIA emend GAFFNEY & MEYLAN, 1988

MAGNORDER: PLEURODIRA GAFFNEY & MEYLAN, 1988

ORDER: PLEURODIRIDA COPE, 1864

SUBORDER: EUPLEURODIRA GAFFNEY & MEYLAN, 1988

Superfamily: PELOMEDUSOIDES Cope, 1868 Cope, 1868

Family: PELOMEDUSIDAE Cope, 1868

Gen. sp undescribed

Subfamily: PELOMEDUSINAE Cope, 1868

Bothremyididae gen sp indet

_____________________________________________________________________________________


MADAGASCAR

MAHAJANGA

MAJUNGA PROVINCE

BERIVOTRO

BERIVOTRO FORMATION

SENONIAN OR CAMPANIAN STAGE

SENONIAN SUBEPOCH

GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1903

SUPERLEGION: NEODIAPSIDA BENTON, 1985

LEGION: ARCHOSAUROMORPHA HUENE, 1946

SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985

INFRALEGION: DINOSAURIFORMES NOVAS, 1992

SUPERCOHORT: DINOSAURIA OWEN, 1842

COHORT: THEROPODA EMEND MARSH, 1881

AVEPODA PAUL, 2002

SUPERORDER: NEOTHEROPODA BAKKER, 1986

GRANDORDER: CERATOSAURIA NOVAS, 1992

AVEROSTRA PAUL, 2001

ORDER: CERATOSAURIA MARSH, 1884

Superfamily: ABELISAUROIDEA Novas, 1989

Family: ABELISAURIDAE Bonaparte & Novas, 1985

Subfamily: CARNOTAURINAE Sereno, 1998

*Majungasaurus crenatissimus
= Megalosaurus crenatissimus

COHORT: ORNITHISCHIFORMES COOPER, 1985

SUPERORDER: SAUROPODOMORPHA HUENE, 1932

Incertae sedis

Gen. sp indet.


_____________________________________________________________________________________

 

MADAGASCAR

MAHAJANGA PROVINCE

MAHAJANGA BASIN

JUSE EAST OF BERIVOTRA VILLAGE

MAEVARENO SANDSTONE

CAMPANIAN STAGE?

SENONIAN SUBEPOCH

UPPER GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: MAMMALIA LINNAEUS, 1758

SUBCLASS: INCERTAE SEDIS

Gen. sp indet.

SUBCLASS: THERIIFORMES (ROWE, 1988) McKENNA & BELL, 1997

INFRACLASS: ALLOTHERIA (MARSH, 1880) McKENNA & BELL, 1997

ORDER: MULTITUBERCULATA COPE, 1884

Incertae sedis

Gen. sp indet.

SUBORDER: GONDWANATHERIA Mones, 1987

Family: SUDAMERICICIDAE Scillato-Yane & Prasad, 1984

Lavanify miolaka

_____________________________________________________________________________________

N. W. MADAGASCAR

MAHAJANGA PROVINCE

MAHAJANGA BASIN

MAEVARANO FORMATION

CAMPANIAN STAGE?

SENONIAN SUBEPOCH

UPPER GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1903

INFRACLASS: NEODIAPSIDA BENTON, 1985

LEGION: LEPIDOSAUROMORPHA BENTON, 1983

SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)

MAGNORDER: SQUAMATA, OPPEL, 1811

SUPERORDER: LACERTILIA OWEN, 1842

GRANDORDER: SCINCOGEKKONOMORPHA SUKHANOV, 1961

MICRORDER: SCINCOMORPHA CAMP, 1923

ORDER: LEPTOGLOSSA COPE, 1900

SUBORDER: ZONUROMORPHA ALIFANOV, 2000

INFRAORDER: ZONURIA FURBRINGER, 1900

Superfamily: CORDYLOIDEA Fritzinger, 1826

Family: CORDYLIDAE? Gray, 1837

Gen. sp indet.

_____________________________________________________________________________________

NORTH-WEST MADAGASCAR

MAHAJANGA PROVINCE

MAHAJANGA BASIN

APPROXIMATELY 1 KM SOUTH-EAST OF THE VILLAGE OF BERIVOTRA

MAEVARANO FORMATION

NEAR THE TOP OF THE CAMPANIAN STAGE?

SENONIAN SUBEPOCH

GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1903

SUPERLEGION: NEODIAPSIDA BENTON, 1985

LEGION: ARCHOSAUROMORPHA HUENE, 1946

SUPERCOHORT: CRUROTARSI SERENO & ARCUCCI, 1990

COHORT: CROCODYLOTARSI BENTON & CLARK, 1988

MAGNORDER: SUCHIA KREBS, 1974

SUPERORDER: CROCODYLOMORPHA WALKER, 1968

GRANDORDER: CROCODILIFORMES HAY, 1930

MICRORDER: MESOEUROCROCODYLIA WHETSTONE & WHYBROW, 1983

SUBORDER: MESOEUROCROCODYLIA WHETSTONE & WHYBROW, 1983

MESOEUROCRODYLIA: NEW FAMILES?

Also a long snouted form.

Incertae sedis

GONDWANASUCHIA DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004

PEIROSAURIMORPHA DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004

PEIROSAURIFORMES DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004

Superfamily: PEIROSAURIDAE Gasparini, 1982 emend de Souza Carvalho, Ribeiro & dos Santos Avilla, 2004

Family: PEIROSAURIDAE Gasparini, 1982

Subfamily: PEIROSAURIDAE Gasparini, 1982

Gen. nova?

 

NEOSUCHIA CLARK, IN BENTON & CLARK, 1988

Family: TREMATOCHAMPSIDAE Buffetaut, 1974

Trematochampsa oblita

_____________________________________________________________________________________

MADAGASCAR

MAJUNGA PROVINCE

46 KM S OF MAJUNGA

BETSIBOKA

“GRES DE MAEVARENO”

CAMPANIAN STAGE

SENONIAN SUBEPOCH

GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1903

SUPERLEGION: NEODIAPSIDA BENTON, 1985

LEGION: ARCHOSAUROMORPHA HUENE, 1946

SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985

INFRALEGION: DINOSAURIFORMES NOVAS, 1992

SUPERCOHORT: DINOSAURIA OWEN, 1842

COHORT: ORNITHISCHIFORMES COOPER, 1985

SUPERORDER: SAUROPODOMORPHA HUENE, 1932

GRANDORDER: BRONTOSAURIA OLSHEVSKY, 1991 (emend)

MICRORDER: SAUROPODA MARSH, 1878

ORDER: EUSAUROPODA

SUBORDER: NEOSAUROPODA BONAPARTE, 1986

INFRAORDER: MACRONARIA WILSON & SERENO, 1998

PARVORDER: TITANOSAURIFORMES SALGADO ET AL, 1997

Superfamily: TITANOSAUROIDEA Lydekker, 1885 (sensu Upchurch, 1999)

Family: TITANOSAURIDAE Lydekker, 1885

Incertae sedis

non Titanosaurus, T? madagascariensis
= Laplatasaurus madagascariensis

_____________________________________________________________________________________

MADAGASCAR

MAJUNGA PROVINCE

‘GRES DE MAEVANO’

PROBABLY CAMPANIAN STAGE

SENONIAN SUBEPOCH

GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1903

SUPERLEGION: NEODIAPSIDA BENTON, 1985

LEGION: ARCHOSAUROMORPHA HUENE, 1946

SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985

INFRALEGION: DINOSAURIFORMES NOVAS, 1992

SUPERCOHORT: DINOSAURIA OWEN, 1842

COHORT: THEROPODA EMEND MARSH, 1881

AVEPODA PAUL, 2002

SUPERORDER: NEOTHEROPODA BAKKER, 1986

GRANDORDER: CERATOSAURIA NOVAS, 1992

AVEROSTRA PAUL, 2001

ORDER: CERATOSAURIA MARSH, 1884

Superfamily: ABELISAUROIDEA Novas, 1989

Family: ABELISAURIDAE Bonaparte & Novas, 1985

Subfamily: MAJUNGASAURINAE Tortosa, Buffetaut, Vialle, Dutour, Turini, & Cheylan, 2014

*Majungatholus atopus


_____________________________________________________________________________________

MADAGASCAR

MAJUNGA PROVINCE

MAHAJANGA BASIN

CAMPANIAN STAGE?

SENONIAN SUBEPOCH

GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1903

SUPERLEGION: NEODIAPSIDA BENTON, 1985

LEGION: ARCHOSAUROMORPHA HUENE, 1946

SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985

INFRALEGION: DINOSAURIFORMES NOVAS, 1992

SUPERCOHORT: DINOSAURIA OWEN, 1842

COHORT: THEROPODA EMEND MARSH, 1881

AVEPODA PAUL, 2002

SUPERORDER: NEOTHEROPODA BAKKER, 1986

GRANDORDER: CERATOSAURIA NOVAS, 1992

AVEROSTRA PAUL, 2001

ORDER: CERATOSAURIA MARSH, 1884

Superfamily: ABELISAUROIDEA Novas, 1989

Family: ABELISAURIDAE Bonaparte & Novas, 1985

Subfamily: CARNOTAURINAE Sereno, 1998

Majungasaurus crenatissimus
= Megalosaurus crenatissimus

_____________________________________________________________________________________

MADAGASCAR

MAHAJANGA PROVINCE

MAHAJANGA BASIN

S TO THE RN4 AND BERIVOTRA VILLAGE

ANEMBALEMBA MEMBER

MAEVARANO FORMATION

MAASTRICHTIAN STAGE

UPPERMOST SENONIAN SUBEPOCH

UPPERMOST GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1904

SUPERLEGION: NEODIAPSIDA BENTON, 1985

LEGION: ARCHOSAUROMORPHA HUENE, 1946

SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985

INFRALEGION: DINOSAURIFORMES NOVAS, 1992

SUPERCOHORT: DINOSAURIA OWEN, 1842

COHORT: THEROPODA EMEND MARSH, 1881

AVEPODA PAUL, 2002

SUPERORDER: NEOTHEROPODA BAKKER, 1986

GRANDORDER: CERATOSAURIA NOVAS, 1992

AVEROSTRA PAUL, 2001

ORDER: CERATOSAURIA MARSH, 1884

Superfamily: ABELISAUROIDEA Bonaparte & Novas, 1985

Family: ABELISAURIDAE Bonaparte & Novas, 1985

Gen. sp indet.

_____________________________________________________________________________________

MADAGASCAR

MAHAJANGA PROVINCE

MAHAJANGA BASIN

ABOUT 500 M NE TO THE BERIVOTRA VILLAGE

LEFT BANK OF A SMALL STREAM "VAVARANONANTASHALVA"

ANEMBALEMBA MEMBER

MAEVARANO FORMATION

MAASTRICHTIAN STAGE

UPPERMOST SENONIAN SUBEPOCH

UPPERMOST GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1904

SUPERLEGION: NEODIAPSIDA BENTON, 1985

LEGION: ARCHOSAUROMORPHA HUENE, 1946

SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985

INFRALEGION: DINOSAURIFORMES NOVAS, 1992

SUPERCOHORT: DINOSAURIA OWEN, 1842

COHORT: THEROPODA EMEND MARSH, 1881

AVEPODA PAUL, 2002

SUPERORDER: NEOTHEROPODA BAKKER, 1986

GRANDORDER: CERATOSAURIA NOVAS, 1992

AVEROSTRA PAUL, 2001

ORDER: CERATOSAURIA MARSH, 1884

Superfamily: ABELISAUROIDEA Bonaparte & Novas, 1985

Family: ABELISAURIDAE Bonaparte & Novas, 1985

Gen. sp indet.

_____________________________________________________________________________________

MADAGASCAR

MAHAJANGA PROVINCE

MAHAJANGA BASIN

ANEMBALEMBA MEMBER

MAEVARANO FORMATION

MAASTRICHTIAN STAGE

SENONIAN SUBEPOCH

GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: AVES LINNAEUS, 1758

Incertae sedis

Gen. sp indet.

CLASS: MAMMALIA LINNAEUS, 1758

SUBCLASS: PROTOTHERIA GILL, 1872

INFRACLASS: TRECHNOTHERIA McKENNA, 1975

LEGION: CLADOTHERIA McKENNA, 1975

SUBLEGION: ZATHERIA MCKENNA, 1975

INFRALEGION: TRIBOSPHENIDA (McKENNA, 1975) McKENNA & BELL, 1997

SUPERCOHORT: THERIA (PARKER & HASWELL, 1987) McKENNA & BELL, 1997

COHORT: PLACENTALIA (OWEN, 1837) McKENNA & BELL, 1997

MAGNORDER: EPITHERIA (McKENNA, 1975) McKENNA & BELL, 1997

SUPERORDER: PREPTOTHERIA McKENNA, 1975

GRANDORDER: UNGULATOMORPHA ARCHIBALD, 1996

Family: ZHELESTIDAE Nesov, 1990

Gen. sp indet.

_____________________________________________________________________________________

 

MADAGASCAR

MAJUNGA PROVINCE

MAHAJANGA BASIN

MAEVARANO FORMATION

ANEMBALEMBA MEMBER

MAASTRICHTIAN STAGE

SENONIAN SUBEPOCH

GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1903

SUPERLEGION: NEODIAPSIDA BENTON, 1985

LEGION: ARCHOSAUROMORPHA HUENE, 1946

SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985

INFRALEGION: DINOSAURIFORMES NOVAS, 1992

SUPERCOHORT: DINOSAURIA OWEN, 1842

COHORT: ORNITHISCHIFORMES COOPER, 1985

SUPERORDER: SAUROPODOMORPHA HUENE, 1932

GRANDORDER: BRONTOSAURIA OLSHEVSKY, 1991 (emend)

MICRORDER: SAUROPODA MARSH, 1878

ORDER: EUSAUROPODA

SUBORDER: NEOSAUROPODA BONAPARTE, 1986

INFRAORDER: MACRONARIA WILSON & SERENO, 1998

PARVORDER: TITANOSAURIFORMES SALGADO ET AL, 1997

Superfamily: TITANOSAUROIDEA Lydekker, 1885 (sensu Upchurch, 1999)

Family: TITANOSAURIDAE Lydekker, 1885

Subfamily: SALTASAURINAE Powell, 1986 vide Bonaparte, 1987

Gen. sp indet.

 

_____________________________________________________________________________________

MADAGASCAR

MAJUNGA PROVINCE

MAHAJANGA BASIN

NEAR THE VILLAGE OF BERIVOTRA

MAEVARANO FORMATION

ANEMBALEMBA MEMBER

MAASTRICHTIAN STAGE

SENONIAN SUBEPOCH

GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1903

SUPERLEGION: NEODIAPSIDA BENTON, 1985

LEGION: ARCHOSAUROMORPHA HUENE, 1946

SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985

INFRALEGION: DINOSAURIFORMES NOVAS, 1992

SUPERCOHORT: DINOSAURIA OWEN, 1842

COHORT: THEROPODA EMEND MARSH, 1881

AVEPODA PAUL, 2002

SUPERORDER: NEOTHEROPODA BAKKER, 1986

GRANDORDER: CERATOSAURIA NOVAS, 1992

AVEROSTRA PAUL, 2001

ORDER: CERATOSAURIA MARSH, 1884

Superfamily: ABELISAUROIDEA Novas, 1989

Family: NOASAURIDAE Bonaparte & Powell, 1980

Subfamily: VELOCISAURIINAE Bonaparte, 1991 (emend AGNOLIN, NOVAS & APESTEGUIA, 2003)

*Masiakasaurus knopfleri

_____________________________________________________________________________________

NORTHEASTERN MADAGASCAR

MAHAJANA PROVINCE

MAHAJANGA BASIN

BERIVOTRA

ANEBALEMBA MEMBER

MAEVARANO FORMATION

MAASTRICHTIAN STAGE

UPPERMOST SENONIAN SUBEPOCH

UPPERMOST GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1903

SUPERLEGION: NEODIAPSIDA BENTON, 1985

LEGION: ARCHOSAUROMORPHA HUENE, 1946

SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985

INFRALEGION: DINOSAURIFORMES NOVAS, 1992

SUPERCOHORT: DINOSAURIA OWEN, 1842

COHORT: THEROPODA EMEND MARSH, 1881

AVEPODA PAUL, 2002

SUPERORDER: NEOTHEROPODA BAKKER, 1986

GRANDORDER: CERATOSAURIA NOVAS, 1992

AVEROSTRA PAUL, 2001

ORDER: CERATOSAURIA MARSH, 1884

Superfamily: ABELISAUROIDEA Novas, 1989

Family: ABELISAURIDAE Bonaparte & Novas, 1985

Subfamily: MAJUNGASAURINAE Tortosa, Buffetaut, Vialle, Dutour, Turini, & Cheylan, 2014

Majungatholus atopus

Family: NOASAURIDAE Bonaparte & Powell, 1980

Subfamily: VELOCISAURIINAE Bonaparte, 1991 (emend AGNOLIN, NOVAS & APESTEGUIA, 2003)

Masiakasaurus knopfleri

GRANDORDER: TETANURAE GAUTHIER, 1986

MICRORDER: COELUROSAURIA HUENE, 1914

ORDER: MANIRAPTORFORMES HOLTZ, 1996

AVETHEROPOA PAUL, 2002

AVEPECTORA PAUL, 2002

SUBORDER: MANIRAPTORA GAUTHIER, 1986

INFRAORDER: PARVES SERENO, 1997

PARVORDER: DEINONYCHOSAURIA COLBERT & D. A. RUSSELL, 1969

Family: DROMAEOSAURIDAE Russell, 1969

Gen. sp indet.

_____________________________________________________________________________________

MADAGASCAR

MAJUNGA PROVINCE

MAHAJANGA BASIN

UNSPECIFIED LOCALITY IN THE BERIVOTRA STUDY AREA

MAEVARANO FORMATION

ANEMBALEMBA MEMBER

MAASTRICHTIAN STAGE

SENONIAN SUBEPOCH

GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1903

SUPERLEGION: NEODIAPSIDA BENTON, 1985

LEGION: ARCHOSAUROMORPHA HUENE, 1946

SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985

INFRALEGION: DINOSAURIFORMES NOVAS, 1992

SUPERCOHORT: DINOSAURIA OWEN, 1842

COHORT: THEROPODA EMEND MARSH, 1881

AVEPODA PAUL, 2002

SUPERORDER: NEOTHEROPODA BAKKER, 1986

GRANDORDER: CERATOSAURIA NOVAS, 1992

AVEROSTRA PAUL, 2001

ORDER: CERATOSAURIA MARSH, 1884

Superfamily: ABELISAUROIDEA Novas, 1989

Family: ABELISAURIDAE Bonaparte & Novas, 1985

Subfamily: MAJUNGASAURINAE Tortosa, Buffetaut, Vialle, Dutour, Turini, & Cheylan, 2014

Majungatholus atopus


_____________________________________________________________________________________

MADAGASCAR

MAJUNGA PROVINCE

MAHAJANGA BASIN

UNSPECIFIED LOCALITY IN THE 'MAJUNGA DISTRICT'

MAEVARANO FORMATION

ANEMBALEMBA MEMBER

MAASTRICHTIAN STAGE

SENONIAN SUBEPOCH

GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1903

SUPERLEGION: NEODIAPSIDA BENTON, 1985

LEGION: ARCHOSAUROMORPHA HUENE, 1946

SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985

INFRALEGION: DINOSAURIFORMES NOVAS, 1992

SUPERCOHORT: DINOSAURIA OWEN, 1842

COHORT: THEROPODA EMEND MARSH, 1881

AVEPODA PAUL, 2002

SUPERORDER: NEOTHEROPODA BAKKER, 1986

GRANDORDER: CERATOSAURIA NOVAS, 1992

AVEROSTRA PAUL, 2001

ORDER: CERATOSAURIA MARSH, 1884

Superfamily: ABELISAUROIDEA Novas, 1989

Family: ABELISAURIDAE Bonaparte & Novas, 1985

Subfamily: MAJUNGASAURINAE Tortosa, Buffetaut, Vialle, Dutour, Turini, & Cheylan, 2014

Majungatholus atopus


_____________________________________________________________________________________

MADAGASCAR

MAHAJANGA BASIN

NORTH OF BERIVORTA

"GITE DU GUIDE"

MAEVARANO FORMATION

MAASTRICHTIAN STAGE

UPPER SENONIAN SUBEPOCH

UPPER UPPER GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1903

INFRACLASS: NEODIAPSIDA BENTON, 1985

LEGION: LEPIDOSAUROMORPHA BENTON, 1983

SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)

MAGNORDER: SQUAMATA, OPPEL, 1811

SUPERORDER: SERPENTES LINNAEUS, 1759

ORDER: OPHIDIA BRONGNIART, 1800

SUBORDER: ALETHINOPHIDIA NOPCSCA, 1923

Superfamily: BOOIDEA Gray, 1825

Subfamily: BOOIDINAE Gray, 1825

Infrafamily: BOOINIEA Gray, 1925

Tribe: MATSOINI Hoffstetter, 1961

*Madtsoia madagascariensis

 

_____________________________________________________________________________________

MADAGASCAR

MAJUNGA PROVINCE

MAHAJANGA BASIN

"EAST OF MAMMAL"

MAEVARANO FORMATION

ANEMBALEMBA MEMBER

MAASTRICHTIAN STAGE

SENONIAN SUBEPOCH

GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1903

SUPERLEGION: NEODIAPSIDA BENTON, 1985

LEGION: ARCHOSAUROMORPHA HUENE, 1946

SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985

INFRALEGION: DINOSAURIFORMES NOVAS, 1992

SUPERCOHORT: DINOSAURIA OWEN, 1842

COHORT: THEROPODA EMEND MARSH, 1881

AVEPODA PAUL, 2002

SUPERORDER: NEOTHEROPODA BAKKER, 1986

GRANDORDER: CERATOSAURIA NOVAS, 1992

AVEROSTRA PAUL, 2001

ORDER: CERATOSAURIA MARSH, 1884

Superfamily: ABELISAUROIDEA Novas, 1989

Family: NOASAURIDAE Bonaparte & Powell, 1980

Subfamily: VELOCISAURIINAE Bonaparte, 1991 (emend AGNOLIN, NOVAS & APESTEGUIA, 2003)

Masiakasaurus knopfleri

_____________________________________________________________________________________

 

MADAGASCAR

MAHAJANGA BASIN

SOUTH OF BERIVORTA

MNHN MAJ 8

MAEVARANO FORMATION

MAASTRICHTIAN STAGE

UPPER SENONIAN SUBEPOCH

UPPER UPPER GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1903

INFRACLASS: NEODIAPSIDA BENTON, 1985

LEGION: LEPIDOSAUROMORPHA BENTON, 1983

SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)

MAGNORDER: SQUAMATA, OPPEL, 1811

SUPERORDER: SERPENTES LINNAEUS, 1759

ORDER: OPHIDIA BRONGNIART, 1800

SUBORDER: ALETHINOPHIDIA NOPCSCA, 1923

Superfamily: BOOIDEA Gray, 1825

Subfamily: BOOIDINAE Gray, 1825

Infrafamily: BOOINIEA Gray, 1925

Tribe: MATSOINI Hoffstetter, 1961

Madtsoia madagascariensis

 

_____________________________________________________________________________________

MADAGASCAR

MAHAJANGA BASIN

SOUTH OF BERIVORTA

MNHN MAJ 9

MAEVARANO FORMATION

MAASTRICHTIAN STAGE

UPPER SENONIAN SUBEPOCH

UPPER UPPER GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1903

INFRACLASS: NEODIAPSIDA BENTON, 1985

LEGION: LEPIDOSAUROMORPHA BENTON, 1983

SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)

MAGNORDER: SQUAMATA, OPPEL, 1811

SUPERORDER: SERPENTES LINNAEUS, 1759

ORDER: OPHIDIA BRONGNIART, 1800

SUBORDER: ALETHINOPHIDIA NOPCSCA, 1923

Superfamily: BOOIDEA Gray, 1825

Subfamily: BOOIDINAE Gray, 1825

Infrafamily: BOOINIEA Gray, 1925

Tribe: MATSOINI Hoffstetter, 1961

Madtsoia madagascariensis

 

_____________________________________________________________________________________


MADAGASCAR

MAHAJANGA BASIN

SOUTH OF BERIVORTA

MNHN MAJ 10

MAEVARANO FORMATION

MAASTRICHTIAN STAGE

UPPER SENONIAN SUBEPOCH

UPPER UPPER GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1903

INFRACLASS: NEODIAPSIDA BENTON, 1985

LEGION: LEPIDOSAUROMORPHA BENTON, 1983

SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)

MAGNORDER: SQUAMATA, OPPEL, 1811

SUPERORDER: SERPENTES LINNAEUS, 1759

ORDER: OPHIDIA BRONGNIART, 1800

SUBORDER: ALETHINOPHIDIA NOPCSCA, 1923

Superfamily: BOOIDEA Gray, 1825

Subfamily: BOOIDINAE Gray, 1825

Infrafamily: BOOINIEA Gray, 1925

Tribe: MATSOINI Hoffstetter, 1961

Madtsoia madagascariensis

 

_____________________________________________________________________________________


MADAGASCAR

MAHAJANGA BASIN

NORTH OF MAHAJANGA-AMBALABE ROAD, BETWEEN KM 20 AND 25

MNHN MAJ 6

MAEVARANO FORMATION

MAASTRICHTIAN STAGE

UPPER SENONIAN SUBEPOCH

UPPER UPPER GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1903

INFRACLASS: NEODIAPSIDA BENTON, 1985

LEGION: LEPIDOSAUROMORPHA BENTON, 1983

SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)

MAGNORDER: SQUAMATA, OPPEL, 1811

SUPERORDER: SERPENTES LINNAEUS, 1759

ORDER: OPHIDIA BRONGNIART, 1800

SUBORDER: ALETHINOPHIDIA NOPCSCA, 1923

Superfamily: BOOIDEA Gray, 1825

Subfamily: BOOIDINAE Gray, 1825

Infrafamily: BOOINIEA Gray, 1925

Tribe: MATSOINI Hoffstetter, 1961

Madtsoia madagascariensis

 

_____________________________________________________________________________________

MADAGASCAR

MAHAJANGA BASIN

NORTH OF MAHAJANGA-AMBALABE ROAD, BETWEEN KM 20 AND 25

MNHN MAJ 7

MAEVARANO FORMATION

MAASTRICHTIAN STAGE

UPPER SENONIAN SUBEPOCH

UPPER UPPER GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1903

INFRACLASS: NEODIAPSIDA BENTON, 1985

LEGION: LEPIDOSAUROMORPHA BENTON, 1983

SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)

MAGNORDER: SQUAMATA, OPPEL, 1811

SUPERORDER: SERPENTES LINNAEUS, 1759

ORDER: OPHIDIA BRONGNIART, 1800

SUBORDER: ALETHINOPHIDIA NOPCSCA, 1923

Superfamily: BOOIDEA Gray, 1825

Subfamily: BOOIDINAE Gray, 1825

Infrafamily: BOOINIEA Gray, 1925

Tribe: MATSOINI Hoffstetter, 1961

Madtsoia madagascariensis

 

_____________________________________________________________________________________

MADAGASCAR

MAJUNGA PROVINCE

MAHAJANGA BASIN

MAD 01-03

MAEVARANO FORMATION

ANEMBALEMBA MEMBER

MAASTRICHTIAN STAGE

SENONIAN SUBEPOCH

GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1903

INFRACLASS: NEODIAPSIDA BENTON, 1985

LEGION: LEPIDOSAUROMORPHA BENTON, 1983

SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)

MAGNORDER: SQUAMATA, OPPEL, 1811

SUPERORDER: SERPENTES LINNAEUS, 1759

ORDER: OPHIDIA BRONGNIART, 1800

SUBORDER: ALETHINOPHIDIA NOPCSCA, 1923

Superfamily: BOOIDEA Gray, 1825

Subfamily: BOOIDINAE Gray, 1825

Infrafamily: BOOINIEA Gray, 1925

Tribe: MATSOINI Hoffstetter, 1961

Madtsoia madagascariensis

 

_____________________________________________________________________________________

MADAGASCAR

MAJUNGA PROVINCE

MAHAJANGA BASIN

MAD 01-05

MAEVARANO FORMATION

ANEMBALEMBA MEMBER

MAASTRICHTIAN STAGE

SENONIAN SUBEPOCH

GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1903

SUPERLEGION: NEODIAPSIDA BENTON, 1985

LEGION: ARCHOSAUROMORPHA HUENE, 1946

SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985

INFRALEGION: DINOSAURIFORMES NOVAS, 1992

SUPERCOHORT: DINOSAURIA OWEN, 1842

COHORT: THEROPODA EMEND MARSH, 1881

AVEPODA PAUL, 2002

SUPERORDER: NEOTHEROPODA BAKKER, 1986

GRANDORDER: CERATOSAURIA NOVAS, 1992

AVEROSTRA PAUL, 2001

ORDER: CERATOSAURIA MARSH, 1884

Superfamily: ABELISAUROIDEA Novas, 1989

Family: ABELISAURIDAE Bonaparte & Novas, 1985

Subfamily: CARNOTAURINAE Sereno, 1998

Majungatholus atopus


_____________________________________________________________________________________

MADAGASCAR

MAJUNGA PROVINCE

MAHAJANGA BASIN

MAD 03-03

MAEVARANO FORMATION

ANEMBALEMBA MEMBER

MAASTRICHTIAN STAGE

SENONIAN SUBEPOCH

GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1903

INFRACLASS: NEODIAPSIDA BENTON, 1985

LEGION: LEPIDOSAUROMORPHA BENTON, 1983

SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)

MAGNORDER: SQUAMATA, OPPEL, 1811

SUPERORDER: SERPENTES LINNAEUS, 1759

ORDER: OPHIDIA BRONGNIART, 1800

SUBORDER: ALETHINOPHIDIA NOPCSCA, 1923

Superfamily: BOOIDEA Gray, 1825

Subfamily: BOOIDINAE Gray, 1825

Infrafamily: BOOINIEA Gray, 1925

Tribe: MATSOINI Hoffstetter, 1961

Madtsoia madagascariensis

LEGION: ARCHOSAUROMORPHA HUENE, 1946

SUPERCOHORT: CRUROTARSI SERENO & ARCUCCI, 1990

COHORT: CROCODYLOTARSI BENTON & CLARK, 1988

MAGNORDER: SUCHIA KREBS, 1974

SUPERORDER: CROCODYLOMORPHA WALKER, 1968

GRANDORDER: CROCODILIFORMES HAY, 1930

MICRORDER: MESOEUROCROCODYLIA WHETSTONE & WHYBROW, 1983

ORDER: MESOSUCHIA EMEND HUXLEY, 1875

PARVORDER: METASUCHIA BENTON & CLARK, 1988

GONDWANASUCHIA DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004

NOTOSUCHIMORPHA DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004

NOTOSUCHIFORMES DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004

TERRIASUCHIA DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004

INFRAORDER: NOTOSUCHIA GASPARINI, 1971

Family: NOTOSUCHIDAE Dollo, 1924

Simosuchus clarki

SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985

INFRALEGION: DINOSAURIFORMES NOVAS, 1992

SUPERCOHORT: DINOSAURIA OWEN, 1842

COHORT: THEROPODA EMEND MARSH, 1881

AVEPODA PAUL, 2002

SUPERORDER: NEOTHEROPODA BAKKER, 1986

GRANDORDER: CERATOSAURIA NOVAS, 1992

AVEROSTRA PAUL, 2001

ORDER: CERATOSAURIA MARSH, 1884

Superfamily: ABELISAUROIDEA Novas, 1989

Family: NOASAURIDAE Bonaparte & Powell, 1980

Subfamily: VELOCISAURIINAE Bonaparte, 1991 (emend AGNOLIN, NOVAS & APESTEGUIA, 2003)

Masiakasaurus knopfleri

_____________________________________________________________________________________

MADAGASCAR

MAJUNGA PROVINCE

MAHAJANGA BASIN

MAD 03-04

MAEVARANO FORMATION

ANEMBALEMBA MEMBER

MAASTRICHTIAN STAGE

SENONIAN SUBEPOCH

GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1903

INFRACLASS: NEODIAPSIDA BENTON, 1985

LEGION: LEPIDOSAUROMORPHA BENTON, 1983

SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)

MAGNORDER: SQUAMATA, OPPEL, 1811

SUPERORDER: SERPENTES LINNAEUS, 1759

ORDER: OPHIDIA BRONGNIART, 1800

SUBORDER: ALETHINOPHIDIA NOPCSCA, 1923

Superfamily: BOOIDEA Gray, 1825

Subfamily: BOOIDINAE Gray, 1825

Infrafamily: BOOINIEA Gray, 1925

Tribe: MATSOINI Hoffstetter, 1961

Madtsoia madagascariensis

 

_____________________________________________________________________________________

MADAGASCAR

MAJUNGA PROVINCE

MAHAJANGA BASIN

MAD 03-05

MAEVARANO FORMATION

ANEMBALEMBA MEMBER

MAASTRICHTIAN STAGE

SENONIAN SUBEPOCH

GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: AMPHIBIA LINNAEUS, 1758

SUBCLASS: LISSAMPHIBIA, HAECKEL, 1866?

SUPERORDER: SALIENTIA LAURENTI, 1768

ORDER: ANURA RAFINESQUE, 1815

SUBORDER: NEOBATRACHIA REIG, 1958

Superfamily: HYLOIDEA Rafinesque, 1815 (sensu Wied, 1856)

Family: LEPTODACTYLIDAE Werner, 1896

Subfamily: CERATOPHYRINAE Tschudi, 1838

Beelzebufo ampinga

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1903

INFRACLASS: NEODIAPSIDA BENTON, 1985

LEGION: LEPIDOSAUROMORPHA BENTON, 1983

SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)

MAGNORDER: SQUAMATA, OPPEL, 1811

SUPERORDER: SERPENTES LINNAEUS, 1759

ORDER: OPHIDIA BRONGNIART, 1800

SUBORDER: ALETHINOPHIDIA NOPCSCA, 1923

Superfamily: BOOIDEA Gray, 1825

Subfamily: BOOIDINAE Gray, 1825

Infrafamily: BOOINIEA Gray, 1925

Tribe: MATSOINI Hoffstetter, 1961

Madtsoia madagascariensis

 

_____________________________________________________________________________________

MADAGASCAR

MAJUNGA PROVINCE

MAHAJANGA BASIN

MAD 03-09

MAEVARANO FORMATION

ANEMBALEMBA MEMBER

MAASTRICHTIAN STAGE

SENONIAN SUBEPOCH

GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1903

INFRACLASS: NEODIAPSIDA BENTON, 1985

LEGION: LEPIDOSAUROMORPHA BENTON, 1983

SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)

MAGNORDER: SQUAMATA, OPPEL, 1811

SUPERORDER: SERPENTES LINNAEUS, 1759

ORDER: OPHIDIA BRONGNIART, 1800

SUBORDER: ALETHINOPHIDIA NOPCSCA, 1923

Superfamily: BOOIDEA Gray, 1825

Subfamily: BOOIDINAE Gray, 1825

Infrafamily: BOOINIEA Gray, 1925

Tribe: MATSOINI Hoffstetter, 1961

Madtsoia madagascariensis

LEGION: ARCHOSAUROMORPHA HUENE, 1946

SUPERCOHORT: CRUROTARSI SERENO & ARCUCCI, 1990

COHORT: CROCODYLOTARSI BENTON & CLARK, 1988

MAGNORDER: SUCHIA KREBS, 1974

SUPERORDER: CROCODYLOMORPHA WALKER, 1968

GRANDORDER: CROCODILIFORMES HAY, 1930

MICRORDER: MESOEUROCROCODYLIA WHETSTONE & WHYBROW, 1983

ORDER: MESOSUCHIA EMEND HUXLEY, 1875

PARVORDER: METASUCHIA BENTON & CLARK, 1988

GONDWANASUCHIA DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004

NOTOSUCHIMORPHA DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004

NOTOSUCHIFORMES DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004

TERRIASUCHIA DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004

INFRAORDER: NOTOSUCHIA GASPARINI, 1971

Family: NOTOSUCHIDAE Dollo, 1924

Simosuchus clarki

_____________________________________________________________________________________

MADAGASCAR

MAJUNGA PROVINCE

MAHAJANGA BASIN

MAD 03-10

MAEVARANO FORMATION

ANEMBALEMBA MEMBER

MAASTRICHTIAN STAGE

SENONIAN SUBEPOCH

GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: AMPHIBIA LINNAEUS, 1758

SUBCLASS: LISSAMPHIBIA, HAECKEL, 1866?

SUPERORDER: SALIENTIA LAURENTI, 1768

ORDER: ANURA RAFINESQUE, 1815

SUBORDER: NEOBATRACHIA REIG, 1958

Superfamily: HYLOIDEA Rafinesque, 1815 (sensu Wied, 1856)

Family: LEPTODACTYLIDAE Werner, 1896

Subfamily: CERATOPHYRINAE Tschudi, 1838

Beelzebufo ampinga

_____________________________________________________________________________________

MADAGASCAR

MAJUNGA PROVINCE

MAHAJANGA BASIN

MASIAKAKOHO STUDY AREA

MAD 03-17

MAEVARANO FORMATION

ANEMBALEMBA MEMBER

MAASTRICHTIAN STAGE

SENONIAN SUBEPOCH

GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1903

LEGION: ARCHOSAUROMORPHA HUENE, 1946

SUPERCOHORT: CRUROTARSI SERENO & ARCUCCI, 1990

COHORT: CROCODYLOTARSI BENTON & CLARK, 1988

MAGNORDER: SUCHIA KREBS, 1974

SUPERORDER: CROCODYLOMORPHA WALKER, 1968

GRANDORDER: CROCODILIFORMES HAY, 1930

MICRORDER: MESOEUROCROCODYLIA WHETSTONE & WHYBROW, 1983

ORDER: MESOSUCHIA EMEND HUXLEY, 1875

PARVORDER: METASUCHIA BENTON & CLARK, 1988

GONDWANASUCHIA DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004

NOTOSUCHIMORPHA DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004

NOTOSUCHIFORMES DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004

TERRIASUCHIA DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004

INFRAORDER: NOTOSUCHIA GASPARINI, 1971

Family: NOTOSUCHIDAE Dollo, 1924

Simosuchus clarki

_____________________________________________________________________________________

MADAGASCAR

MAJUNGA PROVINCE

MAHAJANGA BASIN

MAD 03-18

MAEVARANO FORMATION

ANEMBALEMBA MEMBER

MAASTRICHTIAN STAGE

SENONIAN SUBEPOCH

GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: AMPHIBIA LINNAEUS, 1758

SUBCLASS: LISSAMPHIBIA, HAECKEL, 1866?

SUPERORDER: SALIENTIA LAURENTI, 1768

ORDER: ANURA RAFINESQUE, 1815

SUBORDER: NEOBATRACHIA REIG, 1958

Superfamily: HYLOIDEA Rafinesque, 1815 (sensu Wied, 1856)

Family: LEPTODACTYLIDAE Werner, 1896

Subfamily: CERATOPHYRINAE Tschudi, 1838

Beelzebufo ampinga

_____________________________________________________________________________________

MADAGASCAR

MAJUNGA PROVINCE

MAHAJANGA BASIN

MAD 03-18

MAEVARANO FORMATION

ANEMBALEMBA MEMBER

MAASTRICHTIAN STAGE

SENONIAN SUBEPOCH

GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: AMPHIBIA LINNAEUS, 1758

SUBCLASS: LISSAMPHIBIA, HAECKEL, 1866?

SUPERORDER: SALIENTIA LAURENTI, 1768

ORDER: ANURA RAFINESQUE, 1815

SUBORDER: NEOBATRACHIA REIG, 1958

Superfamily: HYLOIDEA Rafinesque, 1815 (sensu Wied, 1856)

Family: LEPTODACTYLIDAE Werner, 1896

Subfamily: CERATOPHYRINAE Tschudi, 1838

Beelzebufo ampinga

_____________________________________________________________________________________

MADAGASCAR

MAJUNGA PROVINCE

MAHAJANGA BASIN

MAD 03-40

MAEVARANO FORMATION

ANEMBALEMBA MEMBER

MAASTRICHTIAN STAGE

SENONIAN SUBEPOCH

GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1903

SUPERLEGION: NEODIAPSIDA BENTON, 1985

LEGION: ARCHOSAUROMORPHA HUENE, 1946

SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985

INFRALEGION: DINOSAURIFORMES NOVAS, 1992

SUPERCOHORT: DINOSAURIA OWEN, 1842

COHORT: THEROPODA EMEND MARSH, 1881

AVEPODA PAUL, 2002

SUPERORDER: NEOTHEROPODA BAKKER, 1986

GRANDORDER: CERATOSAURIA NOVAS, 1992

AVEROSTRA PAUL, 2001

ORDER: CERATOSAURIA MARSH, 1884

Superfamily: ABELISAUROIDEA Novas, 1989

Family: NOASAURIDAE Bonaparte & Powell, 1980

Subfamily: VELOCISAURIINAE Bonaparte, 1991 (emend AGNOLIN, NOVAS & APESTEGUIA, 2003)

Masiakasaurus knopfleri

_____________________________________________________________________________________

MADAGASCAR

MAJUNGA PROVINCE

MAHAJANGA BASIN

MAD 03-66

MAEVARANO FORMATION

ANEMBALEMBA MEMBER

MAASTRICHTIAN STAGE

SENONIAN SUBEPOCH

GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1903

SUPERLEGION: NEODIAPSIDA BENTON, 1985

LEGION: ARCHOSAUROMORPHA HUENE, 1946

SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985

INFRALEGION: DINOSAURIFORMES NOVAS, 1992

SUPERCOHORT: DINOSAURIA OWEN, 1842

COHORT: THEROPODA EMEND MARSH, 1881

AVEPODA PAUL, 2002

SUPERORDER: NEOTHEROPODA BAKKER, 1986

GRANDORDER: CERATOSAURIA NOVAS, 1992

AVEROSTRA PAUL, 2001

ORDER: CERATOSAURIA MARSH, 1884

Superfamily: ABELISAUROIDEA Novas, 1989

Family: NOASAURIDAE Bonaparte & Powell, 1980

Subfamily: VELOCISAURIINAE Bonaparte, 1991 (emend AGNOLIN, NOVAS & APESTEGUIA, 2003)

Masiakasaurus knopfleri

_____________________________________________________________________________________

MADAGASCAR

MAJUNGA PROVINCE

MAHAJANGA BASIN

MAD 05-04

MAEVARANO FORMATION

ANEMBALEMBA MEMBER

MAASTRICHTIAN STAGE

SENONIAN SUBEPOCH

GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: CHELONOMORPHA KUHN, 1960

SUPERCOHORT: TESTUDINES LINNAEUS, 1758

COHORT: CASICHELYDIA emend GAFFNEY & MEYLAN, 1988

MAGNORDER: PLEURODIRA GAFFNEY & MEYLAN, 1988

ORDER: PLEURODIRIDA COPE, 1864

SUBORDER: EUPLEURODIRA GAFFNEY & MEYLAN, 1988

Superfamily: PELOMEDUSOIDES Cope, 1868 Cope, 1868

Superfamily: BOTHREMYDINURA Baur, 1891

Epifamily: BOTHERMYDINURA Baur, 1891

Family: BOTHREMYDIDAE Baur, 1891

Subfamily: BOTHREMYDINAE Baur, 1891

Infrafamily: BOTHREMYDODDA Baur, 1891

Tribe: KURMADEMYDINI Gaffney, Tong & Meylan, 2006

Kinkonychelys rogersi

SUBCLASS: DIAPSIDA OSBORN, 1903

INFRACLASS: NEODIAPSIDA BENTON, 1985

LEGION: LEPIDOSAUROMORPHA BENTON, 1983

SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)

MAGNORDER: SQUAMATA, OPPEL, 1811

SUPERORDER: SERPENTES LINNAEUS, 1759

ORDER: OPHIDIA BRONGNIART, 1800

SUBORDER: ALETHINOPHIDIA NOPCSCA, 1923

Superfamily: BOOIDEA Gray, 1825

Family: BOIDAE Gray, 1825

Subfamily: BOOIDINAE Gray, 1825

Infrafamily: BOOINIEA Gray, 1925

Tribe: MATSOINI Hoffstetter, 1961

Menarana nosymena

LEGION: ARCHOSAUROMORPHA HUENE, 1946

SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985

INFRALEGION: DINOSAURIFORMES NOVAS, 1992

SUPERCOHORT: DINOSAURIA OWEN, 1842

COHORT: THEROPODA EMEND MARSH, 1881

AVEPODA PAUL, 2002

SUPERORDER: NEOTHEROPODA BAKKER, 1986

GRANDORDER: CERATOSAURIA NOVAS, 1992

AVEROSTRA PAUL, 2001

ORDER: CERATOSAURIA MARSH, 1884

Superfamily: ABELISAUROIDEA Novas, 1989

Family: NOASAURIDAE Bonaparte & Powell, 1980

Subfamily: VELOCISAURIINAE Bonaparte, 1991 (emend AGNOLIN, NOVAS & APESTEGUIA, 2003)

Masiakasaurus knopfleri

_____________________________________________________________________________________

MADAGASCAR

MAJUNGA PROVINCE

MAHAJANGA BASIN

MAD 05-36

MAEVARANO FORMATION

ANEMBALEMBA MEMBER

MAASTRICHTIAN STAGE

SENONIAN SUBEPOCH

GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1903

LEGION: ARCHOSAUROMORPHA HUENE, 1946

SUPERCOHORT: CRUROTARSI SERENO & ARCUCCI, 1990

COHORT: CROCODYLOTARSI BENTON & CLARK, 1988

MAGNORDER: SUCHIA KREBS, 1974

SUPERORDER: CROCODYLOMORPHA WALKER, 1968

GRANDORDER: CROCODILIFORMES HAY, 1930

MICRORDER: MESOEUROCROCODYLIA WHETSTONE & WHYBROW, 1983

ORDER: MESOSUCHIA EMEND HUXLEY, 1875

PARVORDER: METASUCHIA BENTON & CLARK, 1988

GONDWANASUCHIA DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004

NOTOSUCHIMORPHA DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004

NOTOSUCHIFORMES DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004

TERRIASUCHIA DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004

INFRAORDER: NOTOSUCHIA GASPARINI, 1971

Family: NOTOSUCHIDAE Dollo, 1924

Simosuchus clarki

_____________________________________________________________________________________

MADAGASCAR

MAJUNGA PROVINCE

MAHAJANGA BASIN

MAD 05-37

MAEVARANO FORMATION

ANEMBALEMBA MEMBER

MAASTRICHTIAN STAGE

SENONIAN SUBEPOCH

GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1903

LEGION: ARCHOSAUROMORPHA HUENE, 1946

SUPERCOHORT: CRUROTARSI SERENO & ARCUCCI, 1990

COHORT: CROCODYLOTARSI BENTON & CLARK, 1988

MAGNORDER: SUCHIA KREBS, 1974

SUPERORDER: CROCODYLOMORPHA WALKER, 1968

GRANDORDER: CROCODILIFORMES HAY, 1930

MICRORDER: MESOEUROCROCODYLIA WHETSTONE & WHYBROW, 1983

ORDER: MESOSUCHIA EMEND HUXLEY, 1875

PARVORDER: METASUCHIA BENTON & CLARK, 1988

GONDWANASUCHIA DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004

NOTOSUCHIMORPHA DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004

NOTOSUCHIFORMES DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004

TERRIASUCHIA DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004

INFRAORDER: NOTOSUCHIA GASPARINI, 1971

Family: NOTOSUCHIDAE Dollo, 1924

Simosuchus clarki

_____________________________________________________________________________________

MADAGASCAR

MAJUNGA PROVINCE

MAHAJANGA BASIN

BOENY REGION

MAD 05-38

MAEVARANO FORMATION

ANEMBALEMBA MEMBER

MAASTRICHTIAN STAGE

SENONIAN SUBEPOCH

GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: CHELONOMORPHA KUHN, 1960

SUPERCOHORT: TESTUDINES LINNAEUS, 1758

COHORT: CASICHELYDIA emend GAFFNEY & MEYLAN, 1988

MAGNORDER: PLEURODIRA GAFFNEY & MEYLAN, 1988

ORDER: PLEURODIRIDA COPE, 1864

SUBORDER: EUPLEURODIRA GAFFNEY & MEYLAN, 1988

Superfamily: PELOMEDUSOIDES Cope, 1868 Cope, 1868

Hyperfamily: PODOCNEMIDERA Cope, 1868

CLADE: PAN-PODOCNEMIDIDAE

Family: SAHONACHELYIDAE Joyce, Rollot, Evers, Lyson, Rahantariosa & Krause, 2021

*Sahonachelys mailakavava

_____________________________________________________________________________________

MADAGASCAR

MAJUNGA PROVINCE

MAHAJANGA BASIN

MAD 05-42

MAEVARANO FORMATION

ANEMBALEMBA MEMBER

MAASTRICHTIAN STAGE

SENONIAN SUBEPOCH

GULF EPOCH

LATE CRETACEOUS

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1903

LEGION: ARCHOSAUROMORPHA HUENE, 1946

SUPERCOHORT: CRUROTARSI SERENO & ARCUCCI, 1990

COHORT: CROCODYLOTARSI BENTON & CLARK, 1988

MAGNORDER: SUCHIA KREBS, 1974

SUPERORDER: CROCODYLOMORPHA WALKER, 1968

GRANDORDER: CROCODILIFORMES HAY, 1930

MICRORDER: MESOEUROCROCODYLIA WHETSTONE & WHYBROW, 1983

ORDER: MESOSUCHIA EMEND HUXLEY, 1875

PARVORDER: METASUCHIA BENTON & CLARK, 1988

GONDWANASUCHIA DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004

NOTOSUCHIMORPHA DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004

NOTOSUCHIFORMES DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004

TERRIASUCHIA DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004

INFRAORDER: NOTOSUCHIA GASPARINI, 1971

Family: NOTOSUCHIDAE Dollo, 1924

Simosuchus clarki

SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985

INFRALEGION: DINOSAURIFORMES NOVAS, 1992

SUPERCOHORT: DINOSAURIA OWEN, 1842

COHORT: THEROPODA EMEND MARSH, 1881

AVEPODA PAUL, 2002

SUPERORDER: NEOTHEROPODA BAKKER, 1986

GRANDORDER: CERATOSAURIA NOVAS, 1992

AVEROSTRA PAUL, 2001

ORDER: CERATOSAURIA MARSH, 1884

Superfamily: ABELISAUROIDEA Novas, 1989

Family: ABELISAURIDAE Bonaparte & Novas, 1985

Subfamily: CARNOTAURINAE Sereno, 1998

Majungatholus atopus

Family: NOASAURIDAE Bonaparte & Powell, 1980

Subfamily: VELOCISAURIINAE Bonaparte, 1991 (emend AGNOLIN, NOVAS & APESTEGUIA, 2003)

Masiakasaurus knopfleri

CLASS: AVES LINNAEUS, 1758

Incertae sedis

Gen.sp indet.

SUBCLASS: SAURORNITHES NICHOLSON, 1879

AVEBREVICAUDA PAUL, 2001

INFRACLASS: ORNITHOTHORACES NOVA

Incertae sedis

Gen.sp indet.

LEGION: ORNITHURAE NOVA

COHORT: ENANTIORNITHES NOVA

Incertae sedis

*Falcatakely forsterae

Gen.sp indet.

SUPERORDER: EUENANTIORNITHES CHIAPPE, 2002

ORDER: EUORNITHIFORMES KUROCHKIN, 1996

Family: CONCORNITHIDAE Kurochkin, 1996

Vorona berivotrensis

_____________________________________________________________________________________

MADAGASCAR

MAJUNGA PROVINCE

MAHAJANGA BASIN

MASIAKAKOHO STUDY AREA

MAD 05-59

MAEVARANO FORMATION

ANEMBALEMBA MEMBER

MAASTRICHTIAN STAGE

SENONIAN SUBEPOCH

GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1903

LEGION: ARCHOSAUROMORPHA HUENE, 1946

SUPERCOHORT: CRUROTARSI SERENO & ARCUCCI, 1990

COHORT: CROCODYLOTARSI BENTON & CLARK, 1988

MAGNORDER: SUCHIA KREBS, 1974

SUPERORDER: CROCODYLOMORPHA WALKER, 1968

GRANDORDER: CROCODILIFORMES HAY, 1930

MICRORDER: MESOEUROCROCODYLIA WHETSTONE & WHYBROW, 1983

ORDER: MESOSUCHIA EMEND HUXLEY, 1875

PARVORDER: METASUCHIA BENTON & CLARK, 1988

GONDWANASUCHIA DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004

NOTOSUCHIMORPHA DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004

NOTOSUCHIFORMES DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004

TERRIASUCHIA DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004

INFRAORDER: NOTOSUCHIA GASPARINI, 1971

Family: NOTOSUCHIDAE Dollo, 1924

Simosuchus clarki

_____________________________________________________________________________________

MADAGASCAR

MAJUNGA PROVINCE

MAHAJANGA BASIN

MASIAKAKOHO STUDY AREA

MAD 05-61

MAEVARANO FORMATION

MAASTRICHTIAN STAGE

SENONIAN SUBEPOCH

GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: CHELONOMORPHA KUHN, 1960

SUPERCOHORT: TESTUDINES LINNAEUS, 1758

COHORT: CASICHELYDIA emend GAFFNEY & MEYLAN, 1988

MAGNORDER: PLEURODIRA GAFFNEY & MEYLAN, 1988

ORDER: PLEURODIRIDA COPE, 1864

SUBORDER: EUPLEURODIRA GAFFNEY & MEYLAN, 1988

Superfamily: PELOMEDUSOIDES Cope, 1868 Cope, 1868

Hyperfamily: PODOCNEMIDERA Cope, 1868

CLADE: PAN-PODOCNEMIDIDAE

Family: SAHONACHELYIDAE Joyce, Rollot, Evers, Lyson, Rahantariosa & Krause, 2021

Sokatra antitra

_____________________________________________________________________________________

MADAGASCAR

MAJUNGA PROVINCE

MAHAJANGA BASIN

MAD 05-64

MAEVARANO FORMATION

ANEMBALEMBA MEMBER

MAASTRICHTIAN STAGE

SENONIAN SUBEPOCH

GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: AMPHIBIA LINNAEUS, 1758

SUBCLASS: LISSAMPHIBIA, HAECKEL, 1866?

SUPERORDER: SALIENTIA LAURENTI, 1768

ORDER: ANURA RAFINESQUE, 1815

SUBORDER: NEOBATRACHIA REIG, 1958

Superfamily: HYLOIDEA Rafinesque, 1815 (sensu Wied, 1856)

Family: LEPTODACTYLIDAE Werner, 1896

Subfamily: CERATOPHYRINAE Tschudi, 1838

Beelzebufo ampinga

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1903

INFRACLASS: NEODIAPSIDA BENTON, 1985

LEGION: LEPIDOSAUROMORPHA BENTON, 1983

SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)

MAGNORDER: SQUAMATA, OPPEL, 1811

SUPERORDER: SERPENTES LINNAEUS, 1759

ORDER: OPHIDIA BRONGNIART, 1800

SUBORDER: ALETHINOPHIDIA NOPCSCA, 1923

Superfamily: BOOIDEA Gray, 1825

Subfamily: BOOIDINAE Gray, 1825

Infrafamily: BOOINIEA Gray, 1925

Tribe: MATSOINI Hoffstetter, 1961

Madtsoia madagascariensis

 

_____________________________________________________________________________________

MADAGASCAR

MAJUNGA PROVINCE

MAHAJANGA BASIN

MAD 07-15

MAEVARANO FORMATION

ANEMBALEMBA MEMBER

MAASTRICHTIAN STAGE

SENONIAN SUBEPOCH

GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: AMPHIBIA LINNAEUS, 1758

SUBCLASS: LISSAMPHIBIA, HAECKEL, 1866?

SUPERORDER: SALIENTIA LAURENTI, 1768

ORDER: ANURA RAFINESQUE, 1815

SUBORDER: NEOBATRACHIA REIG, 1958

Superfamily: HYLOIDEA Rafinesque, 1815 (sensu Wied, 1856)

Family: LEPTODACTYLIDAE Werner, 1896

Subfamily: CERATOPHYRINAE Tschudi, 1838

Beelzebufo ampinga

_____________________________________________________________________________________

MADAGASCAR

MAJUNGA PROVINCE

MAHAJANGA BASIN

MAD 07-16

MAEVARANO FORMATION

ANEMBALEMBA MEMBER

MAASTRICHTIAN STAGE

SENONIAN SUBEPOCH

GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1903

SUPERLEGION: NEODIAPSIDA BENTON, 1985

LEGION: ARCHOSAUROMORPHA HUENE, 1946

SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985

INFRALEGION: DINOSAURIFORMES NOVAS, 1992

SUPERCOHORT: DINOSAURIA OWEN, 1842

COHORT: THEROPODA EMEND MARSH, 1881

AVEPODA PAUL, 2002

SUPERORDER: NEOTHEROPODA BAKKER, 1986

GRANDORDER: CERATOSAURIA NOVAS, 1992

AVEROSTRA PAUL, 2001

ORDER: CERATOSAURIA MARSH, 1884

Superfamily: ABELISAUROIDEA Novas, 1989

Family: NOASAURIDAE Bonaparte & Powell, 1980

Subfamily: VELOCISAURIINAE Bonaparte, 1991 (emend AGNOLIN, NOVAS & APESTEGUIA, 2003)

Masiakasaurus knopfleri

_____________________________________________________________________________________

MADAGASCAR

MAJUNGA PROVINCE

MAHAJANGA BASIN

MAD 07-17

MAEVARANO FORMATION

ANEMBALEMBA MEMBER

MAASTRICHTIAN STAGE

SENONIAN SUBEPOCH

GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1903

SUPERLEGION: NEODIAPSIDA BENTON, 1985

LEGION: ARCHOSAUROMORPHA HUENE, 1946

SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985

INFRALEGION: DINOSAURIFORMES NOVAS, 1992

SUPERCOHORT: DINOSAURIA OWEN, 1842

COHORT: THEROPODA EMEND MARSH, 1881

AVEPODA PAUL, 2002

SUPERORDER: NEOTHEROPODA BAKKER, 1986

GRANDORDER: CERATOSAURIA NOVAS, 1992

AVEROSTRA PAUL, 2001

ORDER: CERATOSAURIA MARSH, 1884

Superfamily: ABELISAUROIDEA Novas, 1989

Family: NOASAURIDAE Bonaparte & Powell, 1980

Subfamily: VELOCISAURIINAE Bonaparte, 1991 (emend AGNOLIN, NOVAS & APESTEGUIA, 2003)

Masiakasaurus knopfleri

_____________________________________________________________________________________

MADAGASCAR

MAJUNGA PROVINCE

MAHAJANGA BASIN

MAD 07-20

MAEVARANO FORMATION

ANEMBALEMBA MEMBER

MAASTRICHTIAN STAGE

SENONIAN SUBEPOCH

GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: AMPHIBIA LINNAEUS, 1758

SUBCLASS: LISSAMPHIBIA, HAECKEL, 1866?

SUPERORDER: SALIENTIA LAURENTI, 1768

ORDER: ANURA RAFINESQUE, 1815

SUBORDER: NEOBATRACHIA REIG, 1958

Superfamily: HYLOIDEA Rafinesque, 1815 (sensu Wied, 1856)

Family: LEPTODACTYLIDAE Werner, 1896

Subfamily: CERATOPHYRINAE Tschudi, 1838

Beelzebufo ampinga

_____________________________________________________________________________________

NORTHEASTERN MADAGASCAR

MAHAJANA PROVINCE

MAHAJANGA BASIN

APPROXIMATELY 2 KM NORTH OF THE WESTERN END OF LAC KINKONY AND 4.9 KM NORTHEAST OF THE VILLAGE OF ANTONGOMENA

LAC KINKONY STUDY AREA

MAD 07-25

UNNAMED MEMBER

MAEVARANO FORMATION

MAASTRICHTIAN STAGE

SENONIAN SUBEPOCH

GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: CHELONOMORPHA KUHN, 1960

SUPERCOHORT: TESTUDINES LINNAEUS, 1758

COHORT: CASICHELYDIA emend GAFFNEY & MEYLAN, 1988

MAGNORDER: PLEURODIRA GAFFNEY & MEYLAN, 1988

ORDER: PLEURODIRIDA COPE, 1864

SUBORDER: EUPLEURODIRA GAFFNEY & MEYLAN, 1988

Superfamily: PELOMEDUSOIDES Cope, 1868 Cope, 1868

Superfamily: BOTHREMYDINURA Baur, 1891

Epifamily: BOTHERMYDINURA Baur, 1891

Family: BOTHREMYDIDAE Baur, 1891

Subfamily: BOTHREMYDINAE Baur, 1891

Infrafamily: BOTHREMYDODDA Baur, 1891

Tribe: KURMADEMYDINI Gaffney, Tong & Meylan, 2006

*Kinkonychelys rogersi

_____________________________________________________________________________________

MADAGASCAR

MAJUNGA PROVINCE

MAHAJANGA BASIN

BERIVOTRA STUDY AREA

MAD 07-66

MAEVARANO FORMATION

MAASTRICHTIAN STAGE

SENONIAN SUBEPOCH

GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: CHELONOMORPHA KUHN, 1960

SUPERCOHORT: TESTUDINES LINNAEUS, 1758

COHORT: CASICHELYDIA emend GAFFNEY & MEYLAN, 1988

MAGNORDER: PLEURODIRA GAFFNEY & MEYLAN, 1988

ORDER: PLEURODIRIDA COPE, 1864

SUBORDER: EUPLEURODIRA GAFFNEY & MEYLAN, 1988

Superfamily: PELOMEDUSOIDES Cope, 1868 Cope, 1868

Hyperfamily: PODOCNEMIDERA Cope, 1868

CLADE: PAN-PODOCNEMIDIDAE

Family: SAHONACHELYIDAE Joyce, Rollot, Evers, Lyson, Rahantariosa & Krause, 2021

Sokatra antitra

_____________________________________________________________________________________

MADAGASCAR

MAJUNGA PROVINCE

MAHAJANGA BASIN

MAD 10-13

MAEVARANO FORMATION

ANEMBALEMBA MEMBER

MAASTRICHTIAN STAGE

SENONIAN SUBEPOCH

GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: AMPHIBIA LINNAEUS, 1758

SUBCLASS: LISSAMPHIBIA, HAECKEL, 1866?

SUPERORDER: SALIENTIA LAURENTI, 1768

ORDER: ANURA RAFINESQUE, 1815

SUBORDER: NEOBATRACHIA REIG, 1958

Superfamily: HYLOIDEA Rafinesque, 1815 (sensu Wied, 1856)

Family: LEPTODACTYLIDAE Werner, 1896

Subfamily: CERATOPHYRINAE Tschudi, 1838

Beelzebufo ampinga

_____________________________________________________________________________________

MADAGASCAR

MAJUNGA PROVINCE

MAHAJANGA BASIN

MAD 10-24

MAEVARANO FORMATION

LAC KINKOY MEMBER

MAASTRICHTIAN STAGE

SENONIAN SUBEPOCH

GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: AMPHIBIA LINNAEUS, 1758

SUBCLASS: LISSAMPHIBIA, HAECKEL, 1866?

SUPERORDER: SALIENTIA LAURENTI, 1768

ORDER: ANURA RAFINESQUE, 1815

SUBORDER: NEOBATRACHIA REIG, 1958

Superfamily: HYLOIDEA Rafinesque, 1815 (sensu Wied, 1856)

Family: LEPTODACTYLIDAE Werner, 1896

Subfamily: CERATOPHYRINAE Tschudi, 1838

Beelzebufo ampinga

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1903

INFRACLASS: NEODIAPSIDA BENTON, 1985

LEGION: LEPIDOSAUROMORPHA BENTON, 1983

SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)

MAGNORDER: SQUAMATA, OPPEL, 1811

SUPERORDER: SERPENTES LINNAEUS, 1759

Incertae sedis

Gen. sp indet

ORDER: OPHIDIA BRONGNIART, 1800

SUBORDER: ALETHINOPHIDIA NOPCSCA, 1923

Superfamily: BOOIDEA Gray, 1825

Family: ACROCHORDOIDAE Bonaparte, 1838

Subfamily: ?NIGEROPHEINAE Rage, 1975

*Indophis fanambinana

CLASS: MAMMALIA LINNAEUS, 1758

SUBCLASS: THERIIFORMES (ROWE, 1988) McKENNA & BELL, 1997

INFRACLASS: ALLOTHERIA (MARSH, 1880) McKENNA & BELL, 1997

ORDER: MULTITUBERCULATA COPE, 1884

SUBORDER: GONDWANATHERIA Mones, 1987

Family: SUDAMERICICIDAE Scillato-Yane & Prasad, 1984

*Vintana sertichi

_____________________________________________________________________________________

MADAGASCAR

MAJUNGA PROVINCE

MAHAJANGA BASIN

MAD 93-01

MAEVARANO FORMATION

ANEMBALEMBA MEMBER

MAASTRICHTIAN STAGE

SENONIAN SUBEPOCH

GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: AMPHIBIA LINNAEUS, 1758

SUBCLASS: LISSAMPHIBIA, HAECKEL, 1866?

SUPERORDER: SALIENTIA LAURENTI, 1768

ORDER: ANURA RAFINESQUE, 1815

SUBORDER: NEOBATRACHIA REIG, 1958

Superfamily: HYLOIDEA Rafinesque, 1815 (sensu Wied, 1856)

Family: LEPTODACTYLIDAE Werner, 1896

Subfamily: CERATOPHYRINAE Tschudi, 1838

Beelzebufo ampinga

SUBORDER: MESOBATRACHIA LAURENT, 1979

Superfamily: PELOBATOIDEA Stanius, 1856 , nom. corr. Spinar, 1971 (sensu Bolkay, 1919)

Family: PELOBATIDAE Stanius, 1856 , nom. corr. Spinar, 1971

Gen. sp indet.

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1903

INFRACLASS: NEODIAPSIDA BENTON, 1985

LEGION: LEPIDOSAUROMORPHA BENTON, 1983

SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)

MAGNORDER: SQUAMATA, OPPEL, 1811

SUPERORDER: SERPENTES LINNAEUS, 1759

Incertae sedis

Gen. sp indet.

ORDER: OPHIDIA BRONGNIART, 1800

SUBORDER: ALETHINOPHIDIA NOPCSCA, 1923

Superfamily: BOOIDEA Gray, 1825

Family: BOIDAE Gray, 1825

Subfamily: BOOIDINAE Gray, 1825

Infrafamily: BOOINIEA Gray, 1925

Tribe: MATSOINI Hoffstetter, 1961

Madtsoia madagascariensis

Family; ACROCHORDOIDAE Bonaparte, 1838

Subfamily: ?NIGEROPHEINAE Rage, 1975

Kelyophis hechti

LEGION: ARCHOSAUROMORPHA HUENE, 1946

SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985

INFRALEGION: DINOSAURIFORMES NOVAS, 1992

SUPERCOHORT: DINOSAURIA OWEN, 1842

COHORT: THEROPODA EMEND MARSH, 1881

AVEPODA PAUL, 2002

SUPERORDER: NEOTHEROPODA BAKKER, 1986

GRANDORDER: CERATOSAURIA NOVAS, 1992

AVEROSTRA PAUL, 2001

ORDER: CERATOSAURIA MARSH, 1884

Superfamily: ABELISAUROIDEA Novas, 1989

Family: NOASAURIDAE Bonaparte & Powell, 1980

Subfamily: VELOCISAURIINAE Bonaparte, 1991 (emend AGNOLIN, NOVAS & APESTEGUIA, 2003)

Masiakasaurus knopfleri

Family: ABELISAURIDAE Bonaparte & Novas, 1985

Subfamily: CARNOTAURINAE Sereno, 1998

Majungatholus atopus


_____________________________________________________________________________________

MADAGASCAR

MAJUNGA PROVINCE

MAHAJANGA BASIN

MAD 93-06

MAEVARANO FORMATION

ANEMBALEMBA MEMBER

MAASTRICHTIAN STAGE

SENONIAN SUBEPOCH

GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: AMPHIBIA LINNAEUS, 1758

SUBCLASS: LISSAMPHIBIA, HAECKEL, 1866?

SUPERORDER: SALIENTIA LAURENTI, 1768

ORDER: ANURA RAFINESQUE, 1815

SUBORDER: NEOBATRACHIA REIG, 1958

Superfamily: HYLOIDEA Rafinesque, 1815 (sensu Wied, 1856)

Family: LEPTODACTYLIDAE Werner, 1896

Subfamily: CERATOPHYRINAE Tschudi, 1838

Beelzebufo ampinga

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1903

SUPERLEGION: NEODIAPSIDA BENTON, 1985

LEGION: ARCHOSAUROMORPHA HUENE, 1946

SUPERCOHORT: CRUROTARSI SERENO & ARCUCCI, 1990

COHORT: CROCODYLOTARSI BENTON & CLARK, 1988

MAGNORDER: SUCHIA KREBS, 1974

SUPERORDER: CROCODYLOMORPHA WALKER, 1968

GRANDORDER: CROCODILIFORMES HAY, 1930

MICRORDER: MESOEUROCROCODYLIA WHETSTONE & WHYBROW, 1983

ORDER: MESOSUCHIA EMEND HUXLEY, 1875

PARVORDER: METASUCHIA BENTON & CLARK, 1988

GONDWANASUCHIA DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004

NOTOSUCHIMORPHA DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004

NOTOSUCHIFORMES DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004

TERRIASUCHIA DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004

INFRAORDER: NOTOSUCHIA GASPARINI, 1971

Family: NOTOSUCHIDAE Dollo, 1924

Simosuchus clarki

_____________________________________________________________________________________

MADAGASCAR

MAJUNGA PROVINCE

MAHAJANGA BASIN

MAD 93-09

MAEVARANO FORMATION

ANEMBALEMBA MEMBER

MAASTRICHTIAN STAGE

SENONIAN SUBEPOCH

GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1903

INFRACLASS: NEODIAPSIDA BENTON, 1985

LEGION: LEPIDOSAUROMORPHA BENTON, 1983

SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)

MAGNORDER: SQUAMATA, OPPEL, 1811

SUPERORDER: SERPENTES LINNAEUS, 1759

Incertae sedis

Gen. sp indet.

ORDER: OPHIDIA BRONGNIART, 1800

SUBORDER: ALETHINOPHIDIA NOPCSCA, 1923

Superfamily: BOOIDEA Gray, 1825

Family: BOIDAE Gray, 1825

Subfamily: BOOIDINAE Gray, 1825

Infrafamily: BOOINIEA Gray, 1925

Tribe: MATSOINI Hoffstetter, 1961

Madtsoia madagascariensis

Family; ACROCHORDOIDAE Bonaparte, 1838

Subfamily: ?NIGEROPHEINAE Rage, 1975

Kelyophis hechti

LEGION: ARCHOSAUROMORPHA HUENE, 1946

SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985

INFRALEGION: DINOSAURIFORMES NOVAS, 1992

SUPERCOHORT: DINOSAURIA OWEN, 1842

COHORT: THEROPODA EMEND MARSH, 1881

AVEPODA PAUL, 2002

SUPERORDER: NEOTHEROPODA BAKKER, 1986

GRANDORDER: CERATOSAURIA NOVAS, 1992

AVEROSTRA PAUL, 2001

ORDER: CERATOSAURIA MARSH, 1884

Superfamily: ABELISAUROIDEA Novas, 1989

Family: NOASAURIDAE Bonaparte & Powell, 1980

Subfamily: VELOCISAURIINAE Bonaparte, 1991 (emend AGNOLIN, NOVAS & APESTEGUIA, 2003)

Masiakasaurus knopfleri

_____________________________________________________________________________________

MADAGASCAR

MAJUNGA PROVINCE

MAHAJANGA BASIN

MAD 93-09

MAEVARANO FORMATION

ANEMBALEMBA MEMBER

MAASTRICHTIAN STAGE

SENONIAN SUBEPOCH

GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1903

INFRACLASS: NEODIAPSIDA BENTON, 1985

LEGION: LEPIDOSAUROMORPHA BENTON, 1983

SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)

MAGNORDER: SQUAMATA, OPPEL, 1811

SUPERORDER: SERPENTES LINNAEUS, 1759

ORDER: OPHIDIA BRONGNIART, 1800

SUBORDER: ALETHINOPHIDIA NOPCSCA, 1923

Superfamily: BOOIDEA Gray, 1825

Family: BOIDAE Gray, 1825

Subfamily: BOOIDINAE Gray, 1825

Infrafamily: BOOINIEA Gray, 1925

Tribe: MATSOINI Hoffstetter, 1961

Madtsoia madagascariensis

LEGION: ARCHOSAUROMORPHA HUENE, 1946

SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985

INFRALEGION: DINOSAURIFORMES NOVAS, 1992

SUPERCOHORT: DINOSAURIA OWEN, 1842

COHORT: THEROPODA EMEND MARSH, 1881

AVEPODA PAUL, 2002

SUPERORDER: NEOTHEROPODA BAKKER, 1986

GRANDORDER: CERATOSAURIA NOVAS, 1992

AVEROSTRA PAUL, 2001

ORDER: CERATOSAURIA MARSH, 1884

Superfamily: ABELISAUROIDEA Novas, 1989

Family: NOASAURIDAE Bonaparte & Powell, 1980

Subfamily: VELOCISAURIINAE Bonaparte, 1991 (emend AGNOLIN, NOVAS & APESTEGUIA, 2003)

Masiakasaurus knopfleri

_____________________________________________________________________________________

MADAGASCAR

MAJUNGA PROVINCE

MAHAJANGA BASIN

MAD 93-14

MAEVARANO FORMATION

ANEMBALEMBA MEMBER

MAASTRICHTIAN STAGE

SENONIAN SUBEPOCH

GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: AMPHIBIA LINNAEUS, 1758

SUBCLASS: LISSAMPHIBIA, HAECKEL, 1866?

SUPERORDER: SALIENTIA LAURENTI, 1768

ORDER: ANURA RAFINESQUE, 1815

SUBORDER: NEOBATRACHIA REIG, 1958

Superfamily: HYLOIDEA Rafinesque, 1815 (sensu Wied, 1856)

Family: LEPTODACTYLIDAE Werner, 1896

Subfamily: CERATOPHYRINAE Tschudi, 1838

Beelzebufo ampinga

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1903

INFRACLASS: NEODIAPSIDA BENTON, 1985

LEGION: LEPIDOSAUROMORPHA BENTON, 1983

SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)

MAGNORDER: SQUAMATA, OPPEL, 1811

SUPERORDER: SERPENTES LINNAEUS, 1759

ORDER: OPHIDIA BRONGNIART, 1800

SUBORDER: ALETHINOPHIDIA NOPCSCA, 1923

Superfamily: BOOIDEA Gray, 1825

Family: BOIDAE Gray, 1825

Subfamily: BOOIDINAE Gray, 1825

Infrafamily: BOOINIEA Gray, 1925

Tribe: MATSOINI Hoffstetter, 1961

Madtsoia madagascariensis

*Menarana nosymena

LEGION: ARCHOSAUROMORPHA HUENE, 1946

SUPERCOHORT: CRUROTARSI SERENO & ARCUCCI, 1990

COHORT: CROCODYLOTARSI BENTON & CLARK, 1988

MAGNORDER: SUCHIA KREBS, 1974

SUPERORDER: CROCODYLOMORPHA WALKER, 1968

GRANDORDER: CROCODILIFORMES HAY, 1930

MICRORDER: MESOEUROCROCODYLIA WHETSTONE & WHYBROW, 1983

ORDER: MESOSUCHIA EMEND HUXLEY, 1875

PARVORDER: METASUCHIA BENTON & CLARK, 1988

GONDWANASUCHIA DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004

NOTOSUCHIMORPHA DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004

NOTOSUCHIFORMES DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004

TERRIASUCHIA DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004

INFRAORDER: NOTOSUCHIA GASPARINI, 1971

Family: NOTOSUCHIDAE Dollo, 1924

Simosuchus clarki

_____________________________________________________________________________________

MADAGASCAR

MAHAJANGA BASIN

SOUTH OF BERIVORTA

MAD 93-16

MAEVARANO FORMATION

MAASTRICHTIAN STAGE

UPPER SENONIAN SUBEPOCH

UPPER UPPER GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1903

INFRACLASS: NEODIAPSIDA BENTON, 1985

LEGION: LEPIDOSAUROMORPHA BENTON, 1983

SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)

MAGNORDER: SQUAMATA, OPPEL, 1811

SUPERORDER: SERPENTES LINNAEUS, 1759

ORDER: OPHIDIA BRONGNIART, 1800

SUBORDER: ALETHINOPHIDIA NOPCSCA, 1923

Superfamily: BOOIDEA Gray, 1825

Family: BOIDAE Gray, 1825

Subfamily: BOOIDINAE Gray, 1825

Infrafamily: BOOINIEA Gray, 1925

Tribe: MATSOINI Hoffstetter, 1961

Menarana nosymena

 

_____________________________________________________________________________________

MADAGASCAR

MAHAJANGA BASIN

SOUTH OF BERIVORTA

MAD 93-17

MAEVARANO FORMATION

MAASTRICHTIAN STAGE

UPPER SENONIAN SUBEPOCH

UPPER UPPER GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: AMPHIBIA LINNAEUS, 1758

SUBCLASS: LISSAMPHIBIA, HAECKEL, 1866?

SUPERORDER: SALIENTIA LAURENTI, 1768

ORDER: ANURA RAFINESQUE, 1815

SUBORDER: NEOBATRACHIA REIG, 1958

Superfamily: HYLOIDEA Rafinesque, 1815 (sensu Wied, 1856)

Family: LEPTODACTYLIDAE Werner, 1896

Subfamily: CERATOPHYRINAE Tschudi, 1838

Beelzebufo ampinga

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1903

INFRACLASS: NEODIAPSIDA BENTON, 1985

LEGION: LEPIDOSAUROMORPHA BENTON, 1983

SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)

MAGNORDER: SQUAMATA, OPPEL, 1811

SUPERORDER: SERPENTES LINNAEUS, 1759

ORDER: OPHIDIA BRONGNIART, 1800

SUBORDER: ALETHINOPHIDIA NOPCSCA, 1923

Superfamily: BOOIDEA Gray, 1825

Family: BOIDAE Gray, 1825

Subfamily: BOOIDINAE Gray, 1825

Infrafamily: BOOINIEA Gray, 1925

Tribe: MATSOINI Hoffstetter, 1961

Madtsoia madagascariensis

 

_____________________________________________________________________________________

MADAGASCAR

MAJUNGA PROVINCE

MAHAJANGA BASIN

MAD 93-17

MAEVARANO FORMATION

ANEMBALEMBA MEMBER

MAASTRICHTIAN STAGE

SENONIAN SUBEPOCH

GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1903

INFRACLASS: NEODIAPSIDA BENTON, 1985

LEGION: LEPIDOSAUROMORPHA BENTON, 1983

SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)

MAGNORDER: SQUAMATA, OPPEL, 1811

SUPERORDER: SERPENTES LINNAEUS, 1759

ORDER: OPHIDIA BRONGNIART, 1800

SUBORDER: ALETHINOPHIDIA NOPCSCA, 1923

Superfamily: BOOIDEA Gray, 1825

Family: BOIDAE Gray, 1825

Subfamily: BOOIDINAE Gray, 1825

Infrafamily: BOOINIEA Gray, 1925

Tribe: MATSOINI Hoffstetter, 1961

Madtsoia madagascariensis

 

_____________________________________________________________________________________

MADAGASCAR

MAJUNGA PROVINCE

MAHAJANGA BASIN

MAD 93-18

MAEVARANO FORMATION

ANEMBALEMBA MEMBER

MAASTRICHTIAN STAGE

SENONIAN SUBEPOCH

GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: CHELONOMORPHA KUHN, 1960

SUPERCOHORT: TESTUDINES LINNAEUS, 1758

COHORT: CASICHELYDIA emend GAFFNEY & MEYLAN, 1988

MAGNORDER: PLEURODIRA GAFFNEY & MEYLAN, 1988

ORDER: PLEURODIRIDA COPE, 1864

SUBORDER: EUPLEURODIRA GAFFNEY & MEYLAN, 1988

Superfamily: PELOMEDUSOIDES Cope, 1868 Cope, 1868

Superfamily: BOTHREMYDINURA Baur, 1891

Epifamily: BOTHERMYDINURA Baur, 1891

Family: BOTHREMYDIDAE Baur, 1891

Subfamily: BOTHREMYDINAE Baur, 1891

Infrafamily: BOTHREMYDODDA Baur, 1891

Tribe: KURMADEMYDINI Gaffney, Tong & Meylan, 2006

Kinkonychelys rogersi

SUBCLASS: DIAPSIDA OSBORN, 1903

INFRACLASS: NEODIAPSIDA BENTON, 1985

LEGION: LEPIDOSAUROMORPHA BENTON, 1983

SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)

MAGNORDER: SQUAMATA, OPPEL, 1811

SUPERORDER: SERPENTES LINNAEUS, 1759

ORDER: OPHIDIA BRONGNIART, 1800

SUBORDER: ALETHINOPHIDIA NOPCSCA, 1923

Superfamily: BOOIDEA Gray, 1825

Family: BOIDAE Gray, 1825

Subfamily: BOOIDINAE Gray, 1825

Infrafamily: BOOINIEA Gray, 1925

Tribe: MATSOINI Hoffstetter, 1961

Madtsoia madagascariensis

LEGION: ARCHOSAUROMORPHA HUENE, 1946

SUPERCOHORT: CRUROTARSI SERENO & ARCUCCI, 1990

COHORT: CROCODYLOTARSI BENTON & CLARK, 1988

MAGNORDER: SUCHIA KREBS, 1974

SUPERORDER: CROCODYLOMORPHA WALKER, 1968

GRANDORDER: CROCODILIFORMES HAY, 1930

MICRORDER: MESOEUROCROCODYLIA WHETSTONE & WHYBROW, 1983

ORDER: MESOSUCHIA EMEND HUXLEY, 1875

PARVORDER: METASUCHIA BENTON & CLARK, 1988

GONDWANASUCHIA DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004

NOTOSUCHIMORPHA DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004

NOTOSUCHIFORMES DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004

TERRIASUCHIA DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004

INFRAORDER: NOTOSUCHIA GASPARINI, 1971

Family: NOTOSUCHIDAE Dollo, 1924

Simosuchus clarki

SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985

INFRALEGION: DINOSAURIFORMES NOVAS, 1992

SUPERCOHORT: DINOSAURIA OWEN, 1842

COHORT: THEROPODA EMEND MARSH, 1881

AVEPODA PAUL, 2002

SUPERORDER: NEOTHEROPODA BAKKER, 1986

GRANDORDER: CERATOSAURIA NOVAS, 1992

AVEROSTRA PAUL, 2001

ORDER: CERATOSAURIA MARSH, 1884

Superfamily: ABELISAUROIDEA Novas, 1989

Family: ABELISAURIDAE Bonaparte & Novas, 1985

Subfamily: CARNOTAURINAE Sereno, 1998

Majungatholus atopus

GRANDORDER: TETANURAE GAUTHIER, 1986

MICRORDER: COELUROSAURIA HUENE, 1914

ORDER: MANIRAPTORFORMES HOLTZ, 1996

AVETHEROPOA PAUL, 2002

AVEPECTORA PAUL, 2002

SUBORDER: MANIRAPTORA GAUTHIER, 1986

INFRAORDER: PARVES SERENO, 1997

PARVORDER: DEINONYCHOSAURIA COLBERT & D. A. RUSSELL, 1969

Family: DROMAEOSAURIDAE Russell, 1969

Subfamily: UNENLAGIINAE Bonaparte, 1999

*Rahonavis ostromi
= Rahona ostromi

COHORT: ORNITHISCHIFORMES COOPER, 1985
SUPERORDER: SAUROPODOMORPHA HUENE, 1932
GRANDORDER: BRONTOSAURIA OLSHEVSKY, 1991 (emend)
MICRORDER: SAUROPODA MARSH, 1878
ORDER: EUSAUROPODA
SUBORDER: NEOSAUROPODA BONAPARTE, 1986
INFRAORDER: MACRONARIA WILSON & SERENO, 1998
PARVORDER: TITANOSAURIFORMES SALGADO ET AL, 1997
Superfamily: TITANOSAUROIDEA Lydekker, 1885 (sensu Upchurch, 1999)
Family: TITANOSAURIDAE Lydekker, 1885
Incertae sedis
Vahiny depereti

 

CLASS: AVES LINNAEUS, 1758

Incertae sedis

Gen.sp indet.

SUBCLASS: SAURORNITHES NICHOLSON, 1879

AVEBREVICAUDA PAUL, 2001

INFRACLASS: ORNITHOTHORACES NOVA

Incertae sedis

Gen.sp indet.

LEGION: ORNITHURAE NOVA

Incertae sedis

Gen.sp indet.

COHORT: ENANTIORNITHES NOVA

Incertae sedis

Gen.sp indet.

SUPERORDER: EUENANTIORNITHES CHIAPPE, 2002

ORDER: EUORNITHIFORMES KUROCHKIN, 1996

Family: CONCORNITHIDAE Kurochkin, 1996

*Vorona berivotrensis

_____________________________________________________________________________________

MADAGASCAR

MAJUNGA PROVINCE

MAHAJANGA BASIN

MAD 93-19

MAEVARANO FORMATION

ANEMBALEMBA MEMBER

MAASTRICHTIAN STAGE

SENONIAN SUBEPOCH

GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1903

INFRACLASS: NEODIAPSIDA BENTON, 1985

LEGION: LEPIDOSAUROMORPHA BENTON, 1983

SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)

MAGNORDER: SQUAMATA, OPPEL, 1811

SUPERORDER: SERPENTES LINNAEUS, 1759

Incertae sedis

Gen. sp indet.

LEGION: ARCHOSAUROMORPHA HUENE, 1946

SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985

INFRALEGION: DINOSAURIFORMES NOVAS, 1992

SUPERCOHORT: DINOSAURIA OWEN, 1842

COHORT: THEROPODA EMEND MARSH, 1881

AVEPODA PAUL, 2002

SUPERORDER: NEOTHEROPODA BAKKER, 1986

GRANDORDER: CERATOSAURIA NOVAS, 1992

AVEROSTRA PAUL, 2001

ORDER: CERATOSAURIA MARSH, 1884

Superfamily: ABELISAUROIDEA Novas, 1989

Family: ABELISAURIDAE Bonaparte & Novas, 1985

Subfamily: CARNOTAURINAE Sereno, 1998

Majungatholus atopus


_____________________________________________________________________________________

MADAGASCAR

MAJUNGA PROVINCE

MAHAJANGA BASIN

MAD 93-25

MAEVARANO FORMATION

ANEMBALEMBA MEMBER

MAASTRICHTIAN STAGE

SENONIAN SUBEPOCH

GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: AMPHIBIA LINNAEUS, 1758

SUBCLASS: LISSAMPHIBIA, HAECKEL, 1866?

SUPERORDER: SALIENTIA LAURENTI, 1768

ORDER: ANURA RAFINESQUE, 1815

SUBORDER: NEOBATRACHIA REIG, 1958

Superfamily: HYLOIDEA Rafinesque, 1815 (sensu Wied, 1856)

Family: LEPTODACTYLIDAE Werner, 1896

Subfamily: CERATOPHYRINAE Tschudi, 1838

*Beelzebufo ampinga

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1903

INFRACLASS: NEODIAPSIDA BENTON, 1985

LEGION: LEPIDOSAUROMORPHA BENTON, 1983

SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)

MAGNORDER: SQUAMATA, OPPEL, 1811

SUPERORDER: SERPENTES LINNAEUS, 1759

ORDER: OPHIDIA BRONGNIART, 1800

SUBORDER: ALETHINOPHIDIA NOPCSCA, 1923

Superfamily: BOOIDEA Gray, 1825

Family: BOIDAE Gray, 1825

Subfamily: BOOIDINAE Gray, 1825

Infrafamily: BOOINIEA Gray, 1925

Tribe: MATSOINI Hoffstetter, 1961

Madtsoia madagascariensis

 

_____________________________________________________________________________________

MADAGASCAR

MAJUNGA PROVINCE

MAHAJANGA BASIN

MAD 93-25

MAEVARANO FORMATION

ANEMBALEMBA MEMBER

MAASTRICHTIAN STAGE

SENONIAN SUBEPOCH

GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1903

INFRACLASS: NEODIAPSIDA BENTON, 1985

LEGION: LEPIDOSAUROMORPHA BENTON, 1983

SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)

MAGNORDER: SQUAMATA, OPPEL, 1811

SUPERORDER: SERPENTES LINNAEUS, 1759

ORDER: OPHIDIA BRONGNIART, 1800

SUBORDER: ALETHINOPHIDIA NOPCSCA, 1923

Superfamily: BOOIDEA Gray, 1825

Family: BOIDAE Gray, 1825

Subfamily: BOOIDINAE Gray, 1825

Infrafamily: BOOINIEA Gray, 1925

Tribe: MATSOINI Hoffstetter, 1961

Madtsoia madagascariensis

 

_____________________________________________________________________________________

MADAGASCAR

MAJUNGA PROVINCE

MAHAJANGA BASIN

MAD 93-26

MAEVARANO FORMATION

ANEMBALEMBA MEMBER

MAASTRICHTIAN STAGE

SENONIAN SUBEPOCH

GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1903

SUPERLEGION: NEODIAPSIDA BENTON, 1985

LEGION: ARCHOSAUROMORPHA HUENE, 1946

SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985

INFRALEGION: DINOSAURIFORMES NOVAS, 1992

SUPERCOHORT: DINOSAURIA OWEN, 1842

COHORT: THEROPODA EMEND MARSH, 1881

AVEPODA PAUL, 2002

SUPERORDER: NEOTHEROPODA BAKKER, 1986

GRANDORDER: CERATOSAURIA NOVAS, 1992

AVEROSTRA PAUL, 2001

ORDER: CERATOSAURIA MARSH, 1884

Superfamily: ABELISAUROIDEA Novas, 1989

Family: NOASAURIDAE Bonaparte & Powell, 1980

Subfamily: VELOCISAURIINAE Bonaparte, 1991 (emend AGNOLIN, NOVAS & APESTEGUIA, 2003)

Masiakasaurus knopfleri

_____________________________________________________________________________________

MADAGASCAR

MAJUNGA PROVINCE

MAHAJANGA BASIN

MAD 93-27

MAEVARANO FORMATION

ANEMBALEMBA MEMBER

MAASTRICHTIAN STAGE

SENONIAN SUBEPOCH

GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1903

INFRACLASS: NEODIAPSIDA BENTON, 1985

LEGION: LEPIDOSAUROMORPHA BENTON, 1983

SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)

MAGNORDER: SQUAMATA, OPPEL, 1811

SUPERORDER: SERPENTES LINNAEUS, 1759

Incertae sedis

Gen. sp indet.

 

_____________________________________________________________________________________

MADAGASCAR

MAHAJANGA BASIN

SOUTH OF BERIVORTA

MAD 93-28

MAEVARANO FORMATION

MAASTRICHTIAN STAGE

UPPER SENONIAN SUBEPOCH

UPPER UPPER GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1903

INFRACLASS: NEODIAPSIDA BENTON, 1985

LEGION: LEPIDOSAUROMORPHA BENTON, 1983

SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)

MAGNORDER: SQUAMATA, OPPEL, 1811

SUPERORDER: SERPENTES LINNAEUS, 1759

ORDER: OPHIDIA BRONGNIART, 1800

SUBORDER: ALETHINOPHIDIA NOPCSCA, 1923

Superfamily: BOOIDEA Gray, 1825

Family: BOIDAE Gray, 1825

Subfamily: BOOIDINAE Gray, 1825

Infrafamily: BOOINIEA Gray, 1925

Tribe: MATSOINI Hoffstetter, 1961

Madtsoia madagascariensis

 

_____________________________________________________________________________________

MADAGASCAR

MAJUNGA PROVINCE

MAHAJANGA BASIN

MAD 93-30

MAEVARANO FORMATION

ANEMBALEMBA MEMBER

MAASTRICHTIAN STAGE

SENONIAN SUBEPOCH

GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1903

INFRACLASS: NEODIAPSIDA BENTON, 1985

LEGION: LEPIDOSAUROMORPHA BENTON, 1983

SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)

MAGNORDER: SQUAMATA, OPPEL, 1811

SUPERORDER: SERPENTES LINNAEUS, 1759

ORDER: OPHIDIA BRONGNIART, 1800

SUBORDER: ALETHINOPHIDIA NOPCSCA, 1923

Superfamily: BOOIDEA Gray, 1825

Family: BOIDAE Gray, 1825

Subfamily: BOOIDINAE Gray, 1825

Infrafamily: BOOINIEA Gray, 1925

Tribe: MATSOINI Hoffstetter, 1961

Madtsoia madagascariensis

LEGION: ARCHOSAUROMORPHA HUENE, 1946

SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985

INFRALEGION: DINOSAURIFORMES NOVAS, 1992

SUPERCOHORT: DINOSAURIA OWEN, 1842

COHORT: THEROPODA EMEND MARSH, 1881

AVEPODA PAUL, 2002

SUPERORDER: NEOTHEROPODA BAKKER, 1986

GRANDORDER: CERATOSAURIA NOVAS, 1992

AVEROSTRA PAUL, 2001

ORDER: CERATOSAURIA MARSH, 1884

Superfamily: ABELISAUROIDEA Novas, 1989

Family: NOASAURIDAE Bonaparte & Powell, 1980

Subfamily: VELOCISAURIINAE Bonaparte, 1991 (emend AGNOLIN, NOVAS & APESTEGUIA, 2003)

Masiakasaurus knopfleri

_____________________________________________________________________________________

MADAGASCAR

MAJUNGA PROVINCE

MAHAJANGA BASIN

MAD 93-33

MAEVARANO FORMATION

ANEMBALEMBA MEMBER

MAASTRICHTIAN STAGE

SENONIAN SUBEPOCH

GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: AMPHIBIA LINNAEUS, 1758

SUBCLASS: LISSAMPHIBIA, HAECKEL, 1866?

SUPERORDER: SALIENTIA LAURENTI, 1768

ORDER: ANURA RAFINESQUE, 1815

SUBORDER: MESOBATRACHIA LAURENT, 1979

Superfamily: PELOBATOIDEA Stanius, 1856 , nom. corr. Spinar, 1971 (sensu Bolkay, 1919)

Family: PELOBATIDAE Stanius, 1856 , nom. corr. Spinar, 1971

Gen. sp indet.

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1903 OSBORN, 1903

INFRACLASS: NEODIAPSIDA BENTON, 1985

LEGION: LEPIDOSAUROMORPHA BENTON, 1983

SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)

MAGNORDER: SQUAMATA, OPPEL, 1811

SUPERORDER: LACERTILIA OWEN, 1842

GRANDORDER: SCINCOGEKKONOMORPHA SUKHANOV, 1961

MICRORDER: SCINCOMORPHA CAMP, 1923

ORDER: LEPTOGLOSSA COPE, 1900

SUBORDER: ZONUROMORPHA ALIFANOV, 2000

INFRAORDER: ZONURIA FURBRINGER, 1900

Superfamily: CORDYLOIDEA Fritzinger, 1826

Family: CORDYLIDAE? Gray, 1837

*Konkasaurus mahalana

SUPERORDER: SERPENTES LINNAEUS, 1759

ORDER: OPHIDIA BRONGNIART, 1800

SUBORDER: ALETHINOPHIDIA NOPCSCA, 1923

Superfamily: BOOIDEA Gray, 1825

Family: BOIDAE Gray, 1825

Subfamily: BOOIDINAE Gray, 1825

Infrafamily: BOOINIEA Gray, 1925

Tribe: MATSOINI Hoffstetter, 1961

Madtsoia madagascariensis

 

LEGION: ARCHOSAUROMORPHA HUENE, 1946

SUPERCOHORT: CRUROTARSI SERENO & ARCUCCI, 1990

COHORT: CROCODYLOTARSI BENTON & CLARK, 1988

MAGNORDER: SUCHIA KREBS, 1974

SUPERORDER: CROCODYLOMORPHA WALKER, 1968

GRANDORDER: CROCODILIFORMES HAY, 1930

MICRORDER: MESOEUROCROCODYLIA WHETSTONE & WHYBROW, 1983

ORDER: MESOSUCHIA EMEND HUXLEY, 1875

PARVORDER: METASUCHIA BENTON & CLARK, 1988

GONDWANASUCHIA DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004

NOTOSUCHIMORPHA DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004

NOTOSUCHIFORMES DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004

Family: URUGUAYSUCHIDAE Gasparini, 1971

*Araripesuchus tsangatsangana

TERRIASUCHIA DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004

INFRAORDER: NOTOSUCHIA GASPARINI, 1971

Family: NOTOSUCHIDAE Dollo, 1924

Simosuchus clarki

SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985

INFRALEGION: DINOSAURIFORMES NOVAS, 1992

SUPERCOHORT: DINOSAURIA OWEN, 1842

COHORT: THEROPODA EMEND MARSH, 1881

AVEPODA PAUL, 2002

SUPERORDER: NEOTHEROPODA BAKKER, 1986

GRANDORDER: CERATOSAURIA NOVAS, 1992

AVEROSTRA PAUL, 2001

ORDER: CERATOSAURIA MARSH, 1884

Superfamily: ABELISAUROIDEA Novas, 1989

Family: ABELISAURIDAE Bonaparte & Novas, 1985

Subfamily: CARNOTAURINAE Sereno, 1998

Majungatholus atopus

 

_____________________________________________________________________________________

MADAGASCAR

MAJUNGA PROVINCE

MAHAJANGA BASIN

MAD 93-33

MAEVARANO FORMATION

ANEMBALEMBA MEMBER

MAASTRICHTIAN STAGE

SENONIAN SUBEPOCH

GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: AMPHIBIA LINNAEUS, 1758

SUBCLASS: LISSAMPHIBIA, HAECKEL, 1866?

SUPERORDER: SALIENTIA LAURENTI, 1768

ORDER: ANURA RAFINESQUE, 1815

SUBORDER: NEOBATRACHIA REIG, 1958

Superfamily: HYLOIDEA Rafinesque, 1815 (sensu Wied, 1856)

Family: LEPTODACTYLIDAE Werner, 1896

Subfamily: CERATOPHYRINAE Tschudi, 1838

Beelzebufo ampinga

_____________________________________________________________________________________

MADAGASCAR

MAJUNGA PROVINCE

MAHAJANGA BASIN

MAD 93-34

MAEVARANO FORMATION

ANEMBALEMBA MEMBER

MAASTRICHTIAN STAGE

SENONIAN SUBEPOCH

GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: AMPHIBIA LINNAEUS, 1758

SUBCLASS: LISSAMPHIBIA, HAECKEL, 1866?

SUPERORDER: SALIENTIA LAURENTI, 1768

ORDER: ANURA RAFINESQUE, 1815

SUBORDER: NEOBATRACHIA REIG, 1958

Superfamily: HYLOIDEA Rafinesque, 1815 (sensu Wied, 1856)

Family: LEPTODACTYLIDAE Werner, 1896

Subfamily: CERATOPHYRINAE Tschudi, 1838

Beelzebufo ampinga

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: CHELONOMORPHA KUHN, 1960

SUPERCOHORT: TESTUDINES LINNAEUS, 1758

COHORT: CASICHELYDIA emend GAFFNEY & MEYLAN, 1988

MAGNORDER: PLEURODIRA GAFFNEY & MEYLAN, 1988

ORDER: PLEURODIRIDA COPE, 1864

SUBORDER: EUPLEURODIRA GAFFNEY & MEYLAN, 1988

Superfamily: PELOMEDUSOIDES Cope, 1868 Cope, 1868

Hyperfamily: PODOCNEMIDERA Cope, 1868

CLADE: PAN-PODOCNEMIDIDAE

Family: SAHONACHELYIDAE Joyce, Rollot, Evers, Lyson, Rahantariosa & Krause, 2021

Sokatra antitra

SUBCLASS: DIAPSIDA OSBORN, 1903

INFRACLASS: NEODIAPSIDA BENTON, 1985

LEGION: LEPIDOSAUROMORPHA BENTON, 1983

SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)

MAGNORDER: SQUAMATA, OPPEL, 1811

SUPERORDER: SERPENTES LINNAEUS, 1759

ORDER: OPHIDIA BRONGNIART, 1800

SUBORDER: ALETHINOPHIDIA NOPCSCA, 1923

Superfamily: BOOIDEA Gray, 1825

Family: BOIDAE Gray, 1825

Subfamily: BOOIDINAE Gray, 1825

Infrafamily: BOOINIEA Gray, 1925

Tribe: MATSOINI Hoffstetter, 1961

Madtsoia madagascariensis

LEGION: ARCHOSAUROMORPHA HUENE, 1946

SUPERCOHORT: CRUROTARSI SERENO & ARCUCCI, 1990

COHORT: CROCODYLOTARSI BENTON & CLARK, 1988

MAGNORDER: SUCHIA KREBS, 1974

SUPERORDER: CROCODYLOMORPHA WALKER, 1968

GRANDORDER: CROCODILIFORMES HAY, 1930

MICRORDER: MESOEUROCROCODYLIA WHETSTONE & WHYBROW, 1983

SUBORDER: MESOEUROCROCODYLIA WHETSTONE & WHYBROW, 1983

MESOEUROCRODYLIA: NEW FAMILES?

GONDWANASUCHIA DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004

NOTOSUCHIMORPHA DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004

NOTOSUCHIFORMES DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004

TERRIASUCHIA DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004

INFRAORDER: NOTOSUCHIA GASPARINI, 1971

Family: NOTOSUCHIDAE Dollo, 1924

Simosuchus clarki

PEIROSAURIMORPHA DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004

PEIROSAURIFORMES DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004

Superfamily: PEIROSAURIDAE Gasparini, 1982 emend de Souza Carvalho, Ribeiro & dos Santos Avilla, 2004

Family: PEIROSAURIDAE Gasparini, 1982

Subfamily: LOMASUCHINAE de Souza Carvalho, Ribeiro & dos Santos Avilla, 2004

Tribe MAHAJANGASUCHINI de Souza Carvalho, Ribeiro & dos Santos Avilla, 2004

Mahajangasuchus insignis

LEGION: ARCHOSAUROMORPHA HUENE, 1946

SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985

INFRALEGION: DINOSAURIFORMES NOVAS, 1992

SUPERCOHORT: DINOSAURIA OWEN, 1842

COHORT: THEROPODA EMEND MARSH, 1881

AVEPODA PAUL, 2002

SUPERORDER: NEOTHEROPODA BAKKER, 1986

GRANDORDER: CERATOSAURIA NOVAS, 1992

AVEROSTRA PAUL, 2001

ORDER: CERATOSAURIA MARSH, 1884

Superfamily: ABELISAUROIDEA Novas, 1989

Family: NOASAURIDAE Bonaparte & Powell, 1980

Subfamily: VELOCISAURIINAE Bonaparte, 1991 (emend AGNOLIN, NOVAS & APESTEGUIA, 2003)

Masiakasaurus knopfleri

_____________________________________________________________________________________

MADAGASCAR

MAHAJANGA PROVINCE

MAHAJANGA BASIN

NEAR BERIVOTRA

MAD 93-35

MAEVARANO FORMATION

CAMPANIAN STAGE?

SENONIAN SUBEPOCH

UPPER GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: AMPHIBIA LINNAEUS, 1758

SUBCLASS: LISSAMPHIBIA, HAECKEL, 1866?

SUPERORDER: SALIENTIA LAURENTI, 1768

ORDER: ANURA RAFINESQUE, 1815

SUBORDER: NEOBATRACHIA REIG, 1958

Superfamily: HYLOIDEA Rafinesque, 1815 (sensu Wied, 1856)

Family: LEPTODACTYLIDAE Werner, 1896

Subfamily: CERATOPHYRINAE Tschudi, 1838

Beelzebufo ampinga

SUBORDER: MESOBATRACHIA LAURENT, 1979

Superfamily: PELOBATOIDEA Stanius, 1856 , nom. corr. Spinar, 1971 (sensu Bolkay, 1919)

Family: PELOBATIDAE Stanius, 1856 , nom. corr. Spinar, 1971

Gen. sp indet.

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: CHELONOMORPHA KUHN, 1960

SUPERCOHORT: TESTUDINES LINNAEUS, 1758

COHORT: CASICHELYDIA emend GAFFNEY & MEYLAN, 1988

MAGNORDER: PLEURODIRA GAFFNEY & MEYLAN, 1988

ORDER: PLEURODIRIDA COPE, 1864

SUBORDER: EUPLEURODIRA GAFFNEY & MEYLAN, 1988

Superfamily: PELOMEDUSOIDES Cope, 1868 Cope, 1868

Hyperfamily: PELOMEDUSERA Cope, 1868

Family: PELOMEDUSIDAE Cope, 1868

Subfamily: PELOMEDUSINAE Cope, 1868

Subfamily: Erymnochelus group

Cf. Erymnochelys

Hyperfamily: PODOCNEMIDERA Cope, 1868

CLADE: PAN-PODOCNEMIDIDAE

Family: SAHONACHELYIDAE Joyce, Rollot, Evers, Lyson, Rahantariosa & Krause, 2021

*Sokatra antitra

SUBCLASS: DIAPSIDA OSBORN, 1903

INFRACLASS: NEODIAPSIDA BENTON, 1985

LEGION: LEPIDOSAUROMORPHA BENTON, 1983

SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)

MAGNORDER: SQUAMATA, OPPEL, 1811

SUPERORDER: SERPENTES LINNAEUS, 1759

Incertae sedis

Gen. sp indet.

ORDER: OPHIDIA BRONGNIART, 1800

SUBORDER: ALETHINOPHIDIA NOPCSCA, 1923

Superfamily: BOOIDEA Gray, 1825

Family: BOIDAE Gray, 1825

Subfamily: BOOIDINAE Gray, 1825

Infrafamily: BOOINIEA Gray, 1925

Tribe: MATSOINI Hoffstetter, 1961

Madtsoia madagascariensis

Menarana nosymena

Family; ACROCHORDOIDAE Bonaparte, 1838

Subfamily: ?NIGEROPHEINAE Rage, 1975

Kelyophis hechti

LEGION: ARCHOSAUROMORPHA HUENE, 1946

SUPERCOHORT: CRUROTARSI SERENO & ARCUCCI, 1990

COHORT: CROCODYLOTARSI BENTON & CLARK, 1988

MAGNORDER: SUCHIA KREBS, 1974

SUPERORDER: CROCODYLOMORPHA WALKER, 1968

GRANDORDER: CROCODILIFORMES HAY, 1930

MICRORDER: MESOEUROCROCODYLIA WHETSTONE & WHYBROW, 1983

SUBORDER: MESOEUROCROCODYLIA WHETSTONE & WHYBROW, 1983

MESOEUROCRODYLIA: NEW FAMILES?

GONDWANASUCHIA DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004

NOTOSUCHIMORPHA DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004

NOTOSUCHIFORMES DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004

TERRIASUCHIA DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004

INFRAORDER: NOTOSUCHIA GASPARINI, 1971

Family: NOTOSUCHIDAE Dollo, 1924

Simosuchus clarki

PEIROSAURIMORPHA DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004

PEIROSAURIFORMES DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004

Superfamily: PEIROSAURIDAE Gasparini, 1982 emend de Souza Carvalho, Ribeiro & dos Santos Avilla, 2004

Family: PEIROSAURIDAE Gasparini, 1982

Subfamily: LOMASUCHINAE de Souza Carvalho, Ribeiro & dos Santos Avilla, 2004

Tribe MAHAJANGASUCHINI de Souza Carvalho, Ribeiro & dos Santos Avilla, 2004

*Mahajangasuchus insignis

LEGION: ARCHOSAUROMORPHA HUENE, 1946

SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985

INFRALEGION: DINOSAURIFORMES NOVAS, 1992

SUPERCOHORT: DINOSAURIA OWEN, 1842

COHORT: THEROPODA EMEND MARSH, 1881

AVEPODA PAUL, 2002

SUPERORDER: NEOTHEROPODA BAKKER, 1986

GRANDORDER: CERATOSAURIA NOVAS, 1992

AVEROSTRA PAUL, 2001

ORDER: CERATOSAURIA MARSH, 1884

Superfamily: ABELISAUROIDEA Novas, 1989

Family: NOASAURIDAE Bonaparte & Powell, 1980

Subfamily: VELOCISAURIINAE Bonaparte, 1991 (emend AGNOLIN, NOVAS & APESTEGUIA, 2003)

Masiakasaurus knopfleri

CLASS: AVES LINNAEUS, 1758

Incertae sedis

Gen.sp indet.

_____________________________________________________________________________________

MADAGASCAR

MAHAJANGA BASIN

SOUTH OF BERIVORTA

MAD 93-36

MAEVARANO FORMATION

MAASTRICHTIAN STAGE

UPPER SENONIAN SUBEPOCH

UPPER UPPER GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: AMPHIBIA LINNAEUS, 1758

SUBCLASS: LISSAMPHIBIA, HAECKEL, 1866?

SUPERORDER: SALIENTIA LAURENTI, 1768

ORDER: ANURA RAFINESQUE, 1815

SUBORDER: NEOBATRACHIA REIG, 1958

Superfamily: HYLOIDEA Rafinesque, 1815 (sensu Wied, 1856)

Family: LEPTODACTYLIDAE Werner, 1896

Subfamily: CERATOPHYRINAE Tschudi, 1838

Beelzebufo ampinga

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1903

INFRACLASS: NEODIAPSIDA BENTON, 1985

LEGION: LEPIDOSAUROMORPHA BENTON, 1983

SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)

MAGNORDER: SQUAMATA, OPPEL, 1811

SUPERORDER: SERPENTES LINNAEUS, 1759

ORDER: OPHIDIA BRONGNIART, 1800

SUBORDER: ALETHINOPHIDIA NOPCSCA, 1923

Superfamily: BOOIDEA Gray, 1825

Family: BOIDAE Gray, 1825

Subfamily: BOOIDINAE Gray, 1825

Infrafamily: BOOINIEA Gray, 1925

Tribe: MATSOINI Hoffstetter, 1961

Madtsoia madagascariensis

 

CLASS: AVES LINNAEUS, 1758

SUBCLASS: SAURORNITHES NICHOLSON, 1879

AVEBREVICAUDA PAUL, 2001

INFRACLASS: ORNITHOTHORACES NOVA

LEGION: ORNITHURAE NOVA

COHORT: ENANTIORNITHES

SUPERORDER: EUENANTIORNITHES CHIAPPE, 2002

ORDER: EUORNITHIFORMES KUROCHKIN, 1996

Family: CONCORNITHIDAE Kurochkin, 1996

Vorona berivotrensis

_____________________________________________________________________________________

MADAGASCAR

MAHAJANGA PROVINCE

MAHAJANGA BASIN

NEAR BERIVOTRA

MAD 93-37

MAEVARANO FORMATION

CAMPANIAN STAGE?

SENONIAN SUBEPOCH

UPPER GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: AMPHIBIA LINNAEUS, 1758

SUBCLASS: LISSAMPHIBIA, HAECKEL, 1866?

SUPERORDER: SALIENTIA LAURENTI, 1768

ORDER: ANURA RAFINESQUE, 1815

SUBORDER: MESOBATRACHIA LAURENT, 1979

Superfamily: PELOBATOIDEA Stanius, 1856 , nom. corr. Spinar, 1971 (sensu Bolkay, 1919)

Family: PELOBATIDAE Stanius, 1856 , nom. corr. Spinar, 1971

Gen. sp indet.

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1903

INFRACLASS: NEODIAPSIDA BENTON, 1985

LEGION: LEPIDOSAUROMORPHA BENTON, 1983

SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)

MAGNORDER: SQUAMATA, OPPEL, 1811

SUPERORDER: SERPENTES LINNAEUS, 1759

ORDER: OPHIDIA BRONGNIART, 1800

SUBORDER: ALETHINOPHIDIA NOPCSCA, 1923

Superfamily: BOOIDEA Gray, 1825

Family; ACROCHORDOIDAE Bonaparte, 1838

Subfamily: ?NIGEROPHEINAE Rage, 1975

Kelyophis hechti

_____________________________________________________________________________________

MADAGASCAR

MAJUNGA PROVINCE

MAHAJANGA BASIN

MAD 93-38

MAEVARANO FORMATION

ANEMBALEMBA MEMBER

MAASTRICHTIAN STAGE

SENONIAN SUBEPOCH

GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1903

INFRACLASS: NEODIAPSIDA BENTON, 1985

LEGION: LEPIDOSAUROMORPHA BENTON, 1983

SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)

MAGNORDER: SQUAMATA, OPPEL, 1811

SUPERORDER: SERPENTES LINNAEUS, 1759

ORDER: OPHIDIA BRONGNIART, 1800

SUBORDER: ALETHINOPHIDIA NOPCSCA, 1923

Superfamily: BOOIDEA Gray, 1825

Family: BOIDAE Gray, 1825

Subfamily: BOOIDINAE Gray, 1825

Infrafamily: BOOINIEA Gray, 1925

Tribe: MATSOINI Hoffstetter, 1961

Madtsoia madagascariensis

Gen. sp indet.

LEGION: ARCHOSAUROMORPHA HUENE, 1946

SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985

INFRALEGION: DINOSAURIFORMES NOVAS, 1992

SUPERCOHORT: DINOSAURIA OWEN, 1842

COHORT: THEROPODA EMEND MARSH, 1881

AVEPODA PAUL, 2002

SUPERORDER: NEOTHEROPODA BAKKER, 1986

GRANDORDER: CERATOSAURIA NOVAS, 1992

AVEROSTRA PAUL, 2001

ORDER: CERATOSAURIA MARSH, 1884

Superfamily: ABELISAUROIDEA Novas, 1989

Family: NOASAURIDAE Bonaparte & Powell, 1980

Subfamily: VELOCISAURIINAE Bonaparte, 1991 (emend AGNOLIN, NOVAS & APESTEGUIA, 2003)

Masiakasaurus knopfleri

_____________________________________________________________________________________

MADAGASCAR

MAJUNGA PROVINCE

MAHAJANGA BASIN

MAD 93-40

MAEVARANO FORMATION

ANEMBALEMBA MEMBER

MAASTRICHTIAN STAGE

SENONIAN SUBEPOCH

GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1903

INFRACLASS: NEODIAPSIDA BENTON, 1985

LEGION: LEPIDOSAUROMORPHA BENTON, 1983

SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)

MAGNORDER: SQUAMATA, OPPEL, 1811

SUPERORDER: SERPENTES LINNAEUS, 1759

Incertae sedis

Gen. sp indet.

 

_____________________________________________________________________________________

MADAGASCAR

MAJUNGA PROVINCE

MAHAJANGA BASIN

MAD 93-44

MAEVARANO FORMATION

ANEMBALEMBA MEMBER

MAASTRICHTIAN STAGE

SENONIAN SUBEPOCH

GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1903

INFRACLASS: NEODIAPSIDA BENTON, 1985

LEGION: LEPIDOSAUROMORPHA BENTON, 1983

SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)

MAGNORDER: SQUAMATA, OPPEL, 1811

SUPERORDER: SERPENTES LINNAEUS, 1759

Incertae sedis

Gen. sp indet.

 

_____________________________________________________________________________________

MADAGASCAR

MAJUNGA PROVINCE

MAHAJANGA BASIN

MAD 93-73

MAEVARANO FORMATION

ANEMBALEMBA MEMBER

MAASTRICHTIAN STAGE

SENONIAN SUBEPOCH

GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: AMPHIBIA LINNAEUS, 1758

SUBCLASS: LISSAMPHIBIA, HAECKEL, 1866?

SUPERORDER: SALIENTIA LAURENTI, 1768

ORDER: ANURA RAFINESQUE, 1815

SUBORDER: NEOBATRACHIA REIG, 1958

Superfamily: HYLOIDEA Rafinesque, 1815 (sensu Wied, 1856)

Family: LEPTODACTYLIDAE Werner, 1896

Subfamily: CERATOPHYRINAE Tschudi, 1838

Beelzebufo ampinga

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1903

INFRACLASS: NEODIAPSIDA BENTON, 1985

LEGION: LEPIDOSAUROMORPHA BENTON, 1983

SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)

MAGNORDER: SQUAMATA, OPPEL, 1811

SUPERORDER: SERPENTES LINNAEUS, 1759

ORDER: OPHIDIA BRONGNIART, 1800

SUBORDER: ALETHINOPHIDIA NOPCSCA, 1923

Superfamily: BOOIDEA Gray, 1825

Family: BOIDAE Gray, 1825

Subfamily: BOOIDINAE Gray, 1825

Infrafamily: BOOINIEA Gray, 1925

Tribe: MATSOINI Hoffstetter, 1961

*Adinophis fisaka

LEGION: ARCHOSAUROMORPHA HUENE, 1946

SUPERCOHORT: CRUROTARSI SERENO & ARCUCCI, 1990

COHORT: CROCODYLOTARSI BENTON & CLARK, 1988

MAGNORDER: SUCHIA KREBS, 1974

SUPERORDER: CROCODYLOMORPHA WALKER, 1968

GRANDORDER: CROCODILIFORMES HAY, 1930

MICRORDER: MESOEUROCROCODYLIA WHETSTONE & WHYBROW, 1983

ORDER: MESOSUCHIA EMEND HUXLEY, 1875

PARVORDER: METASUCHIA BENTON & CLARK, 1988

GONDWANASUCHIA DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004

NOTOSUCHIMORPHA DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004

NOTOSUCHIFORMES DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004

TERRIASUCHIA DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004

INFRAORDER: NOTOSUCHIA GASPARINI, 1971

Family: NOTOSUCHIDAE Dollo, 1924

Simosuchus clarki

SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985

INFRALEGION: DINOSAURIFORMES NOVAS, 1992

SUPERCOHORT: DINOSAURIA OWEN, 1842

COHORT: THEROPODA EMEND MARSH, 1881

AVEPODA PAUL, 2002

SUPERORDER: NEOTHEROPODA BAKKER, 1986

GRANDORDER: CERATOSAURIA NOVAS, 1992

AVEROSTRA PAUL, 2001

ORDER: CERATOSAURIA MARSH, 1884

Superfamily: ABELISAUROIDEA Novas, 1989

Family: ABELISAURIDAE Bonaparte & Novas, 1985

Subfamily: CARNOTAURINAE Sereno, 1998

Majungatholus atopus


_____________________________________________________________________________________

MADAGASCAR

MAJUNGA PROVINCE

MAHAJANGA BASIN

MAD 93-74

MAEVARANO FORMATION

ANEMBALEMBA MEMBER

MAASTRICHTIAN STAGE

SENONIAN SUBEPOCH

GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1903

SUPERLEGION: NEODIAPSIDA BENTON, 1985

LEGION: ARCHOSAUROMORPHA HUENE, 1946

SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985

INFRALEGION: DINOSAURIFORMES NOVAS, 1992

SUPERCOHORT: DINOSAURIA OWEN, 1842

COHORT: THEROPODA EMEND MARSH, 1881

AVEPODA PAUL, 2002

SUPERORDER: NEOTHEROPODA BAKKER, 1986

GRANDORDER: CERATOSAURIA NOVAS, 1992

AVEROSTRA PAUL, 2001

ORDER: CERATOSAURIA MARSH, 1884

Superfamily: ABELISAUROIDEA Novas, 1989

Family: NOASAURIDAE Bonaparte & Powell, 1980

Subfamily: VELOCISAURIINAE Bonaparte, 1991 (emend AGNOLIN, NOVAS & APESTEGUIA, 2003)

Masiakasaurus knopfleri

_____________________________________________________________________________________

MADAGASCAR

MAHAJANGA PROVINCE

MAHAJANGA BASIN

NEAR BERIVOTRA

MAD 93-79

MAEVARANO FORMATION

CAMPANIAN STAGE?

SENONIAN SUBEPOCH

UPPER GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: CHELONOMORPHA KUHN, 1960

SUPERCOHORT: TESTUDINES LINNAEUS, 1758

COHORT: CASICHELYDIA emend GAFFNEY & MEYLAN, 1988

MAGNORDER: PLEURODIRA GAFFNEY & MEYLAN, 1988

ORDER: PLEURODIRIDA COPE, 1864

SUBORDER: EUPLEURODIRA GAFFNEY & MEYLAN, 1988

Superfamily: PELOMEDUSOIDES Cope, 1868 Cope, 1868

Family: PELOMEDUSIDAE Cope, 1868

Subfamily: PELOMEDUSINAE Cope, 1868

Bothremyididae gen sp indet

_____________________________________________________________________________________

MADAGASCAR

MAJUNGA PROVINCE

MAHAJANGA BASIN

MAD 93-81

MAEVARANO FORMATION

ANEMBALEMBA MEMBER

MAASTRICHTIAN STAGE

SENONIAN SUBEPOCH

GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1903

INFRACLASS: NEODIAPSIDA BENTON, 1985

LEGION: LEPIDOSAUROMORPHA BENTON, 1983

SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)

MAGNORDER: SQUAMATA, OPPEL, 1811

SUPERORDER: SERPENTES LINNAEUS, 1759

Incertae sedis

Gen. sp indet.

ORDER: OPHIDIA BRONGNIART, 1800

SUBORDER: ALETHINOPHIDIA NOPCSCA, 1923

Superfamily: BOOIDEA Gray, 1825

Family: BOIDAE Gray, 1825

Subfamily: BOOIDINAE Gray, 1825

Infrafamily: BOOINIEA Gray, 1925

Tribe: MATSOINI Hoffstetter, 1961

Madtsoia madagascariensis

 
 

_____________________________________________________________________________________

MADAGASCAR

MAJUNGA PROVINCE

MAHAJANGA BASIN

MAD 93-86

MAEVARANO FORMATION

ANEMBALEMBA MEMBER

MAASTRICHTIAN STAGE

SENONIAN SUBEPOCH

GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1903

INFRACLASS: NEODIAPSIDA BENTON, 1985

LEGION: LEPIDOSAUROMORPHA BENTON, 1983

SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)

MAGNORDER: SQUAMATA, OPPEL, 1811

SUPERORDER: SERPENTES LINNAEUS, 1759

Incertae sedis

Gen. sp indet.

 

_____________________________________________________________________________________

MADAGASCAR

MAJUNGA PROVINCE

MAHAJANGA BASIN

MAD 93-101

MAEVARANO FORMATION

ANEMBALEMBA MEMBER

MAASTRICHTIAN STAGE

SENONIAN SUBEPOCH

GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1903

SUPERLEGION: NEODIAPSIDA BENTON, 1985

LEGION: ARCHOSAUROMORPHA HUENE, 1946

SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985

INFRALEGION: DINOSAURIFORMES NOVAS, 1992

SUPERCOHORT: DINOSAURIA OWEN, 1842

COHORT: THEROPODA EMEND MARSH, 1881

AVEPODA PAUL, 2002

SUPERORDER: NEOTHEROPODA BAKKER, 1986

GRANDORDER: CERATOSAURIA NOVAS, 1992

AVEROSTRA PAUL, 2001

ORDER: CERATOSAURIA MARSH, 1884

Superfamily: ABELISAUROIDEA Novas, 1989

Family: NOASAURIDAE Bonaparte & Powell, 1980

Subfamily: VELOCISAURIINAE Bonaparte, 1991 (emend AGNOLIN, NOVAS & APESTEGUIA, 2003)

Masiakasaurus knopfleri

_____________________________________________________________________________________

MADAGASCAR

MAJUNGA PROVINCE

MAHAJANGA BASIN

MAD 95-07

MAEVARANO FORMATION

ANEMBALEMBA MEMBER

MAASTRICHTIAN STAGE

SENONIAN SUBEPOCH

GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1903

SUPERLEGION: NEODIAPSIDA BENTON, 1985

LEGION: ARCHOSAUROMORPHA HUENE, 1946

SUPERCOHORT: CRUROTARSI SERENO & ARCUCCI, 1990

COHORT: CROCODYLOTARSI BENTON & CLARK, 1988

MAGNORDER: SUCHIA KREBS, 1974

SUPERORDER: CROCODYLOMORPHA WALKER, 1968

GRANDORDER: CROCODILIFORMES HAY, 1930

MICRORDER: MESOEUROCROCODYLIA WHETSTONE & WHYBROW, 1983

SUBORDER: MESOEUROCROCODYLIA WHETSTONE & WHYBROW, 1983

MESOEUROCRODYLIA: NEW FAMILES?

GONDWANASUCHIA DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004

PEIROSAURIMORPHA DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004

PEIROSAURIFORMES DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004

Superfamily: PEIROSAURIDAE Gasparini, 1982 emend de Souza Carvalho, Ribeiro & dos Santos Avilla, 2004

Family: PEIROSAURIDAE Gasparini, 1982

Subfamily: LOMASUCHINAE de Souza Carvalho, Ribeiro & dos Santos Avilla, 2004

Tribe MAHAJANGASUCHINI de Souza Carvalho, Ribeiro & dos Santos Avilla, 2004

Mahajangasuchus insignis

____________________________________________________________________________________

MADAGASCAR

MAJUNGA PROVINCE

MAHAJANGA BASIN

MAD 95-11

MAEVARANO FORMATION

ANEMBALEMBA MEMBER

MAASTRICHTIAN STAGE

SENONIAN SUBEPOCH

GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1903

LEGION: ARCHOSAUROMORPHA HUENE, 1946

SUPERCOHORT: CRUROTARSI SERENO & ARCUCCI, 1990

COHORT: CROCODYLOTARSI BENTON & CLARK, 1988

MAGNORDER: SUCHIA KREBS, 1974

SUPERORDER: CROCODYLOMORPHA WALKER, 1968

GRANDORDER: CROCODILIFORMES HAY, 1930

MICRORDER: MESOEUROCROCODYLIA WHETSTONE & WHYBROW, 1983

ORDER: MESOSUCHIA EMEND HUXLEY, 1875

PARVORDER: METASUCHIA BENTON & CLARK, 1988

GONDWANASUCHIA DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004

NOTOSUCHIMORPHA DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004

NOTOSUCHIFORMES DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004

TERRIASUCHIA DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004

INFRAORDER: NOTOSUCHIA GASPARINI, 1971

Family: NOTOSUCHIDAE Dollo, 1924

Simosuchus clarki

_____________________________________________________________________________________

MADAGASCAR

MAJUNGA PROVINCE

MAHAJANGA BASIN

MAD 95-14

MAEVARANO FORMATION

ANEMBALEMBA MEMBER

MAASTRICHTIAN STAGE

SENONIAN SUBEPOCH

GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1903

INFRACLASS: NEODIAPSIDA BENTON, 1985

LEGION: LEPIDOSAUROMORPHA BENTON, 1983

SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)

MAGNORDER: SQUAMATA, OPPEL, 1811

SUPERORDER: SERPENTES LINNAEUS, 1759

ORDER: OPHIDIA BRONGNIART, 1800

SUBORDER: ALETHINOPHIDIA NOPCSCA, 1923

Superfamily: BOOIDEA Gray, 1825

Family: BOIDAE Gray, 1825

Subfamily: BOOIDINAE Gray, 1825

Infrafamily: BOOINIEA Gray, 1925

Tribe: MATSOINI Hoffstetter, 1961

Madtsoia madagascariensis

LEGION: ARCHOSAUROMORPHA HUENE, 1946

SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985

INFRALEGION: DINOSAURIFORMES NOVAS, 1992

SUPERCOHORT: DINOSAURIA OWEN, 1842

COHORT: THEROPODA EMEND MARSH, 1881

AVEPODA PAUL, 2002

SUPERORDER: NEOTHEROPODA BAKKER, 1986

GRANDORDER: CERATOSAURIA NOVAS, 1992

AVEROSTRA PAUL, 2001

ORDER: CERATOSAURIA MARSH, 1884

Superfamily: ABELISAUROIDEA Novas, 1989

Family: NOASAURIDAE Bonaparte & Powell, 1980

Subfamily: VELOCISAURIINAE Bonaparte, 1991 (emend AGNOLIN, NOVAS & APESTEGUIA, 2003)

Masiakasaurus knopfleri

Family: ABELISAURIDAE Bonaparte & Novas, 1985

Subfamily: CARNOTAURINAE Sereno, 1998

Majungatholus atopus


_____________________________________________________________________________________

MADAGASCAR

MAJUNGA PROVINCE

MAHAJANGA BASIN

MAD 95-16

MAEVARANO FORMATION

ANEMBALEMBA MEMBER

MAASTRICHTIAN STAGE

SENONIAN SUBEPOCH

GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1903

SUPERLEGION: NEODIAPSIDA BENTON, 1985

LEGION: ARCHOSAUROMORPHA HUENE, 1946

SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985

INFRALEGION: DINOSAURIFORMES NOVAS, 1992

SUPERCOHORT: DINOSAURIA OWEN, 1842

COHORT: THEROPODA EMEND MARSH, 1881

AVEPODA PAUL, 2002

SUPERORDER: NEOTHEROPODA BAKKER, 1986

GRANDORDER: CERATOSAURIA NOVAS, 1992

AVEROSTRA PAUL, 2001

ORDER: CERATOSAURIA MARSH, 1884

Superfamily: ABELISAUROIDEA Novas, 1989

Family: ABELISAURIDAE Bonaparte & Novas, 1985

Subfamily: CARNOTAURINAE Sereno, 1998

Majungatholus atopus


_____________________________________________________________________________________

MADAGASCAR

MAJUNGA PROVINCE

MAHAJANGA BASIN

MAD 96-01

MAEVARANO FORMATION

ANEMBALEMBA MEMBER

MAASTRICHTIAN STAGE

SENONIAN SUBEPOCH

GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1903

INFRACLASS: NEODIAPSIDA BENTON, 1985

LEGION: LEPIDOSAUROMORPHA BENTON, 1983

SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)

MAGNORDER: SQUAMATA, OPPEL, 1811

SUPERORDER: SERPENTES LINNAEUS, 1759

ORDER: OPHIDIA BRONGNIART, 1800

SUBORDER: ALETHINOPHIDIA NOPCSCA, 1923

Superfamily: BOOIDEA Gray, 1825

Family: BOIDAE Gray, 1825

Subfamily: BOOIDINAE Gray, 1825

Infrafamily: BOOINIEA Gray, 1925

Tribe: MATSOINI Hoffstetter, 1961

Madtsoia madagascariensis

LEGION: ARCHOSAUROMORPHA HUENE, 1946

SUPERCOHORT: CRUROTARSI SERENO & ARCUCCI, 1990

COHORT: CROCODYLOTARSI BENTON & CLARK, 1988

MAGNORDER: SUCHIA KREBS, 1974

SUPERORDER: CROCODYLOMORPHA WALKER, 1968

GRANDORDER: CROCODILIFORMES HAY, 1930

MICRORDER: MESOEUROCROCODYLIA WHETSTONE & WHYBROW, 1983

ORDER: MESOSUCHIA EMEND HUXLEY, 1875

PARVORDER: METASUCHIA BENTON & CLARK, 1988

GONDWANASUCHIA DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004

NOTOSUCHIMORPHA DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004

NOTOSUCHIFORMES DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004

TERRIASUCHIA DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004

INFRAORDER: NOTOSUCHIA GASPARINI, 1971

Family: NOTOSUCHIDAE Dollo, 1924

Simosuchus clarki

SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985

INFRALEGION: DINOSAURIFORMES NOVAS, 1992

SUPERCOHORT: DINOSAURIA OWEN, 1842

COHORT: THEROPODA EMEND MARSH, 1881

AVEPODA PAUL, 2002

SUPERORDER: NEOTHEROPODA BAKKER, 1986

GRANDORDER: CERATOSAURIA NOVAS, 1992

AVEROSTRA PAUL, 2001

ORDER: CERATOSAURIA MARSH, 1884

Superfamily: ABELISAUROIDEA Novas, 1989

Family: NOASAURIDAE Bonaparte & Powell, 1980

Subfamily: VELOCISAURIINAE Bonaparte, 1991 (emend AGNOLIN, NOVAS & APESTEGUIA, 2003)

Masiakasaurus knopfleri

Family: ABELISAURIDAE Bonaparte & Novas, 1985

Subfamily: CARNOTAURINAE Sereno, 1998

Majungatholus atopus

CLASS: MAMMALIA LINNAEUS, 1758

SUBCLASS: THERIIFORMES (ROWE, 1988) McKENNA & BELL, 1997

INFRACLASS: ALLOTHERIA (MARSH, 1880) McKENNA & BELL, 1997

ORDER: MULTITUBERCULATA COPE, 1884

SUBORDER: GONDWANATHERIA Mones, 1987

Family: SUDAMERICICIDAE Scillato-Yane & Prasad, 1984

*Lavanify miolaka

_____________________________________________________________________________________

MADAGASCAR

MAJUNGA PROVINCE

MAHAJANGA BASIN

MAD 96-02

MAEVARANO FORMATION

ANEMBALEMBA MEMBER

MAASTRICHTIAN STAGE

SENONIAN SUBEPOCH

GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1903

SUPERLEGION: NEODIAPSIDA BENTON, 1985

LEGION: ARCHOSAUROMORPHA HUENE, 1946

SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985

INFRALEGION: DINOSAURIFORMES NOVAS, 1992

SUPERCOHORT: DINOSAURIA OWEN, 1842

COHORT: ORNITHISCHIFORMES COOPER, 1985

SUPERORDER: SAUROPODOMORPHA HUENE, 1932

GRANDORDER: BRONTOSAURIA OLSHEVSKY, 1991 (emend)

MICRORDER: SAUROPODA MARSH, 1878

ORDER: EUSAUROPODA

SUBORDER: NEOSAUROPODA BONAPARTE, 1986

INFRAORDER: MACRONARIA WILSON & SERENO, 1998

PARVORDER: TITANOSAURIFORMES SALGADO ET AL, 1997

Superfamily: TITANOSAUROIDEA Lydekker, 1885 (sensu Upchurch, 1999)

Family: NEMEGTOSAURIDAE Upchurch, 1995

Subfamily: NEMEGTOSAURINAE Upchurch, 1995

*Rapetosaurus krausei

_____________________________________________________________________________________

MADAGASCAR

MAJUNGA PROVINCE

MAHAJANGA BASIN

MAD 96-03

MAEVARANO FORMATION

ANEMBALEMBA MEMBER

MAASTRICHTIAN STAGE

SENONIAN SUBEPOCH

GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: CHELONOMORPHA KUHN, 1960

SUPERCOHORT: TESTUDINES LINNAEUS, 1758

COHORT: CASICHELYDIA emend GAFFNEY & MEYLAN, 1988

MAGNORDER: PLEURODIRA GAFFNEY & MEYLAN, 1988

ORDER: PLEURODIRIDA COPE, 1864

SUBORDER: EUPLEURODIRA GAFFNEY & MEYLAN, 1988

Superfamily: PELOMEDUSOIDES Cope, 1868 Cope, 1868

Superfamily: BOTHREMYDINURA Baur, 1891

Epifamily: BOTHERMYDINURA Baur, 1891

Family: BOTHREMYDIDAE Baur, 1891

Subfamily: BOTHREMYDINAE Baur, 1891

Infrafamily: BOTHREMYDODDA Baur, 1891

Tribe: KURMADEMYDINI Gaffney, Tong & Meylan, 2006

Kinkonychelys rogersi


_____________________________________________________________________________________

MADAGASCAR

MAHAJANGA BASIN

SOUTH OF BERIVORTA

MAD 96-04

MAEVARANO FORMATION

MAASTRICHTIAN STAGE

UPPER SENONIAN SUBEPOCH

UPPER UPPER GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1903

INFRACLASS: NEODIAPSIDA BENTON, 1985

LEGION: LEPIDOSAUROMORPHA BENTON, 1983

SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)

MAGNORDER: SQUAMATA, OPPEL, 1811

SUPERORDER: SERPENTES LINNAEUS, 1759

ORDER: OPHIDIA BRONGNIART, 1800

SUBORDER: ALETHINOPHIDIA NOPCSCA, 1923

Superfamily: BOOIDEA Gray, 1825

Family: BOIDAE Gray, 1825

Subfamily: BOOIDINAE Gray, 1825

Infrafamily: BOOINIEA Gray, 1925

Tribe: MATSOINI Hoffstetter, 1961

Madtsoia madagascariensis

 

_____________________________________________________________________________________

MADAGASCAR

MAJUNGA PROVINCE

MAHAJANGA BASIN

MAD 96-07

MAEVARANO FORMATION

ANEMBALEMBA MEMBER

MAASTRICHTIAN STAGE

SENONIAN SUBEPOCH

GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1903

SUPERLEGION: NEODIAPSIDA BENTON, 1985

LEGION: ARCHOSAUROMORPHA HUENE, 1946

SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985

INFRALEGION: DINOSAURIFORMES NOVAS, 1992

SUPERCOHORT: DINOSAURIA OWEN, 1842

COHORT: THEROPODA EMEND MARSH, 1881

AVEPODA PAUL, 2002

SUPERORDER: NEOTHEROPODA BAKKER, 1986

GRANDORDER: CERATOSAURIA NOVAS, 1992

AVEROSTRA PAUL, 2001

ORDER: CERATOSAURIA MARSH, 1884

Superfamily: ABELISAUROIDEA Novas, 1989

Family: ABELISAURIDAE Bonaparte & Novas, 1985

Subfamily: CARNOTAURINAE Sereno, 1998

Majungatholus atopus

COHORT: ORNITHISCHIFORMES COOPER, 1985

SUPERORDER: SAUROPODOMORPHA HUENE, 1932

GRANDORDER: BRONTOSAURIA OLSHEVSKY, 1991 (emend)

MICRORDER: SAUROPODA MARSH, 1878

ORDER: EUSAUROPODA

SUBORDER: NEOSAUROPODA BONAPARTE, 1986

INFRAORDER: MACRONARIA WILSON & SERENO, 1998

PARVORDER: TITANOSAURIFORMES SALGADO ET AL, 1997

Superfamily: TITANOSAUROIDEA Lydekker, 1885 (sensu Upchurch, 1999)

Family: TITANOSAURIDAE Lydekker, 1885

Incertae sedis

*Vahiny depereti

_____________________________________________________________________________________

 

MADAGASCAR

MAJUNGA PROVINCE

MAHAJANGA BASIN

MAD 96-18

MAEVARANO FORMATION

ANEMBALEMBA MEMBER

MAASTRICHTIAN STAGE

SENONIAN SUBEPOCH

GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1903

SUPERLEGION: NEODIAPSIDA BENTON, 1985

LEGION: ARCHOSAUROMORPHA HUENE, 1946

SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985

INFRALEGION: DINOSAURIFORMES NOVAS, 1992

SUPERCOHORT: DINOSAURIA OWEN, 1842

COHORT: THEROPODA EMEND MARSH, 1881

AVEPODA PAUL, 2002

SUPERORDER: NEOTHEROPODA BAKKER, 1986

GRANDORDER: CERATOSAURIA NOVAS, 1992

AVEROSTRA PAUL, 2001

ORDER: CERATOSAURIA MARSH, 1884

Superfamily: ABELISAUROIDEA Novas, 1989

Family: ABELISAURIDAE Bonaparte & Novas, 1985

Subfamily: CARNOTAURINAE Sereno, 1998

Majungatholus atopus


_____________________________________________________________________________________

MADAGASCAR

MAJUNGA PROVINCE

MAHAJANGA BASIN

MAD 96-21

MAEVARANO FORMATION

ANEMBALEMBA MEMBER

MAASTRICHTIAN STAGE

SENONIAN SUBEPOCH

GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: AMPHIBIA LINNAEUS, 1758

SUBCLASS: LISSAMPHIBIA, HAECKEL, 1866?

SUPERORDER: SALIENTIA LAURENTI, 1768

ORDER: ANURA RAFINESQUE, 1815

SUBORDER: NEOBATRACHIA REIG, 1958

Superfamily: HYLOIDEA Rafinesque, 1815 (sensu Wied, 1856)

Family: LEPTODACTYLIDAE Werner, 1896

Subfamily: CERATOPHYRINAE Tschudi, 1838

Beelzebufo ampinga

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1903

SUPERLEGION: NEODIAPSIDA BENTON, 1985

LEGION: ARCHOSAUROMORPHA HUENE, 1946

SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985

INFRALEGION: DINOSAURIFORMES NOVAS, 1992

SUPERCOHORT: DINOSAURIA OWEN, 1842

COHORT: THEROPODA EMEND MARSH, 1881

AVEPODA PAUL, 2002

SUPERORDER: NEOTHEROPODA BAKKER, 1986

GRANDORDER: CERATOSAURIA NOVAS, 1992

AVEROSTRA PAUL, 2001

ORDER: CERATOSAURIA MARSH, 1884

Superfamily: ABELISAUROIDEA Novas, 1989

Family: ABELISAURIDAE Bonaparte & Novas, 1985

Subfamily: CARNOTAURINAE Sereno, 1998

Majungatholus atopus


_____________________________________________________________________________________

MADAGASCAR

MAJUNGA PROVINCE

MAHAJANGA BASIN

MAD 96-24

MAEVARANO FORMATION

ANEMBALEMBA MEMBER

MAASTRICHTIAN STAGE

SENONIAN SUBEPOCH

GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: AMPHIBIA LINNAEUS, 1758

SUBCLASS: LISSAMPHIBIA, HAECKEL, 1866?

SUPERORDER: SALIENTIA LAURENTI, 1768

ORDER: ANURA RAFINESQUE, 1815

SUBORDER: NEOBATRACHIA REIG, 1958

Superfamily: HYLOIDEA Rafinesque, 1815 (sensu Wied, 1856)

Family: LEPTODACTYLIDAE Werner, 1896

Subfamily: CERATOPHYRINAE Tschudi, 1838

Beelzebufo ampinga

_____________________________________________________________________________________

MADAGASCAR

MAJUNGA PROVINCE

MAHAJANGA BASIN

MAD 96-41

MAEVARANO FORMATION

ANEMBALEMBA MEMBER

MAASTRICHTIAN STAGE

SENONIAN SUBEPOCH

GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1903

SUPERLEGION: NEODIAPSIDA BENTON, 1985

LEGION: ARCHOSAUROMORPHA HUENE, 1946

SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985

INFRALEGION: DINOSAURIFORMES NOVAS, 1992

SUPERCOHORT: DINOSAURIA OWEN, 1842

COHORT: THEROPODA EMEND MARSH, 1881

AVEPODA PAUL, 2002

SUPERORDER: NEOTHEROPODA BAKKER, 1986

GRANDORDER: CERATOSAURIA NOVAS, 1992

AVEROSTRA PAUL, 2001

ORDER: CERATOSAURIA MARSH, 1884

Superfamily: ABELISAUROIDEA Novas, 1989

Family: NOASAURIDAE Bonaparte & Powell, 1980

Subfamily: VELOCISAURIINAE Bonaparte, 1991 (emend AGNOLIN, NOVAS & APESTEGUIA, 2003)

Masiakasaurus knopfleri

_____________________________________________________________________________________

MADAGASCAR

MAHAJANGA BASIN

SOUTH OF BERIVORTA

MAD 98-08

MAEVARANO FORMATION

MAASTRICHTIAN STAGE

UPPER SENONIAN SUBEPOCH

UPPER UPPER GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1903

INFRACLASS: NEODIAPSIDA BENTON, 1985

LEGION: LEPIDOSAUROMORPHA BENTON, 1983

SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)

MAGNORDER: SQUAMATA, OPPEL, 1811

SUPERORDER: SERPENTES LINNAEUS, 1759

ORDER: OPHIDIA BRONGNIART, 1800

SUBORDER: ALETHINOPHIDIA NOPCSCA, 1923

Superfamily: BOOIDEA Gray, 1825

Family: BOIDAE Gray, 1825

Subfamily: BOOIDINAE Gray, 1825

Infrafamily: BOOINIEA Gray, 1925

Tribe: MATSOINI Hoffstetter, 1961

Madtsoia madagascariensis

 

_____________________________________________________________________________________

MADAGASCAR

MAHAJANGA PROVINCE

MAJUNGA REGION

SE OF BERIVOTRA

MAD 98-17

MAEVERANO FORMATION

MAASTRICHTIAN (PERHAPS LATE MAASTRICHTIAN?) STAGE

UPPERMOST SENONIAN SUBEPOCH

UPPERMOST GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1903

SUPERLEGION: NEODIAPSIDA BENTON, 1985

LEGION: ARCHOSAUROMORPHA HUENE, 1946

SUPERCOHORT: CRUROTARSI SERENO & ARCUCCI, 1990

COHORT: CROCODYLOTARSI BENTON & CLARK, 1988

MAGNORDER: SUCHIA KREBS, 1974

SUPERORDER: CROCODYLOMORPHA WALKER, 1968

GRANDORDER: CROCODILIFORMES HAY, 1930

MICRORDER: MESOEUROCROCODYLIA WHETSTONE & WHYBROW, 1983

ORDER: MESOSUCHIA EMEND HUXLEY, 1875

PARVORDER: METASUCHIA BENTON & CLARK, 1988

GONDWANASUCHIA DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004

NOTOSUCHIMORPHA DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004

NOTOSUCHIFORMES DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004

TERRIASUCHIA DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004

INFRAORDER: NOTOSUCHIA GASPARINI, 1971

Family: NOTOSUCHIDAE Dollo, 1924

*Simosuchus clarki

_____________________________________________________________________________________

MADAGASCAR

MAHAJANGA PROVINCE

WHICH LIE APPROXIMATELY 4-5 KM SOUTHEAST OF THE MIAN FIELD STUDY AREA SURROUNDING THE VILLAGE OF BERIVOTRA

BETWEEN THE TWO MIADANA HILLS

MAD98-19

MAEVARANO FORMATION

CAMPANIAN STAGE

SENONIAN SUBEPOCH

GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1903

SUPERLEGION: NEODIAPSIDA BENTON, 1985

LEGION: ARCHOSAUROMORPHA HUENE, 1946

SUPERCOHORT: CRUROTARSI SERENO & ARCUCCI, 1990

COHORT: CROCODYLOTARSI BENTON & CLARK, 1988

MAGNORDER: SUCHIA KREBS, 1974

SUPERORDER: CROCODYLOMORPHA WALKER, 1968

GRANDORDER: CROCODILIFORMES HAY, 1930

MICRORDER: MESOEUROCROCODYLIA WHETSTONE & WHYBROW, 1983

ORDER: MESOSUCHIA EMEND HUXLEY, 1875

PARVORDER: METASUCHIA BENTON & CLARK, 1988

INFRAORDER: METAMESOSUCHIA HULKE, 1878

NEOSUCHIA CLARK, IN BENTON & CLARK, 1988

Family: TREMATOCHAMPSIDAE Buffetaut, 1974

Miadanasuchus oblita
= Trematochampsa oblita

_____________________________________________________________________________________

MADAGASCAR

MAJUNGA PROVINCE

MAHAJANGA BASIN

MAD 98-25

MAEVARANO FORMATION

ANEMBALEMBA MEMBER

MAASTRICHTIAN STAGE

SENONIAN SUBEPOCH

GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: AMPHIBIA LINNAEUS, 1758

SUBCLASS: LISSAMPHIBIA, HAECKEL, 1866?

SUPERORDER: SALIENTIA LAURENTI, 1768

ORDER: ANURA RAFINESQUE, 1815

SUBORDER: NEOBATRACHIA REIG, 1958

Superfamily: HYLOIDEA Rafinesque, 1815 (sensu Wied, 1856)

Family: LEPTODACTYLIDAE Werner, 1896

Subfamily: CERATOPHYRINAE Tschudi, 1838

Beelzebufo ampinga

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1903

LEGION: ARCHOSAUROMORPHA HUENE, 1946

SUPERCOHORT: CRUROTARSI SERENO & ARCUCCI, 1990

COHORT: CROCODYLOTARSI BENTON & CLARK, 1988

MAGNORDER: SUCHIA KREBS, 1974

SUPERORDER: CROCODYLOMORPHA WALKER, 1968

GRANDORDER: CROCODILIFORMES HAY, 1930

MICRORDER: MESOEUROCROCODYLIA WHETSTONE & WHYBROW, 1983

ORDER: MESOSUCHIA EMEND HUXLEY, 1875

PARVORDER: METASUCHIA BENTON & CLARK, 1988

GONDWANASUCHIA DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004

NOTOSUCHIMORPHA DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004

NOTOSUCHIFORMES DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004

TERRIASUCHIA DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004

INFRAORDER: NOTOSUCHIA GASPARINI, 1971

Family: NOTOSUCHIDAE Dollo, 1924

Simosuchus clarki

_____________________________________________________________________________________

MADAGASCAR

MAJUNGA PROVINCE

MAHAJANGA BASIN

MAD 98-26

MAEVARANO FORMATION

ANEMBALEMBA MEMBER

MAASTRICHTIAN STAGE

SENONIAN SUBEPOCH

GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1903

SUPERLEGION: NEODIAPSIDA BENTON, 1985

LEGION: ARCHOSAUROMORPHA HUENE, 1946

SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985

INFRALEGION: DINOSAURIFORMES NOVAS, 1992

SUPERCOHORT: DINOSAURIA OWEN, 1842

COHORT: THEROPODA EMEND MARSH, 1881

AVEPODA PAUL, 2002

SUPERORDER: NEOTHEROPODA BAKKER, 1986

GRANDORDER: CERATOSAURIA NOVAS, 1992

AVEROSTRA PAUL, 2001

ORDER: CERATOSAURIA MARSH, 1884

Superfamily: ABELISAUROIDEA Novas, 1989

Family: NOASAURIDAE Bonaparte & Powell, 1980

Subfamily: VELOCISAURIINAE Bonaparte, 1991 (emend AGNOLIN, NOVAS & APESTEGUIA, 2003)

Masiakasaurus knopfleri

_____________________________________________________________________________________

MADAGASCAR

MAHAJANGA BASIN

SOUTH OF BERIVORTA

MAD 98-31

MAEVARANO FORMATION

MAASTRICHTIAN STAGE

UPPER SENONIAN SUBEPOCH

UPPER UPPER GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1903

INFRACLASS: NEODIAPSIDA BENTON, 1985

LEGION: LEPIDOSAUROMORPHA BENTON, 1983

SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)

MAGNORDER: SQUAMATA, OPPEL, 1811

SUPERORDER: SERPENTES LINNAEUS, 1759

ORDER: OPHIDIA BRONGNIART, 1800

SUBORDER: ALETHINOPHIDIA NOPCSCA, 1923

Superfamily: BOOIDEA Gray, 1825

Family: BOIDAE Gray, 1825

Subfamily: BOOIDINAE Gray, 1825

Infrafamily: BOOINIEA Gray, 1925

Tribe: MATSOINI Hoffstetter, 1961

Madtsoia madagascariensis

 

_____________________________________________________________________________________

MADAGASCAR

MAJUNGA PROVINCE

MAHAJANGA BASIN

MAD 99-05

MAEVARANO FORMATION

ANEMBALEMBA MEMBER

MAASTRICHTIAN STAGE

SENONIAN SUBEPOCH

GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1903

SUPERLEGION: NEODIAPSIDA BENTON, 1985

LEGION: ARCHOSAUROMORPHA HUENE, 1946

SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985

INFRALEGION: DINOSAURIFORMES NOVAS, 1992

SUPERCOHORT: DINOSAURIA OWEN, 1842

COHORT: THEROPODA EMEND MARSH, 1881

AVEPODA PAUL, 2002

SUPERORDER: NEOTHEROPODA BAKKER, 1986

GRANDORDER: CERATOSAURIA NOVAS, 1992

AVEROSTRA PAUL, 2001

ORDER: CERATOSAURIA MARSH, 1884

Superfamily: ABELISAUROIDEA Novas, 1989

Family: NOASAURIDAE Bonaparte & Powell, 1980

Subfamily: VELOCISAURIINAE Bonaparte, 1991 (emend AGNOLIN, NOVAS & APESTEGUIA, 2003)

Masiakasaurus knopfleri

_____________________________________________________________________________________

MADAGASCAR

MAJUNGA PROVINCE

MAHAJANGA BASIN

MAD 99-06

MAEVARANO FORMATION

ANEMBALEMBA MEMBER

MAASTRICHTIAN STAGE

SENONIAN SUBEPOCH

GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1903

SUPERLEGION: NEODIAPSIDA BENTON, 1985

LEGION: ARCHOSAUROMORPHA HUENE, 1946

SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985

INFRALEGION: DINOSAURIFORMES NOVAS, 1992

SUPERCOHORT: DINOSAURIA OWEN, 1842

COHORT: THEROPODA EMEND MARSH, 1881

AVEPODA PAUL, 2002

SUPERORDER: NEOTHEROPODA BAKKER, 1986

GRANDORDER: CERATOSAURIA NOVAS, 1992

AVEROSTRA PAUL, 2001

ORDER: CERATOSAURIA MARSH, 1884

Superfamily: ABELISAUROIDEA Novas, 1989

Family: NOASAURIDAE Bonaparte & Powell, 1980

Subfamily: VELOCISAURIINAE Bonaparte, 1991 (emend AGNOLIN, NOVAS & APESTEGUIA, 2003)

Masiakasaurus knopfleri

_____________________________________________________________________________________

MADAGASCAR

MAJUNGA PROVINCE

MAHAJANGA BASIN

MAD 99-15

MAEVARANO FORMATION

ANEMBALEMBA MEMBER

MAASTRICHTIAN STAGE

SENONIAN SUBEPOCH

GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: MAMMALIA LINNAEUS, 1758

SUBCLASS: THERIIFORMES (ROWE, 1988) McKENNA & BELL, 1997

INFRACLASS: ALLOTHERIA (MARSH, 1880) McKENNA & BELL, 1997

ORDER: MULTITUBERCULATA COPE, 1884

SUBORDER: GONDWANATHERIA Mones, 1987

Incertae sedis

*Adalatherium hui

_____________________________________________________________________________________

MADAGASCAR

MAJUNGA PROVINCE

MAHAJANGA BASIN

MAD 99-26

MAEVARANO FORMATION

ANEMBALEMBA MEMBER

MAASTRICHTIAN STAGE

SENONIAN SUBEPOCH

GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1903

SUPERLEGION: NEODIAPSIDA BENTON, 1985

LEGION: ARCHOSAUROMORPHA HUENE, 1946

SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985

INFRALEGION: DINOSAURIFORMES NOVAS, 1992

SUPERCOHORT: DINOSAURIA OWEN, 1842

COHORT: THEROPODA EMEND MARSH, 1881

AVEPODA PAUL, 2002

SUPERORDER: NEOTHEROPODA BAKKER, 1986

GRANDORDER: CERATOSAURIA NOVAS, 1992

AVEROSTRA PAUL, 2001

ORDER: CERATOSAURIA MARSH, 1884

Superfamily: ABELISAUROIDEA Novas, 1989

Family: ABELISAURIDAE Bonaparte & Novas, 1985

Subfamily: CARNOTAURINAE Sereno, 1998

Majungatholus atopus


_____________________________________________________________________________________

MADAGASCAR

MAJUNGA PROVINCE

MAHAJANGA BASIN

MAD 99-29

MAEVARANO FORMATION

ANEMBALEMBA MEMBER

MAASTRICHTIAN STAGE

SENONIAN SUBEPOCH

GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: AMPHIBIA LINNAEUS, 1758

SUBCLASS: LISSAMPHIBIA, HAECKEL, 1866?

SUPERORDER: SALIENTIA LAURENTI, 1768

ORDER: ANURA RAFINESQUE, 1815

SUBORDER: NEOBATRACHIA REIG, 1958

Superfamily: HYLOIDEA Rafinesque, 1815 (sensu Wied, 1856)

Family: LEPTODACTYLIDAE Werner, 1896

Subfamily: CERATOPHYRINAE Tschudi, 1838

Beelzebufo ampinga

_____________________________________________________________________________________

MADAGASCAR

MAJUNGA PROVINCE

MAHAJANGA BASIN

MAD 99-30

MAEVARANO FORMATION

ANEMBALEMBA MEMBER

MAASTRICHTIAN STAGE

SENONIAN SUBEPOCH

GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1903

SUPERLEGION: NEODIAPSIDA BENTON, 1985

LEGION: ARCHOSAUROMORPHA HUENE, 1946

SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985

INFRALEGION: DINOSAURIFORMES NOVAS, 1992

SUPERCOHORT: DINOSAURIA OWEN, 1842

COHORT: THEROPODA EMEND MARSH, 1881

AVEPODA PAUL, 2002

SUPERORDER: NEOTHEROPODA BAKKER, 1986

GRANDORDER: CERATOSAURIA NOVAS, 1992

AVEROSTRA PAUL, 2001

ORDER: CERATOSAURIA MARSH, 1884

Superfamily: ABELISAUROIDEA Novas, 1989

Family: NOASAURIDAE Bonaparte & Powell, 1980

Subfamily: VELOCISAURIINAE Bonaparte, 1991 (emend AGNOLIN, NOVAS & APESTEGUIA, 2003)

Masiakasaurus knopfleri

_____________________________________________________________________________________

MADAGASCAR

MAHAJANGA BASIN

SOUTH OF BERIVORTA

MAD 99-31

MAEVARANO FORMATION

MAASTRICHTIAN STAGE

UPPER SENONIAN SUBEPOCH

UPPER UPPER GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1903

INFRACLASS: NEODIAPSIDA BENTON, 1985

LEGION: LEPIDOSAUROMORPHA BENTON, 1983

SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)

MAGNORDER: SQUAMATA, OPPEL, 1811

SUPERORDER: SERPENTES LINNAEUS, 1759

ORDER: OPHIDIA BRONGNIART, 1800

SUBORDER: ALETHINOPHIDIA NOPCSCA, 1923

Superfamily: BOOIDEA Gray, 1825

Family: BOIDAE Gray, 1825

Subfamily: BOOIDINAE Gray, 1825

Infrafamily: BOOINIEA Gray, 1925

Tribe: MATSOINI Hoffstetter, 1961

Madtsoia madagascariensis

 

_____________________________________________________________________________________

MADAGASCAR

MAJUNGA PROVINCE

MAHAJANGA BASIN

MAD 99-31

MAEVARANO FORMATION

ANEMBALEMBA MEMBER

MAASTRICHTIAN STAGE

SENONIAN SUBEPOCH

GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: CHELONOMORPHA KUHN, 1960

SUPERCOHORT: TESTUDINES LINNAEUS, 1758

COHORT: CASICHELYDIA emend GAFFNEY & MEYLAN, 1988

MAGNORDER: PLEURODIRA GAFFNEY & MEYLAN, 1988

ORDER: PLEURODIRIDA COPE, 1864

SUBORDER: EUPLEURODIRA GAFFNEY & MEYLAN, 1988

Superfamily: PELOMEDUSOIDES Cope, 1868 Cope, 1868

Superfamily: BOTHREMYDINURA Baur, 1891

Epifamily: BOTHERMYDINURA Baur, 1891

Family: BOTHREMYDIDAE Baur, 1891

Subfamily: BOTHREMYDINAE Baur, 1891

Infrafamily: BOTHREMYDODDA Baur, 1891

Tribe: KURMADEMYDINI Gaffney, Tong & Meylan, 2006

Kinkonychelys rogersi

SUBCLASS: DIAPSIDA OSBORN, 1903

INFRACLASS: NEODIAPSIDA BENTON, 1985

LEGION: LEPIDOSAUROMORPHA BENTON, 1983

SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)

MAGNORDER: SQUAMATA, OPPEL, 1811

SUPERORDER: SERPENTES LINNAEUS, 1759

ORDER: OPHIDIA BRONGNIART, 1800

SUBORDER: ALETHINOPHIDIA NOPCSCA, 1923

Superfamily: BOOIDEA Gray, 1825

Family: BOIDAE Gray, 1825

Subfamily: BOOIDINAE Gray, 1825

Infrafamily: BOOINIEA Gray, 1925

Tribe: MATSOINI Hoffstetter, 1961

Menarana nosymena

_____________________________________________________________________________________

MADAGASCAR

MAJUNGA PROVINCE

MAHAJANGA BASIN

MAD 99-32

MAEVARANO FORMATION

ANEMBALEMBA MEMBER

MAASTRICHTIAN STAGE

SENONIAN SUBEPOCH

GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1903

SUPERLEGION: NEODIAPSIDA BENTON, 1985

LEGION: ARCHOSAUROMORPHA HUENE, 1946

SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985

INFRALEGION: DINOSAURIFORMES NOVAS, 1992

SUPERCOHORT: DINOSAURIA OWEN, 1842

COHORT: THEROPODA EMEND MARSH, 1881

AVEPODA PAUL, 2002

SUPERORDER: NEOTHEROPODA BAKKER, 1986

GRANDORDER: CERATOSAURIA NOVAS, 1992

AVEROSTRA PAUL, 2001

ORDER: CERATOSAURIA MARSH, 1884

Superfamily: ABELISAUROIDEA Novas, 1989

Family: ABELISAURIDAE Bonaparte & Novas, 1985

Subfamily: CARNOTAURINAE Sereno, 1998

Majungatholus atopus


_____________________________________________________________________________________

MADAGASCAR

MAJUNGA PROVINCE

MAHAJANGA BASIN

MAD 99-33

MAEVARANO FORMATION

ANEMBALEMBA MEMBER

MAASTRICHTIAN STAGE

SENONIAN SUBEPOCH

GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1903

SUPERLEGION: NEODIAPSIDA BENTON, 1985

LEGION: ARCHOSAUROMORPHA HUENE, 1946

SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985

INFRALEGION: DINOSAURIFORMES NOVAS, 1992

SUPERCOHORT: DINOSAURIA OWEN, 1842

COHORT: THEROPODA EMEND MARSH, 1881

AVEPODA PAUL, 2002

SUPERORDER: NEOTHEROPODA BAKKER, 1986

GRANDORDER: CERATOSAURIA NOVAS, 1992

AVEROSTRA PAUL, 2001

ORDER: CERATOSAURIA MARSH, 1884

Superfamily: ABELISAUROIDEA Novas, 1989

Family: NOASAURIDAE Bonaparte & Powell, 1980

Subfamily: VELOCISAURIINAE Bonaparte, 1991 (emend AGNOLIN, NOVAS & APESTEGUIA, 2003)

Masiakasaurus knopfleri

Family: ABELISAURIDAE Bonaparte & Novas, 1985

Subfamily: CARNOTAURINAE Sereno, 1998

Majungatholus atopus


_____________________________________________________________________________________

MADAGASCAR

MAJUNGA PROVINCE

MAHAJANGA BASIN

MAD 99-35

MAEVARANO FORMATION

ANEMBALEMBA MEMBER

MAASTRICHTIAN STAGE

SENONIAN SUBEPOCH

GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1903

SUPERLEGION: NEODIAPSIDA BENTON, 1985

LEGION: ARCHOSAUROMORPHA HUENE, 1946

SUPERCOHORT: CRUROTARSI SERENO & ARCUCCI, 1990

COHORT: CROCODYLOTARSI BENTON & CLARK, 1988

MAGNORDER: SUCHIA KREBS, 1974

SUPERORDER: CROCODYLOMORPHA WALKER, 1968

GRANDORDER: CROCODILIFORMES HAY, 1930

MICRORDER: MESOEUROCROCODYLIA WHETSTONE & WHYBROW, 1983

SUBORDER: MESOEUROCROCODYLIA WHETSTONE & WHYBROW, 1983

MESOEUROCRODYLIA: NEW FAMILES?

GONDWANASUCHIA DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004

PEIROSAURIMORPHA DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004

PEIROSAURIFORMES DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004

Superfamily: PEIROSAURIDAE Gasparini, 1982 emend de Souza Carvalho, Ribeiro & dos Santos Avilla, 2004

Family: PEIROSAURIDAE Gasparini, 1982

Subfamily: LOMASUCHINAE de Souza Carvalho, Ribeiro & dos Santos Avilla, 2004

Tribe MAHAJANGASUCHINI de Souza Carvalho, Ribeiro & dos Santos Avilla, 2004

Mahajangasuchus insignis

____________________________________________________________________________________

MADAGASCAR

MAHAJANGA BASIN

SOUTH OF BERIVORTA

MAD 99-39

MAEVARANO FORMATION

MAASTRICHTIAN STAGE

UPPER SENONIAN SUBEPOCH

UPPER UPPER GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1903

INFRACLASS: NEODIAPSIDA BENTON, 1985

LEGION: LEPIDOSAUROMORPHA BENTON, 1983

SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)

MAGNORDER: SQUAMATA, OPPEL, 1811

SUPERORDER: SERPENTES LINNAEUS, 1759

ORDER: OPHIDIA BRONGNIART, 1800

SUBORDER: ALETHINOPHIDIA NOPCSCA, 1923

Superfamily: BOOIDEA Gray, 1825

Family: BOIDAE Gray, 1825

Subfamily: BOOIDINAE Gray, 1825

Infrafamily: BOOINIEA Gray, 1925

Tribe: MATSOINI Hoffstetter, 1961

Madtsoia madagascariensis

 

_____________________________________________________________________________________

MADAGASCAR

MAJUNGA PROVINCE

MAHAJANGA BASIN

MAD 01-04

MAEVARANO FORMATION

ANEMBALEMBA MEMBER

MAASTRICHTIAN STAGE

SENONIAN SUBEPOCH

GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1903

SUPERLEGION: NEODIAPSIDA BENTON, 1985

LEGION: ARCHOSAUROMORPHA HUENE, 1946

SUPERCOHORT: CRUROTARSI SERENO & ARCUCCI, 1990

COHORT: CROCODYLOTARSI BENTON & CLARK, 1988

MAGNORDER: SUCHIA KREBS, 1974

SUPERORDER: CROCODYLOMORPHA WALKER, 1968

GRANDORDER: CROCODILIFORMES HAY, 1930

MICRORDER: MESOEUROCROCODYLIA WHETSTONE & WHYBROW, 1983

SUBORDER: MESOEUROCROCODYLIA WHETSTONE & WHYBROW, 1983

MESOEUROCRODYLIA: NEW FAMILES?

GONDWANASUCHIA DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004

PEIROSAURIMORPHA DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004

PEIROSAURIFORMES DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004

Superfamily: PEIROSAURIDAE Gasparini, 1982 emend de Souza Carvalho, Ribeiro & dos Santos Avilla, 2004

Family: PEIROSAURIDAE Gasparini, 1982

Subfamily: LOMASUCHINAE de Souza Carvalho, Ribeiro & dos Santos Avilla, 2004

Tribe MAHAJANGASUCHINI de Souza Carvalho, Ribeiro & dos Santos Avilla, 2004

Mahajangasuchus insignis

____________________________________________________________________________________

MADAGASCAR

MAHAJANGA BASIN

MASIAKAKOHO STUDY AREA LOCALITY

MAD03-15

MAEVARANO FORMATION

MAASTRICHTIAN STAGE

UPPER SENONIAN SUBEPOCH

UPPER UPPER GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1903

INFRACLASS: NEODIAPSIDA BENTON, 1985

LEGION: LEPIDOSAUROMORPHA BENTON, 1983

SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)

MAGNORDER: SQUAMATA, OPPEL, 1811

SUPERORDER: SERPENTES LINNAEUS, 1759

ORDER: OPHIDIA BRONGNIART, 1800

SUBORDER: ALETHINOPHIDIA NOPCSCA, 1923

Superfamily: BOOIDEA Gray, 1825

Family; ACROCHORDOIDAE Bonaparte, 1838

Subfamily: ?NIGEROPHEINAE Rage, 1975

Kelyophis hechti

_____________________________________________________________________________________

MADAGASCAR

MAHAJANGA BASIN

MASIAKAKOHO STUDY AREA LOCALITY

MAD03-23

MAEVARANO FORMATION

MAASTRICHTIAN STAGE

UPPER SENONIAN SUBEPOCH

UPPER UPPER GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1903

INFRACLASS: NEODIAPSIDA BENTON, 1985

LEGION: LEPIDOSAUROMORPHA BENTON, 1983

SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)

MAGNORDER: SQUAMATA, OPPEL, 1811

SUPERORDER: SERPENTES LINNAEUS, 1759

ORDER: OPHIDIA BRONGNIART, 1800

SUBORDER: ALETHINOPHIDIA NOPCSCA, 1923

Superfamily: BOOIDEA Gray, 1825

Family: BOIDAE Gray, 1825

Subfamily: BOOIDINAE Gray, 1825

Infrafamily: BOOINIEA Gray, 1925

Tribe: MATSOINI Hoffstetter, 1961

Madtsoia madagascariensis

 

____________________________________________________________________________________

MADAGASCAR

MAJUNGA PROVINCE

MAHAJANGA BASIN

MAD05-59

MAEVARANO FORMATION

ANEMBALEMBA MEMBER

MAASTRICHTIAN STAGE

SENONIAN SUBEPOCH

GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1903

INFRACLASS: NEODIAPSIDA BENTON, 1985

LEGION: LEPIDOSAUROMORPHA BENTON, 1983

SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)

MAGNORDER: SQUAMATA, OPPEL, 1811

SUPERORDER: SERPENTES LINNAEUS, 1759

Incertae sedis

Gen. sp indet.

ORDER: OPHIDIA BRONGNIART, 1800

SUBORDER: ALETHINOPHIDIA NOPCSCA, 1923

Superfamily: BOOIDEA Gray, 1825

Family: BOIDAE Gray, 1825

Subfamily: BOOIDINAE Gray, 1825

Infrafamily: BOOINIEA Gray, 1925

Tribe: MATSOINI Hoffstetter, 1961

Menarana nosymena

Family; ACROCHORDOIDAE Bonaparte, 1838

Subfamily: ?NIGEROPHEINAE Rage, 1975

Kelyophis hechti

_____________________________________________________________________________________

MADAGASCAR

MAJUNGA PROVINCE

MAHAJANGA BASIN

MAD07-37

MAEVARANO FORMATION

ANEMBALEMBA MEMBER

MAASTRICHTIAN STAGE

SENONIAN SUBEPOCH

GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1903

INFRACLASS: NEODIAPSIDA BENTON, 1985

LEGION: LEPIDOSAUROMORPHA BENTON, 1983

SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)

MAGNORDER: SQUAMATA, OPPEL, 1811

SUPERORDER: SERPENTES LINNAEUS, 1759

ORDER: OPHIDIA BRONGNIART, 1800

SUBORDER: ALETHINOPHIDIA NOPCSCA, 1923

Superfamily: BOOIDEA Gray, 1825

Family: BOIDAE Gray, 1825

Subfamily: BOOIDINAE Gray, 1825

Infrafamily: BOOINIEA Gray, 1925

Tribe: MATSOINI Hoffstetter, 1961

Menarana nosymena

 

_____________________________________________________________________________________