96MADAGASCAR
MAHAJANGA BASIN
FROM THE NEIGHBOURING OF AMBONDROMAMY
BATHONIAN STAGE
MIDDLE DOGGER EPOCH
MIDDLE JURASSIC
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
Incertae sedis
Morphotype 2
Gen. sp indet.
_____________________________________________________________________________________
MADAGASCAR
MAHAJANGA BASIN
HILLS W OF AMBONDROMAMY
BATHONIAN STAGE
MIDDLE DOGGER EPOCH
MIDDLE JURASSIC
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
INFRACLASS: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: CRUROTARSI SERENO & ARCUCCI, 1990
COHORT: CROCODYLOTARSI BENTON & CLARK, 1988
MAGNORDER: SUCHIA KREBS, 1974
SUPERORDER: CROCODYLOMORPHA WALKER, 1968
GRANDORDER: CROCODILIFORMES HAY, 1930
MICRORDER: MESOEUROCROCODYLIA WHETSTONE & WHYBROW, 1983
ORDER: MESOSUCHIA EMEND HUXLEY, 1875
PARVORDER: METASUCHIA BENTON & CLARK, 1988
GONDWANASUCHIA DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004
NOTOSUCHIMORPHA DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004
NOTOSUCHIFORMES DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004
TERRIASUCHIA DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004
INFRAORDER: NOTOSUCHIA GASPARINI, 1971
Incertae sedis
*Razanandrongobe sakalavae
_____________________________________________________________________________________
MADAGASCAR
MAHAJANGA BASIN
HILLS N-NW OF AMBONDROMAMY
BATHONIAN STAGE
MIDDLE DOGGER EPOCH
MIDDLE JURASSIC
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
INFRACLASS: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: CRUROTARSI SERENO & ARCUCCI, 1990
COHORT: CROCODYLOTARSI BENTON & CLARK, 1988
MAGNORDER: SUCHIA KREBS, 1974
SUPERORDER: CROCODYLOMORPHA WALKER, 1968
GRANDORDER: CROCODILIFORMES HAY, 1930
MICRORDER: MESOEUROCROCODYLIA WHETSTONE & WHYBROW, 1983
ORDER: MESOSUCHIA EMEND HUXLEY, 1875
PARVORDER: METASUCHIA BENTON & CLARK, 1988
GONDWANASUCHIA DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004
NOTOSUCHIMORPHA DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004
NOTOSUCHIFORMES DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004
TERRIASUCHIA DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004
INFRAORDER: NOTOSUCHIA GASPARINI, 1971
Incertae sedis
Razanandrongobe sakalavae
_____________________________________________________________________________________
MADAGASCAR
MAHAJANGA BASIN
UNDETERMINED LOCALITIES
BATHONIAN STAGE
MIDDLE DOGGER EPOCH
MIDDLE JURASSIC
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
INFRACLASS: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: CRUROTARSI SERENO & ARCUCCI, 1990
COHORT: CROCODYLOTARSI BENTON & CLARK, 1988
MAGNORDER: SUCHIA KREBS, 1974
SUPERORDER: CROCODYLOMORPHA WALKER, 1968
GRANDORDER: CROCODILIFORMES HAY, 1930
MICRORDER: MESOEUROCROCODYLIA WHETSTONE & WHYBROW, 1983
ORDER: MESOSUCHIA EMEND HUXLEY, 1875
PARVORDER: METASUCHIA BENTON & CLARK, 1988
GONDWANASUCHIA DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004
NOTOSUCHIMORPHA DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004
NOTOSUCHIFORMES DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004
TERRIASUCHIA DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004
INFRAORDER: NOTOSUCHIA GASPARINI, 1971
Incertae sedis
Razanandrongobe sakalavae
_____________________________________________________________________________________
MADAGASCAR
AMBILOBE
MAHAJANGA
MORONDAVA
AMBONDROMAMY
AMBONDROMAMY FAUNA
ISALO IIIB
BATHONIAN STAGE
UPPER MIDDLE DOGGER EPOCH
MIDDLE JURASSIC
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: AMPHIBIA LINNAEUS, 1758
SUBCLASS: LISSAMPHIBIA, HAECKEL, 1866?
Incertae sedis
Gen. sp indet.
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: CHELONOMORPHA KUHN, 1960
SUPERCOHORT: TESTUDINES LINNAEUS, 1758
Incertae sedis
Gen. sp indet.
SUBCLASS: DIAPSIDA OSBORN, 1903
INFRACLASS: NEODIAPSIDA BENTON, 1985
DIVISION: LEPIDOSAUROMORPHA BENTON, 1983
SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)
MAGNORDER: SPHENODONTIFORMES Hay, 1930
ORDER: SPHENODONTIA WILLISTON, 1925
Family: SPHENODONTIDAE Cope, 1869
Gen. sp indet.
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: CRUROTARSI SERENO & ARCUCCI, 1990
COHORT: CROCODYLOTARSI BENTON & CLARK, 1988
MAGNORDER: SUCHIA KREBS, 1974
SUPERORDER: CROCODYLOMORPHA WALKER, 1968
GRANDORDER: CROCODILIFORMES HAY, 1930
MICRORDER: MESOEUROCROCODYLIA WHETSTONE & WHYBROW, 1983
ORDER: MESOSUCHIA EMEND HUXLEY, 1875
PARVORDER: METASUCHIA BENTON & CLARK, 1988
INFRAORDER: METAMESOSUCHIA HULKE, 1878
NEOSUCHIA CLARK, IN BENTON & CLARK, 1988
Family: ATOPOSAURIDAE Gervais, 1871
Gen. sp indet.
SUPERCOHORT: AVEMETATARSALIA BENTON, 1999
COHORT: PROLACERTIFORMES CAMP, 1945 NOVA
SUPERORDER: TAPINOPLATIA PETERS, 2000
GRANDORDER: CHARACIOPODA PETERS, 2000
MICRORDER: FENESTRASAURIA PETERS, 2000
ORDER: PTEROSAURIA KAUP, 1834
SUBORDER: RHAMPHORHYNCHOIDEA PLIENINGER, 1901
Incertae sedis
Gen. sp indet.
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
ORDER: Incertae sedis
Gen. sp indet.
COHORT: ORNITHISCHIFORMES COOPER, 1985
SUPERORDER: SAUROPODOMORPHA HUENE, 1932
GRANDORDER: BRONTOSAURIA OLSHEVSKY, 1991 (emend)
MICRORDER: SAUROPODA MARSH, 1878
ORDER: EUSAUROPODA
SUBORDER: NEOSAUROPODA BONAPARTE, 1986
Incertae sedis
Gen. sp indet.
MAGNORDER: ORNITHISCHIA SEELEY, 1888
Incertae sedis
Gen. sp indet.
_____________________________________________________________________________________
NORTHERN MADAGASCAR
MAJUNGA BASIN
NEAR ANDRANOMANY
MANARY-ABO
BATHONIAN STAGE
MIDDLE DOGGER EPOCH
MIDDLE JURASSIC
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: ORNITHISCHIFORMES COOPER, 1985
SUPERORDER: SAUROPODOMORPHA HUENE, 1932
GRANDORDER: BRONTOSAURIA OLSHEVSKY, 1991 (emend)
MICRORDER: SAUROPODA MARSH, 1878
ORDER: EUSAUROPODA
Incertae sedis
non Bothriospondylus, B?madagascariensis
_____________________________________________________________________________________
MADAGASCAR
MAHAJANGA BASIN
FROM AN AREA BETWEEN AMBONDROMAMY AND PORT BERGE
BATHONIAN STAGE
MIDDLE DOGGER EPOCH
MIDDLE JURASSIC
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
Incertae sedis
Morphotype 1
Gen. sp indet.
Morphotype 2
Gen. sp indet.
Morphotype 3
Gen. sp indet.
Morphotype 4
Gen. sp indet.
Morphotype 5
Gen. sp indet.
Morphotype 6
Gen. sp indet.
Morphotype 7
Gen. sp indet.
_____________________________________________________________________________________
MADAGASCAR
MAHAJANGA BASIN
NORTH OF PORT-BERGE
BATHONIAN STAGE
MIDDLE DOGGER EPOCH
MIDDLE JURASSIC
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
Incertae sedis
Morphotype 1
Gen. sp indet.
Morphotype 2
Gen. sp indet.
Morphotype 3
Gen. sp indet.
Morphotype 4
Gen. sp indet.
Morphotype 5
Gen. sp indet.
Morphotype 6
Gen. sp indet.
_____________________________________________________________________________________
MADAGASCAR
MAHAJANGA BASIN
BETWEEN TSINJORAN AND THE BRIDGE ON THE KAMORO RIVER
BATHONIAN STAGE
MIDDLE DOGGER EPOCH
MIDDLE JURASSIC
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
Incertae sedis
Morphotype 8
Gen. sp indet.
_____________________________________________________________________________________
MADAGASCAR
MAHAJANGA BASIN
NEAR THE BRIDGE ON THE KAMORO RIVER ALONG THE WEATHERED SIDES OF A LOW TANETY (LOCAL NAME FOR "HILL") ON THE RIGHT BANK OF THE KAMORO RIVER
BATHONIAN STAGE
MIDDLE DOGGER EPOCH
MIDDLE JURASSIC
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
Incertae sedis
Gen. sp indet.
_____________________________________________________________________________________
MADAGASCAR
MAHAJANGA BASIN
SOUTH-WEST TO AMBONDROMAMY
FROM AN AREA NEAR TSINJORANO
BATHONIAN STAGE
MIDDLE DOGGER EPOCH
MIDDLE JURASSIC
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
Incertae sedis
Morphotype 2
Gen. sp indet.
Morphotype 3
Gen. sp indet.
Morphotype 4
Gen. sp indet.
Morphotype 6
Morphotype 8
Gen. sp indet.
_____________________________________________________________________________________
NORTHWEST MADAGASCAR
MAHAJANGA PROVINCE
UNDETERMINED HILLS WEST OF AMBONDROMAMY
ISALO IIIB SUBUNIT
'FACIES MIXTE DINOSAURIENS’
BATHONIAN STAGE
MIDDLE DOGGER EPOCH
MIDDLE JURASSIC
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
ARCHOSAURIA INCERTAE SEDIS
*Razanadrongobe sakalavae
_____________________________________________________________________________________
NORTHWEST MADAGASCAR
MAHAJANGA PROVINCE
HILLS NNW OF ANDRANOMAMY
ISALO IIIB SUBUNIT
'FACIES MIXTE DINOSAURIENS’
BATHONIAN STAGE
MIDDLE DOGGER EPOCH
MIDDLE JURASSIC
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
ARCHOSAURIA INCERTAE SEDIS
Razanadrongobe sakalavae
_____________________________________________________________________________________
N. W. MADAGASCAR
MAHAJANGA PROVINCE
8 KM (5 MILES) NORTH OF ANABORANO, AND 75 MILES SSW OF DIEGO SUAREZ
AMBARAKARAKA
MIDDLE SAKAMENA FORMATION
INDUAN STAGE
UPPER SCYTHIAN EPOCH
EARLY TRIASSIC
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: AMPHIBIA LINNAEUS, 1758
SUBCLASS: LABYRINTHODONTIA OWEN, 1860
SUPERORDER: TEMNOSPONDYLIA ZITTEL, 1887
MICROORDER: LIMNARCHIA YATES & WARREN, 2000
ORDER: STEREOSPONDYLOMORPHA YATES & WARREN, 2000
Family: LYDEKKERINIDAE Watson, 1919
*Deltacephalus whitei
_____________________________________________________________________________________
MAHAJANGA PROVINCE
JURASSIC?
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: ORNITHISCHIFORMES COOPER, 1985
SUPERORDER: SAUROPODOMORPHA HUENE, 1932
Incertae sedis
Gen. sp indet.
_____________________________________________________________________________________
MADAGASCAR
MAHAJANGA PROVINCE
MAJUNGA REGION
SW OF THE BASIN OF KELIFELY
KANDREHO FORMATION
MIDDLE OR UPPER TOARCIAN STAGE
UPPER LIAS EPOCH
EARLY JURASSIC
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: CRUROTARSI SERENO & ARCUCCI, 1990
COHORT: CROCODYLOTARSI BENTON & CLARK, 1988
MAGNORDER: SUCHIA KREBS, 1974
SUPERORDER: CROCODYLOMORPHA WALKER, 1968
GRANDORDER: CROCODILIFORMES HAY, 1930
MICRORDER: MESOEUROCROCODYLIA WHETSTONE & WHYBROW, 1983
ORDER: MESOSUCHIA EMEND HUXLEY, 1875
ORDER: THALATTOSUCHIA EMEND FRAAS, 1901
Family: TELEOSAURIDAE Geoffroy, 1831
Incertae sedis
Steneosaurus sp
_____________________________________________________________________________________
MADAGASCAR
MAJUNGA PROVINCE
MAJUNGA BASIN
MIDDLE JURASSIC
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: ORNITHISCHIFORMES COOPER, 1985
SUPERORDER: SAUROPODOMORPHA HUENE, 1932
GRANDORDER: BRONTOSAURIA OLSHEVSKY, 1991 (emend)
MICRORDER: SAUROPODA MARSH, 1878
Genus nova.
_____________________________________________________________________________________
NORTHWESTERN MADAGASAR
MAHAJANGA PROVINCE
ABOUT 150 KM SW OF THE COASTAL TOWN OF MAHAJANGA (MAJUNGA)
FROM THE VICINITY OF AMBONDROMAMY
ISALO IIIB FORMATION
BATHONIAN STAGE
MIDDLE DOGGER EPOCH
MIDDLE JURASSIC
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
AVEPODA PAUL, 2002
SUPERORDER: NEOTHEROPODA BAKKER, 1986
GRANDORDER: TETANURAE GAUTHIER, 1986
MICRORDER: COELUROSAURIA HUENE, 1914
ORDER: SEGNOSAURIA BARSBOLD & PERLE, 1980
Family: Nova?
*Archaeodontosaurus descouensi
_____________________________________________________________________________________
MADAGASCAR
MAHAJANGA PROVINCE
20 M EASTWARD OF THE BAY NARINDA
UNNAMED UNIT
BATHONIAN STAGE
MIDDLE DOGGER EPOCH
MIDDLE JURASSIC
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: ORNITHISCHIFORMES COOPER, 1985
SUPERORDER: SAUROPODOMORPHA HUENE, 1932
GRANDORDER: BRONTOSAURIA OLSHEVSKY, 1991 (emend)
MICRORDER: SAUROPODA MARSH, 1878
ORDER: EUSAUROPODA
Incertae sedis
non Bothriospondylus, B?madagascariensis
_____________________________________________________________________________________
MADAGASCAR
MAHAJANGA PROVINCE
MAHAJANGA BASIN
AMBONDROMANY REGION
ISALO III
BATHONIAN STAGE
MIDDLE DOGGER EPOCH
MIDDLE JURASSIC
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: MAMMALIA LINNAEUS, 1758
SUBCLASS: PROTOTHERIA GILL, 1872
ORDER: INCERTAE SEDIS
Incertae sedis
*Ambondro mahabo
_____________________________________________________________________________________
MAHAJANA PROVINCE
MAHAJANA PROVINCE
ANDRANOSAMONTA
LATE JURASSIC
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: ORNITHISCHIFORMES COOPER, 1985
SUPERORDER: SAUROPODOMORPHA HUENE, 1932
GRANDORDER: BRONTOSAURIA OLSHEVSKY, 1991 (emend)
MICRORDER: SAUROPODA MARSH, 1878
ORDER: EUSAUROPODA
Incertae sedis
non Bothriospondylus, B?madagascariensis
_____________________________________________________________________________________
MADAGASCAR
MAHAJANGA PROVINCE
ANKINGANIVALKA
LATE JURASSIC
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: ORNITHISCHIFORMES COOPER, 1985
SUPERORDER: SAUROPODOMORPHA HUENE, 1932
GRANDORDER: BRONTOSAURIA OLSHEVSKY, 1991 (emend)
MICRORDER: SAUROPODA MARSH, 1878
ORDER: EUSAUROPODA
Incertae sedis
non Bothriospondylus, B?madagascariensis
_____________________________________________________________________________________
MADAGASCAR
MAHAJANGA PROVINCE
PORT-BERGE
LATE JURASSIC
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: ORNITHISCHIFORMES COOPER, 1985
SUPERORDER: SAUROPODOMORPHA HUENE, 1932
GRANDORDER: BRONTOSAURIA OLSHEVSKY, 1991 (emend)
MICRORDER: SAUROPODA MARSH, 1878
ORDER: EUSAUROPODA
Incertae sedis
non Bothriospondylus, B?madagascariensis
_____________________________________________________________________________________
MADAGASCAR
MAHAJANGA PROVINCE
NE REPUBLICK MALGACHE
MAJUNGA
LATE JURASSIC
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: ORNITHISCHIFORMES COOPER, 1985
SUPERORDER: SAUROPODOMORPHA HUENE, 1932
GRANDORDER: BRONTOSAURIA OLSHEVSKY, 1991 (emend)
MICRORDER: SAUROPODA MARSH, 1878
ORDER: EUSAUROPODA
Incertae sedis
non Bothriospondylus, B?madagascariensis
_____________________________________________________________________________________
MADAGASCAR
MAHAJANGA PROVINCE
RIGHT BANK OF THE RIVER BETSIBOKA
MAEVARANO
CENOMANIAN STAGE
UPPERMIDDLE GALLIC SUBEPOCH
LOWERMOST GULF EPOCH
EARLY CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
AVEPODA PAUL, 2002
SUPERORDER: NEOTHEROPODA BAKKER, 1986
GRANDORDER: CERATOSAURIA NOVAS, 1992
AVEROSTRA PAUL, 2001
ORDER: CERATOSAURIA MARSH, 1884
Superfamily: ABELISAUROIDEA Novas, 1989
Family: ABELISAURIDAE Bonaparte & Novas, 1985
Subfamily: CARNOTAURINAE Sereno, 1998
Majungasaurus crenatissimus
_____________________________________________________________________________________
MADASCAR
MAHAJANGA PROVINCE
MAJUNGA
LOWER CENOMANIAN STAGE
UPPERMIDDLE GALLIC SUBEPOCH
LOWERMOST GULF EPOCH
EARLY CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: CRUROTARSI SERENO & ARCUCCI, 1990
COHORT: CROCODYLOTARSI BENTON & CLARK, 1988
MAGNORDER: SUCHIA KREBS, 1974
SUPERORDER: CROCODYLOMORPHA WALKER, 1968
GRANDORDER: CROCODILIFORMES HAY, 1930
MICRORDER: MESOEUROCROCODYLIA WHETSTONE & WHYBROW, 1983
ORDER: MESOSUCHIA EMEND HUXLEY, 1875
PARVORDER: METASUCHIA BENTON & CLARK, 1988
INFRAORDER: METAMESOSUCHIA HULKE, 1878
NEOSUCHIA CLARK, IN BENTON & CLARK, 1988
Family: TREMATOCHAMPSIDAE Buffetaut, 1974
*Miadanasuchus oblita
= Trematochampsa oblita
_____________________________________________________________________________________
MADAGASCAR
MAHAJANGA PROVINCE
RIGHT BANKS OF THE RIVER BETSIOBKA
MAEVARANO
MEME LOCAILTY
CENOMANIAN STAGE
UPPERMIDDLE GALLIC SUBEPOCH
LOWERMOST GULF EPOCH
EARLY CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
AVEPODA PAUL, 2002
SUPERORDER: NEOTHEROPODA BAKKER, 1986
GRANDORDER: CERATOSAURIA NOVAS, 1992
AVEROSTRA PAUL, 2001
ORDER: CERATOSAURIA MARSH, 1884
Superfamily: ABELISAUROIDEA Novas, 1989
Family: ABELISAURIDAE Bonaparte & Novas, 1985
Subfamily: CARNOTAURINAE Sereno, 1998
Majungasaurus crenatissimus
COHORT: ORNITHISCHIFORMES COOPER, 1985
SUPERORDER: SAUROPODOMORPHA HUENE, 1932
Incertae sedis
Gen. sp indet.
_____________________________________________________________________________________
MADAGASCAR
MAHAJANGA PROVINCE
MAHAJANGA BASIN
ANKAZOMIHABOKA SANDSTONE
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: CHELONOMORPHA KUHN, 1960
SUPERCOHORT: TESTUDINES LINNAEUS, 1758
COHORT: CASICHELYDIA emend GAFFNEY & MEYLAN, 1988
MAGNORDER: PLEURODIRA GAFFNEY & MEYLAN, 1988
ORDER: PLEURODIRIDA COPE, 1864
SUBORDER: EUPLEURODIRA GAFFNEY & MEYLAN, 1988
Superfamily: PELOMEDUSOIDES Cope, 1868 Cope, 1868
Family: PELOMEDUSIDAE Cope, 1868
Gen. sp indet.
SUBCLASS: DIAPSIDA OSBORN, 1903
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: CRUROTARSI SERENO & ARCUCCI, 1990
COHORT: CROCODYLOTARSI BENTON & CLARK, 1988
MAGNORDER: SUCHIA KREBS, 1974
SUPERORDER: CROCODYLOMORPHA WALKER, 1968
GRANDORDER: CROCODILIFORMES HAY, 1930
MICRORDER: MESOEUROCROCODYLIA WHETSTONE & WHYBROW, 1983
ORDER: MESOSUCHIA EMEND HUXLEY, 1875
PARVORDER: METASUCHIA BENTON & CLARK, 1988
INFRAORDER: METAMESOSUCHIA HULKE, 1878
NEOSUCHIA CLARK, IN BENTON & CLARK, 1988
Family: TREMATOCHAMPSIDAE Buffetaut, 1974
Incertae sedis
Gen. sp indet.
_____________________________________________________________________________________
MADAGASCAR
MAJUNGA PROVINCE
IN A VILLAGE NEAR MAJUNGA
2/3 KM NORTH OF HOTEL BERIVOTRO
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
AVEPODA PAUL, 2002
SUPERORDER: NEOTHEROPODA BAKKER, 1986
GRANDORDER: CERATOSAURIA NOVAS, 1992
AVEROSTRA PAUL, 2001
ORDER: CERATOSAURIA MARSH, 1884
Superfamily: ABELISAUROIDEA Novas, 1989
Family: ABELISAURIDAE Bonaparte & Novas, 1985
Subfamily: CARNOTAURINAE Sereno, 1998
Majungasaurus crenatissimus
= Megalosaurus crenatissimus
_____________________________________________________________________________________
MADAGASCAR
MAHAJANGA PROVINCE
MAHAJANGA BASIN
MAEVARANO FORMATION
CAMPANIAN STAGE?
SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: MAMMALIA LINNAEUS, 1758
SUBCLASS: PROTOTHERIA GILL, 1872
INFRACLASS: TRECHNOTHERIA McKENNA, 1975
LEGION: CLADOTHERIA McKENNA, 1975
SUBLEGION: ZATHERIA MCKENNA, 1975
INFRALEGION: INCERTAE SEDIS
Genus: Nova
_____________________________________________________________________________________
MADAGASCAR
MAHAJANGA PROVINCE
50 KM FROM MAJUNGA
REGION OF BERIVOTRA
UPPER SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: CHELONOMORPHA KUHN, 1960
SUPERCOHORT: TESTUDINES LINNAEUS, 1758
COHORT: CASICHELYDIA emend GAFFNEY & MEYLAN, 1988
MAGNORDER: PLEURODIRA GAFFNEY & MEYLAN, 1988
ORDER: PLEURODIRIDA COPE, 1864
SUBORDER: EUPLEURODIRA GAFFNEY & MEYLAN, 1988
Superfamily: PELOMEDUSOIDES Cope, 1868 Cope, 1868
Family: PELOMEDUSIDAE Cope, 1868
Gen. sp undescribed
Subfamily: PELOMEDUSINAE Cope, 1868
Bothremyididae gen sp indet
_____________________________________________________________________________________
MADAGASCAR
MAHAJANGA
MAJUNGA PROVINCE
BERIVOTRO
BERIVOTRO FORMATION
SENONIAN OR CAMPANIAN STAGE
SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
AVEPODA PAUL, 2002
SUPERORDER: NEOTHEROPODA BAKKER, 1986
GRANDORDER: CERATOSAURIA NOVAS, 1992
AVEROSTRA PAUL, 2001
ORDER: CERATOSAURIA MARSH, 1884
Superfamily: ABELISAUROIDEA Novas, 1989
Family: ABELISAURIDAE Bonaparte & Novas, 1985
Subfamily: CARNOTAURINAE Sereno, 1998
*Majungasaurus crenatissimus
= Megalosaurus crenatissimusCOHORT: ORNITHISCHIFORMES COOPER, 1985
SUPERORDER: SAUROPODOMORPHA HUENE, 1932
Incertae sedis
Gen. sp indet.
_____________________________________________________________________________________
MADAGASCAR
MAHAJANGA PROVINCE
MAHAJANGA BASIN
JUSE EAST OF BERIVOTRA VILLAGE
MAEVARENO SANDSTONE
CAMPANIAN STAGE?
SENONIAN SUBEPOCH
UPPER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: MAMMALIA LINNAEUS, 1758
SUBCLASS: INCERTAE SEDIS
Gen. sp indet.
SUBCLASS: THERIIFORMES (ROWE, 1988) McKENNA & BELL, 1997
INFRACLASS: ALLOTHERIA (MARSH, 1880) McKENNA & BELL, 1997
ORDER: MULTITUBERCULATA COPE, 1884
Incertae sedis
Gen. sp indet.
SUBORDER: GONDWANATHERIA Mones, 1987
Family: SUDAMERICICIDAE Scillato-Yane & Prasad, 1984
Lavanify miolaka
_____________________________________________________________________________________
N. W. MADAGASCAR
MAHAJANGA PROVINCE
MAHAJANGA BASIN
MAEVARANO FORMATION
CAMPANIAN STAGE?
SENONIAN SUBEPOCH
UPPER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
INFRACLASS: NEODIAPSIDA BENTON, 1985
LEGION: LEPIDOSAUROMORPHA BENTON, 1983
SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)
MAGNORDER: SQUAMATA, OPPEL, 1811
SUPERORDER: LACERTILIA OWEN, 1842
GRANDORDER: SCINCOGEKKONOMORPHA SUKHANOV, 1961
MICRORDER: SCINCOMORPHA CAMP, 1923
ORDER: LEPTOGLOSSA COPE, 1900
SUBORDER: ZONUROMORPHA ALIFANOV, 2000
INFRAORDER: ZONURIA FURBRINGER, 1900
Superfamily: CORDYLOIDEA Fritzinger, 1826
Family: CORDYLIDAE? Gray, 1837
Gen. sp indet.
_____________________________________________________________________________________
NORTH-WEST MADAGASCAR
MAHAJANGA PROVINCE
MAHAJANGA BASIN
APPROXIMATELY 1 KM SOUTH-EAST OF THE VILLAGE OF BERIVOTRA
MAEVARANO FORMATION
NEAR THE TOP OF THE CAMPANIAN STAGE?
SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: CRUROTARSI SERENO & ARCUCCI, 1990
COHORT: CROCODYLOTARSI BENTON & CLARK, 1988
MAGNORDER: SUCHIA KREBS, 1974
SUPERORDER: CROCODYLOMORPHA WALKER, 1968
GRANDORDER: CROCODILIFORMES HAY, 1930
MICRORDER: MESOEUROCROCODYLIA WHETSTONE & WHYBROW, 1983
SUBORDER: MESOEUROCROCODYLIA WHETSTONE & WHYBROW, 1983
MESOEUROCRODYLIA: NEW FAMILES?
Also a long snouted form.
Incertae sedis
GONDWANASUCHIA DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004
PEIROSAURIMORPHA DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004
PEIROSAURIFORMES DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004
Superfamily: PEIROSAURIDAE Gasparini, 1982 emend de Souza Carvalho, Ribeiro & dos Santos Avilla, 2004
Family: PEIROSAURIDAE Gasparini, 1982
Subfamily: PEIROSAURIDAE Gasparini, 1982
Gen. nova?
NEOSUCHIA CLARK, IN BENTON & CLARK, 1988
Family: TREMATOCHAMPSIDAE Buffetaut, 1974
Trematochampsa oblita
_____________________________________________________________________________________
MADAGASCAR
MAJUNGA PROVINCE
46 KM S OF MAJUNGA
BETSIBOKA
“GRES DE MAEVARENO”
CAMPANIAN STAGE
SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: ORNITHISCHIFORMES COOPER, 1985
SUPERORDER: SAUROPODOMORPHA HUENE, 1932
GRANDORDER: BRONTOSAURIA OLSHEVSKY, 1991 (emend)
MICRORDER: SAUROPODA MARSH, 1878
ORDER: EUSAUROPODA
SUBORDER: NEOSAUROPODA BONAPARTE, 1986
INFRAORDER: MACRONARIA WILSON & SERENO, 1998
PARVORDER: TITANOSAURIFORMES SALGADO ET AL, 1997
Superfamily: TITANOSAUROIDEA Lydekker, 1885 (sensu Upchurch, 1999)
Family: TITANOSAURIDAE Lydekker, 1885
Incertae sedis
non Titanosaurus, T? madagascariensis
= Laplatasaurus madagascariensis
_____________________________________________________________________________________
MADAGASCAR
MAJUNGA PROVINCE
‘GRES DE MAEVANO’
PROBABLY CAMPANIAN STAGE
SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
AVEPODA PAUL, 2002
SUPERORDER: NEOTHEROPODA BAKKER, 1986
GRANDORDER: CERATOSAURIA NOVAS, 1992
AVEROSTRA PAUL, 2001
ORDER: CERATOSAURIA MARSH, 1884
Superfamily: ABELISAUROIDEA Novas, 1989
Family: ABELISAURIDAE Bonaparte & Novas, 1985
Subfamily: MAJUNGASAURINAE Tortosa, Buffetaut, Vialle, Dutour, Turini, & Cheylan, 2014
*Majungatholus atopus
_____________________________________________________________________________________
MADAGASCAR
MAJUNGA PROVINCE
MAHAJANGA BASIN
CAMPANIAN STAGE?
SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
AVEPODA PAUL, 2002
SUPERORDER: NEOTHEROPODA BAKKER, 1986
GRANDORDER: CERATOSAURIA NOVAS, 1992
AVEROSTRA PAUL, 2001
ORDER: CERATOSAURIA MARSH, 1884
Superfamily: ABELISAUROIDEA Novas, 1989
Family: ABELISAURIDAE Bonaparte & Novas, 1985
Subfamily: CARNOTAURINAE Sereno, 1998
Majungasaurus crenatissimus
= Megalosaurus crenatissimus
_____________________________________________________________________________________
MADAGASCAR
MAHAJANGA PROVINCE
MAHAJANGA BASIN
S TO THE RN4 AND BERIVOTRA VILLAGE
ANEMBALEMBA MEMBER
MAEVARANO FORMATION
MAASTRICHTIAN STAGE
UPPERMOST SENONIAN SUBEPOCH
UPPERMOST GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
AVEPODA PAUL, 2002
SUPERORDER: NEOTHEROPODA BAKKER, 1986
GRANDORDER: CERATOSAURIA NOVAS, 1992
AVEROSTRA PAUL, 2001
ORDER: CERATOSAURIA MARSH, 1884
Superfamily: ABELISAUROIDEA Bonaparte & Novas, 1985
Family: ABELISAURIDAE Bonaparte & Novas, 1985
Gen. sp indet.
_____________________________________________________________________________________
MADAGASCAR
MAHAJANGA PROVINCE
MAHAJANGA BASIN
ABOUT 500 M NE TO THE BERIVOTRA VILLAGE
LEFT BANK OF A SMALL STREAM "VAVARANONANTASHALVA"
ANEMBALEMBA MEMBER
MAEVARANO FORMATION
MAASTRICHTIAN STAGE
UPPERMOST SENONIAN SUBEPOCH
UPPERMOST GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
AVEPODA PAUL, 2002
SUPERORDER: NEOTHEROPODA BAKKER, 1986
GRANDORDER: CERATOSAURIA NOVAS, 1992
AVEROSTRA PAUL, 2001
ORDER: CERATOSAURIA MARSH, 1884
Superfamily: ABELISAUROIDEA Bonaparte & Novas, 1985
Family: ABELISAURIDAE Bonaparte & Novas, 1985
Gen. sp indet.
_____________________________________________________________________________________
MADAGASCAR
MAHAJANGA PROVINCE
MAHAJANGA BASIN
ANEMBALEMBA MEMBER
MAEVARANO FORMATION
MAASTRICHTIAN STAGE
SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: AVES LINNAEUS, 1758
Incertae sedis
Gen. sp indet.
CLASS: MAMMALIA LINNAEUS, 1758
SUBCLASS: PROTOTHERIA GILL, 1872
INFRACLASS: TRECHNOTHERIA McKENNA, 1975
LEGION: CLADOTHERIA McKENNA, 1975
SUBLEGION: ZATHERIA MCKENNA, 1975
INFRALEGION: TRIBOSPHENIDA (McKENNA, 1975) McKENNA & BELL, 1997
SUPERCOHORT: THERIA (PARKER & HASWELL, 1987) McKENNA & BELL, 1997
COHORT: PLACENTALIA (OWEN, 1837) McKENNA & BELL, 1997
MAGNORDER: EPITHERIA (McKENNA, 1975) McKENNA & BELL, 1997
SUPERORDER: PREPTOTHERIA McKENNA, 1975
GRANDORDER: UNGULATOMORPHA ARCHIBALD, 1996
Family: ZHELESTIDAE Nesov, 1990
Gen. sp indet.
_____________________________________________________________________________________
MADAGASCAR
MAJUNGA PROVINCE
MAHAJANGA BASIN
MAEVARANO FORMATION
ANEMBALEMBA MEMBER
MAASTRICHTIAN STAGE
SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: ORNITHISCHIFORMES COOPER, 1985
SUPERORDER: SAUROPODOMORPHA HUENE, 1932
GRANDORDER: BRONTOSAURIA OLSHEVSKY, 1991 (emend)
MICRORDER: SAUROPODA MARSH, 1878
ORDER: EUSAUROPODA
SUBORDER: NEOSAUROPODA BONAPARTE, 1986
INFRAORDER: MACRONARIA WILSON & SERENO, 1998
PARVORDER: TITANOSAURIFORMES SALGADO ET AL, 1997
Superfamily: TITANOSAUROIDEA Lydekker, 1885 (sensu Upchurch, 1999)
Family: TITANOSAURIDAE Lydekker, 1885
Subfamily: SALTASAURINAE Powell, 1986 vide Bonaparte, 1987
Gen. sp indet.
_____________________________________________________________________________________
MADAGASCAR
MAJUNGA PROVINCE
MAHAJANGA BASIN
NEAR THE VILLAGE OF BERIVOTRA
MAEVARANO FORMATION
ANEMBALEMBA MEMBER
MAASTRICHTIAN STAGE
SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
AVEPODA PAUL, 2002
SUPERORDER: NEOTHEROPODA BAKKER, 1986
GRANDORDER: CERATOSAURIA NOVAS, 1992
AVEROSTRA PAUL, 2001
ORDER: CERATOSAURIA MARSH, 1884
Superfamily: ABELISAUROIDEA Novas, 1989
Family: NOASAURIDAE Bonaparte & Powell, 1980
Subfamily: VELOCISAURIINAE Bonaparte, 1991 (emend AGNOLIN, NOVAS & APESTEGUIA, 2003)
*Masiakasaurus knopfleri
_____________________________________________________________________________________
NORTHEASTERN MADAGASCAR
MAHAJANA PROVINCE
MAHAJANGA BASIN
BERIVOTRA
ANEBALEMBA MEMBER
MAEVARANO FORMATION
MAASTRICHTIAN STAGE
UPPERMOST SENONIAN SUBEPOCH
UPPERMOST GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
AVEPODA PAUL, 2002
SUPERORDER: NEOTHEROPODA BAKKER, 1986
GRANDORDER: CERATOSAURIA NOVAS, 1992
AVEROSTRA PAUL, 2001
ORDER: CERATOSAURIA MARSH, 1884
Superfamily: ABELISAUROIDEA Novas, 1989
Family: ABELISAURIDAE Bonaparte & Novas, 1985
Subfamily: MAJUNGASAURINAE Tortosa, Buffetaut, Vialle, Dutour, Turini, & Cheylan, 2014
Majungatholus atopus
Family: NOASAURIDAE Bonaparte & Powell, 1980
Subfamily: VELOCISAURIINAE Bonaparte, 1991 (emend AGNOLIN, NOVAS & APESTEGUIA, 2003)
Masiakasaurus knopfleri
GRANDORDER: TETANURAE GAUTHIER, 1986
MICRORDER: COELUROSAURIA HUENE, 1914
ORDER: MANIRAPTORFORMES HOLTZ, 1996
AVETHEROPOA PAUL, 2002
AVEPECTORA PAUL, 2002
SUBORDER: MANIRAPTORA GAUTHIER, 1986
INFRAORDER: PARVES SERENO, 1997
PARVORDER: DEINONYCHOSAURIA COLBERT & D. A. RUSSELL, 1969
Family: DROMAEOSAURIDAE Russell, 1969
Gen. sp indet.
_____________________________________________________________________________________
MADAGASCAR
MAJUNGA PROVINCE
MAHAJANGA BASIN
UNSPECIFIED LOCALITY IN THE BERIVOTRA STUDY AREA
MAEVARANO FORMATION
ANEMBALEMBA MEMBER
MAASTRICHTIAN STAGE
SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
AVEPODA PAUL, 2002
SUPERORDER: NEOTHEROPODA BAKKER, 1986
GRANDORDER: CERATOSAURIA NOVAS, 1992
AVEROSTRA PAUL, 2001
ORDER: CERATOSAURIA MARSH, 1884
Superfamily: ABELISAUROIDEA Novas, 1989
Family: ABELISAURIDAE Bonaparte & Novas, 1985
Subfamily: MAJUNGASAURINAE Tortosa, Buffetaut, Vialle, Dutour, Turini, & Cheylan, 2014
Majungatholus atopus
_____________________________________________________________________________________
MADAGASCAR
MAJUNGA PROVINCE
MAHAJANGA BASIN
UNSPECIFIED LOCALITY IN THE 'MAJUNGA DISTRICT'
MAEVARANO FORMATION
ANEMBALEMBA MEMBER
MAASTRICHTIAN STAGE
SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
AVEPODA PAUL, 2002
SUPERORDER: NEOTHEROPODA BAKKER, 1986
GRANDORDER: CERATOSAURIA NOVAS, 1992
AVEROSTRA PAUL, 2001
ORDER: CERATOSAURIA MARSH, 1884
Superfamily: ABELISAUROIDEA Novas, 1989
Family: ABELISAURIDAE Bonaparte & Novas, 1985
Subfamily: MAJUNGASAURINAE Tortosa, Buffetaut, Vialle, Dutour, Turini, & Cheylan, 2014
Majungatholus atopus
_____________________________________________________________________________________
MADAGASCAR
MAHAJANGA BASIN
NORTH OF BERIVORTA
"GITE DU GUIDE"
MAEVARANO FORMATION
MAASTRICHTIAN STAGE
UPPER SENONIAN SUBEPOCH
UPPER UPPER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
INFRACLASS: NEODIAPSIDA BENTON, 1985
LEGION: LEPIDOSAUROMORPHA BENTON, 1983
SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)
MAGNORDER: SQUAMATA, OPPEL, 1811
SUPERORDER: SERPENTES LINNAEUS, 1759
ORDER: OPHIDIA BRONGNIART, 1800
SUBORDER: ALETHINOPHIDIA NOPCSCA, 1923
Superfamily: BOOIDEA Gray, 1825
Subfamily: BOOIDINAE Gray, 1825
Infrafamily: BOOINIEA Gray, 1925
Tribe: MATSOINI Hoffstetter, 1961
*Madtsoia madagascariensis
_____________________________________________________________________________________
MADAGASCAR
MAJUNGA PROVINCE
MAHAJANGA BASIN
"EAST OF MAMMAL"MAEVARANO FORMATION
ANEMBALEMBA MEMBER
MAASTRICHTIAN STAGE
SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
AVEPODA PAUL, 2002
SUPERORDER: NEOTHEROPODA BAKKER, 1986
GRANDORDER: CERATOSAURIA NOVAS, 1992
AVEROSTRA PAUL, 2001
ORDER: CERATOSAURIA MARSH, 1884
Superfamily: ABELISAUROIDEA Novas, 1989
Family: NOASAURIDAE Bonaparte & Powell, 1980
Subfamily: VELOCISAURIINAE Bonaparte, 1991 (emend AGNOLIN, NOVAS & APESTEGUIA, 2003)
Masiakasaurus knopfleri
_____________________________________________________________________________________
MADAGASCAR
MAHAJANGA BASIN
SOUTH OF BERIVORTA
MNHN MAJ 8
MAEVARANO FORMATION
MAASTRICHTIAN STAGE
UPPER SENONIAN SUBEPOCH
UPPER UPPER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
INFRACLASS: NEODIAPSIDA BENTON, 1985
LEGION: LEPIDOSAUROMORPHA BENTON, 1983
SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)
MAGNORDER: SQUAMATA, OPPEL, 1811
SUPERORDER: SERPENTES LINNAEUS, 1759
ORDER: OPHIDIA BRONGNIART, 1800
SUBORDER: ALETHINOPHIDIA NOPCSCA, 1923
Superfamily: BOOIDEA Gray, 1825
Subfamily: BOOIDINAE Gray, 1825
Infrafamily: BOOINIEA Gray, 1925
Tribe: MATSOINI Hoffstetter, 1961
Madtsoia madagascariensis
_____________________________________________________________________________________
MADAGASCAR
MAHAJANGA BASIN
SOUTH OF BERIVORTA
MNHN MAJ 9
MAEVARANO FORMATION
MAASTRICHTIAN STAGE
UPPER SENONIAN SUBEPOCH
UPPER UPPER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
INFRACLASS: NEODIAPSIDA BENTON, 1985
LEGION: LEPIDOSAUROMORPHA BENTON, 1983
SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)
MAGNORDER: SQUAMATA, OPPEL, 1811
SUPERORDER: SERPENTES LINNAEUS, 1759
ORDER: OPHIDIA BRONGNIART, 1800
SUBORDER: ALETHINOPHIDIA NOPCSCA, 1923
Superfamily: BOOIDEA Gray, 1825
Subfamily: BOOIDINAE Gray, 1825
Infrafamily: BOOINIEA Gray, 1925
Tribe: MATSOINI Hoffstetter, 1961
Madtsoia madagascariensis
_____________________________________________________________________________________
MADAGASCAR
MAHAJANGA BASIN
SOUTH OF BERIVORTA
MNHN MAJ 10
MAEVARANO FORMATION
MAASTRICHTIAN STAGE
UPPER SENONIAN SUBEPOCH
UPPER UPPER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
INFRACLASS: NEODIAPSIDA BENTON, 1985
LEGION: LEPIDOSAUROMORPHA BENTON, 1983
SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)
MAGNORDER: SQUAMATA, OPPEL, 1811
SUPERORDER: SERPENTES LINNAEUS, 1759
ORDER: OPHIDIA BRONGNIART, 1800
SUBORDER: ALETHINOPHIDIA NOPCSCA, 1923
Superfamily: BOOIDEA Gray, 1825
Subfamily: BOOIDINAE Gray, 1825
Infrafamily: BOOINIEA Gray, 1925
Tribe: MATSOINI Hoffstetter, 1961
Madtsoia madagascariensis
_____________________________________________________________________________________
MADAGASCAR
MAHAJANGA BASIN
NORTH OF MAHAJANGA-AMBALABE ROAD, BETWEEN KM 20 AND 25
MNHN MAJ 6
MAEVARANO FORMATION
MAASTRICHTIAN STAGE
UPPER SENONIAN SUBEPOCH
UPPER UPPER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
INFRACLASS: NEODIAPSIDA BENTON, 1985
LEGION: LEPIDOSAUROMORPHA BENTON, 1983
SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)
MAGNORDER: SQUAMATA, OPPEL, 1811
SUPERORDER: SERPENTES LINNAEUS, 1759
ORDER: OPHIDIA BRONGNIART, 1800
SUBORDER: ALETHINOPHIDIA NOPCSCA, 1923
Superfamily: BOOIDEA Gray, 1825
Subfamily: BOOIDINAE Gray, 1825
Infrafamily: BOOINIEA Gray, 1925
Tribe: MATSOINI Hoffstetter, 1961
Madtsoia madagascariensis
_____________________________________________________________________________________
MADAGASCAR
MAHAJANGA BASIN
NORTH OF MAHAJANGA-AMBALABE ROAD, BETWEEN KM 20 AND 25
MNHN MAJ 7
MAEVARANO FORMATION
MAASTRICHTIAN STAGE
UPPER SENONIAN SUBEPOCH
UPPER UPPER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
INFRACLASS: NEODIAPSIDA BENTON, 1985
LEGION: LEPIDOSAUROMORPHA BENTON, 1983
SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)
MAGNORDER: SQUAMATA, OPPEL, 1811
SUPERORDER: SERPENTES LINNAEUS, 1759
ORDER: OPHIDIA BRONGNIART, 1800
SUBORDER: ALETHINOPHIDIA NOPCSCA, 1923
Superfamily: BOOIDEA Gray, 1825
Subfamily: BOOIDINAE Gray, 1825
Infrafamily: BOOINIEA Gray, 1925
Tribe: MATSOINI Hoffstetter, 1961
Madtsoia madagascariensis
_____________________________________________________________________________________
MADAGASCAR
MAJUNGA PROVINCE
MAHAJANGA BASIN
MAD 01-03
MAEVARANO FORMATION
ANEMBALEMBA MEMBER
MAASTRICHTIAN STAGE
SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
INFRACLASS: NEODIAPSIDA BENTON, 1985
LEGION: LEPIDOSAUROMORPHA BENTON, 1983
SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)
MAGNORDER: SQUAMATA, OPPEL, 1811
SUPERORDER: SERPENTES LINNAEUS, 1759
ORDER: OPHIDIA BRONGNIART, 1800
SUBORDER: ALETHINOPHIDIA NOPCSCA, 1923
Superfamily: BOOIDEA Gray, 1825
Subfamily: BOOIDINAE Gray, 1825
Infrafamily: BOOINIEA Gray, 1925
Tribe: MATSOINI Hoffstetter, 1961
Madtsoia madagascariensis
_____________________________________________________________________________________
MADAGASCAR
MAJUNGA PROVINCE
MAHAJANGA BASIN
MAD 01-05
MAEVARANO FORMATION
ANEMBALEMBA MEMBER
MAASTRICHTIAN STAGE
SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
AVEPODA PAUL, 2002
SUPERORDER: NEOTHEROPODA BAKKER, 1986
GRANDORDER: CERATOSAURIA NOVAS, 1992
AVEROSTRA PAUL, 2001
ORDER: CERATOSAURIA MARSH, 1884
Superfamily: ABELISAUROIDEA Novas, 1989
Family: ABELISAURIDAE Bonaparte & Novas, 1985
Subfamily: CARNOTAURINAE Sereno, 1998
Majungatholus atopus
_____________________________________________________________________________________
MADAGASCAR
MAJUNGA PROVINCE
MAHAJANGA BASIN
MAD 03-03
MAEVARANO FORMATION
ANEMBALEMBA MEMBER
MAASTRICHTIAN STAGE
SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
INFRACLASS: NEODIAPSIDA BENTON, 1985
LEGION: LEPIDOSAUROMORPHA BENTON, 1983
SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)
MAGNORDER: SQUAMATA, OPPEL, 1811
SUPERORDER: SERPENTES LINNAEUS, 1759
ORDER: OPHIDIA BRONGNIART, 1800
SUBORDER: ALETHINOPHIDIA NOPCSCA, 1923
Superfamily: BOOIDEA Gray, 1825
Subfamily: BOOIDINAE Gray, 1825
Infrafamily: BOOINIEA Gray, 1925
Tribe: MATSOINI Hoffstetter, 1961
Madtsoia madagascariensis
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: CRUROTARSI SERENO & ARCUCCI, 1990
COHORT: CROCODYLOTARSI BENTON & CLARK, 1988
MAGNORDER: SUCHIA KREBS, 1974
SUPERORDER: CROCODYLOMORPHA WALKER, 1968
GRANDORDER: CROCODILIFORMES HAY, 1930
MICRORDER: MESOEUROCROCODYLIA WHETSTONE & WHYBROW, 1983
ORDER: MESOSUCHIA EMEND HUXLEY, 1875
PARVORDER: METASUCHIA BENTON & CLARK, 1988
GONDWANASUCHIA DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004
NOTOSUCHIMORPHA DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004
NOTOSUCHIFORMES DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004
TERRIASUCHIA DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004
INFRAORDER: NOTOSUCHIA GASPARINI, 1971
Family: NOTOSUCHIDAE Dollo, 1924
Simosuchus clarki
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
AVEPODA PAUL, 2002
SUPERORDER: NEOTHEROPODA BAKKER, 1986
GRANDORDER: CERATOSAURIA NOVAS, 1992
AVEROSTRA PAUL, 2001
ORDER: CERATOSAURIA MARSH, 1884
Superfamily: ABELISAUROIDEA Novas, 1989
Family: NOASAURIDAE Bonaparte & Powell, 1980
Subfamily: VELOCISAURIINAE Bonaparte, 1991 (emend AGNOLIN, NOVAS & APESTEGUIA, 2003)
Masiakasaurus knopfleri
_____________________________________________________________________________________
MADAGASCAR
MAJUNGA PROVINCE
MAHAJANGA BASIN
MAD 03-04
MAEVARANO FORMATION
ANEMBALEMBA MEMBER
MAASTRICHTIAN STAGE
SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
INFRACLASS: NEODIAPSIDA BENTON, 1985
LEGION: LEPIDOSAUROMORPHA BENTON, 1983
SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)
MAGNORDER: SQUAMATA, OPPEL, 1811
SUPERORDER: SERPENTES LINNAEUS, 1759
ORDER: OPHIDIA BRONGNIART, 1800
SUBORDER: ALETHINOPHIDIA NOPCSCA, 1923
Superfamily: BOOIDEA Gray, 1825
Subfamily: BOOIDINAE Gray, 1825
Infrafamily: BOOINIEA Gray, 1925
Tribe: MATSOINI Hoffstetter, 1961
Madtsoia madagascariensis
_____________________________________________________________________________________
MADAGASCAR
MAJUNGA PROVINCE
MAHAJANGA BASIN
MAD 03-05
MAEVARANO FORMATION
ANEMBALEMBA MEMBER
MAASTRICHTIAN STAGE
SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: AMPHIBIA LINNAEUS, 1758
SUBCLASS: LISSAMPHIBIA, HAECKEL, 1866?
SUPERORDER: SALIENTIA LAURENTI, 1768
ORDER: ANURA RAFINESQUE, 1815
SUBORDER: NEOBATRACHIA REIG, 1958
Superfamily: HYLOIDEA Rafinesque, 1815 (sensu Wied, 1856)
Family: LEPTODACTYLIDAE Werner, 1896
Subfamily: CERATOPHYRINAE Tschudi, 1838
Beelzebufo ampinga
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
INFRACLASS: NEODIAPSIDA BENTON, 1985
LEGION: LEPIDOSAUROMORPHA BENTON, 1983
SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)
MAGNORDER: SQUAMATA, OPPEL, 1811
SUPERORDER: SERPENTES LINNAEUS, 1759
ORDER: OPHIDIA BRONGNIART, 1800
SUBORDER: ALETHINOPHIDIA NOPCSCA, 1923
Superfamily: BOOIDEA Gray, 1825
Subfamily: BOOIDINAE Gray, 1825
Infrafamily: BOOINIEA Gray, 1925
Tribe: MATSOINI Hoffstetter, 1961
Madtsoia madagascariensis
_____________________________________________________________________________________
MADAGASCAR
MAJUNGA PROVINCE
MAHAJANGA BASIN
MAD 03-09
MAEVARANO FORMATION
ANEMBALEMBA MEMBER
MAASTRICHTIAN STAGE
SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
INFRACLASS: NEODIAPSIDA BENTON, 1985
LEGION: LEPIDOSAUROMORPHA BENTON, 1983
SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)
MAGNORDER: SQUAMATA, OPPEL, 1811
SUPERORDER: SERPENTES LINNAEUS, 1759
ORDER: OPHIDIA BRONGNIART, 1800
SUBORDER: ALETHINOPHIDIA NOPCSCA, 1923
Superfamily: BOOIDEA Gray, 1825
Subfamily: BOOIDINAE Gray, 1825
Infrafamily: BOOINIEA Gray, 1925
Tribe: MATSOINI Hoffstetter, 1961
Madtsoia madagascariensis
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: CRUROTARSI SERENO & ARCUCCI, 1990
COHORT: CROCODYLOTARSI BENTON & CLARK, 1988
MAGNORDER: SUCHIA KREBS, 1974
SUPERORDER: CROCODYLOMORPHA WALKER, 1968
GRANDORDER: CROCODILIFORMES HAY, 1930
MICRORDER: MESOEUROCROCODYLIA WHETSTONE & WHYBROW, 1983
ORDER: MESOSUCHIA EMEND HUXLEY, 1875
PARVORDER: METASUCHIA BENTON & CLARK, 1988
GONDWANASUCHIA DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004
NOTOSUCHIMORPHA DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004
NOTOSUCHIFORMES DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004
TERRIASUCHIA DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004
INFRAORDER: NOTOSUCHIA GASPARINI, 1971
Family: NOTOSUCHIDAE Dollo, 1924
Simosuchus clarki
_____________________________________________________________________________________
MADAGASCAR
MAJUNGA PROVINCE
MAHAJANGA BASIN
MAD 03-10
MAEVARANO FORMATION
ANEMBALEMBA MEMBER
MAASTRICHTIAN STAGE
SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: AMPHIBIA LINNAEUS, 1758
SUBCLASS: LISSAMPHIBIA, HAECKEL, 1866?
SUPERORDER: SALIENTIA LAURENTI, 1768
ORDER: ANURA RAFINESQUE, 1815
SUBORDER: NEOBATRACHIA REIG, 1958
Superfamily: HYLOIDEA Rafinesque, 1815 (sensu Wied, 1856)
Family: LEPTODACTYLIDAE Werner, 1896
Subfamily: CERATOPHYRINAE Tschudi, 1838
Beelzebufo ampinga
_____________________________________________________________________________________
MADAGASCAR
MAJUNGA PROVINCE
MAHAJANGA BASIN
MASIAKAKOHO STUDY AREA
MAD 03-17
MAEVARANO FORMATION
ANEMBALEMBA MEMBER
MAASTRICHTIAN STAGE
SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: CRUROTARSI SERENO & ARCUCCI, 1990
COHORT: CROCODYLOTARSI BENTON & CLARK, 1988
MAGNORDER: SUCHIA KREBS, 1974
SUPERORDER: CROCODYLOMORPHA WALKER, 1968
GRANDORDER: CROCODILIFORMES HAY, 1930
MICRORDER: MESOEUROCROCODYLIA WHETSTONE & WHYBROW, 1983
ORDER: MESOSUCHIA EMEND HUXLEY, 1875
PARVORDER: METASUCHIA BENTON & CLARK, 1988
GONDWANASUCHIA DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004
NOTOSUCHIMORPHA DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004
NOTOSUCHIFORMES DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004
TERRIASUCHIA DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004
INFRAORDER: NOTOSUCHIA GASPARINI, 1971
Family: NOTOSUCHIDAE Dollo, 1924
Simosuchus clarki
_____________________________________________________________________________________
MADAGASCAR
MAJUNGA PROVINCE
MAHAJANGA BASIN
MAD 03-18
MAEVARANO FORMATION
ANEMBALEMBA MEMBER
MAASTRICHTIAN STAGE
SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: AMPHIBIA LINNAEUS, 1758
SUBCLASS: LISSAMPHIBIA, HAECKEL, 1866?
SUPERORDER: SALIENTIA LAURENTI, 1768
ORDER: ANURA RAFINESQUE, 1815
SUBORDER: NEOBATRACHIA REIG, 1958
Superfamily: HYLOIDEA Rafinesque, 1815 (sensu Wied, 1856)
Family: LEPTODACTYLIDAE Werner, 1896
Subfamily: CERATOPHYRINAE Tschudi, 1838
Beelzebufo ampinga
_____________________________________________________________________________________
MADAGASCAR
MAJUNGA PROVINCE
MAHAJANGA BASIN
MAD 03-18
MAEVARANO FORMATION
ANEMBALEMBA MEMBER
MAASTRICHTIAN STAGE
SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: AMPHIBIA LINNAEUS, 1758
SUBCLASS: LISSAMPHIBIA, HAECKEL, 1866?
SUPERORDER: SALIENTIA LAURENTI, 1768
ORDER: ANURA RAFINESQUE, 1815
SUBORDER: NEOBATRACHIA REIG, 1958
Superfamily: HYLOIDEA Rafinesque, 1815 (sensu Wied, 1856)
Family: LEPTODACTYLIDAE Werner, 1896
Subfamily: CERATOPHYRINAE Tschudi, 1838
Beelzebufo ampinga
_____________________________________________________________________________________
MADAGASCAR
MAJUNGA PROVINCE
MAHAJANGA BASIN
MAD 03-40
MAEVARANO FORMATION
ANEMBALEMBA MEMBER
MAASTRICHTIAN STAGE
SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
AVEPODA PAUL, 2002
SUPERORDER: NEOTHEROPODA BAKKER, 1986
GRANDORDER: CERATOSAURIA NOVAS, 1992
AVEROSTRA PAUL, 2001
ORDER: CERATOSAURIA MARSH, 1884
Superfamily: ABELISAUROIDEA Novas, 1989
Family: NOASAURIDAE Bonaparte & Powell, 1980
Subfamily: VELOCISAURIINAE Bonaparte, 1991 (emend AGNOLIN, NOVAS & APESTEGUIA, 2003)
Masiakasaurus knopfleri
_____________________________________________________________________________________
MADAGASCAR
MAJUNGA PROVINCE
MAHAJANGA BASIN
MAD 03-66
MAEVARANO FORMATION
ANEMBALEMBA MEMBER
MAASTRICHTIAN STAGE
SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
AVEPODA PAUL, 2002
SUPERORDER: NEOTHEROPODA BAKKER, 1986
GRANDORDER: CERATOSAURIA NOVAS, 1992
AVEROSTRA PAUL, 2001
ORDER: CERATOSAURIA MARSH, 1884
Superfamily: ABELISAUROIDEA Novas, 1989
Family: NOASAURIDAE Bonaparte & Powell, 1980
Subfamily: VELOCISAURIINAE Bonaparte, 1991 (emend AGNOLIN, NOVAS & APESTEGUIA, 2003)
Masiakasaurus knopfleri
_____________________________________________________________________________________
MADAGASCAR
MAJUNGA PROVINCE
MAHAJANGA BASIN
MAD 05-04
MAEVARANO FORMATION
ANEMBALEMBA MEMBER
MAASTRICHTIAN STAGE
SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: CHELONOMORPHA KUHN, 1960
SUPERCOHORT: TESTUDINES LINNAEUS, 1758
COHORT: CASICHELYDIA emend GAFFNEY & MEYLAN, 1988
MAGNORDER: PLEURODIRA GAFFNEY & MEYLAN, 1988
ORDER: PLEURODIRIDA COPE, 1864
SUBORDER: EUPLEURODIRA GAFFNEY & MEYLAN, 1988
Superfamily: PELOMEDUSOIDES Cope, 1868 Cope, 1868
Superfamily: BOTHREMYDINURA Baur, 1891
Epifamily: BOTHERMYDINURA Baur, 1891
Family: BOTHREMYDIDAE Baur, 1891
Subfamily: BOTHREMYDINAE Baur, 1891
Infrafamily: BOTHREMYDODDA Baur, 1891
Tribe: KURMADEMYDINI Gaffney, Tong & Meylan, 2006
Kinkonychelys rogersi
SUBCLASS: DIAPSIDA OSBORN, 1903
INFRACLASS: NEODIAPSIDA BENTON, 1985
LEGION: LEPIDOSAUROMORPHA BENTON, 1983
SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)
MAGNORDER: SQUAMATA, OPPEL, 1811
SUPERORDER: SERPENTES LINNAEUS, 1759
ORDER: OPHIDIA BRONGNIART, 1800
SUBORDER: ALETHINOPHIDIA NOPCSCA, 1923
Superfamily: BOOIDEA Gray, 1825
Family: BOIDAE Gray, 1825
Subfamily: BOOIDINAE Gray, 1825
Infrafamily: BOOINIEA Gray, 1925
Tribe: MATSOINI Hoffstetter, 1961
Menarana nosymena
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
AVEPODA PAUL, 2002
SUPERORDER: NEOTHEROPODA BAKKER, 1986
GRANDORDER: CERATOSAURIA NOVAS, 1992
AVEROSTRA PAUL, 2001
ORDER: CERATOSAURIA MARSH, 1884
Superfamily: ABELISAUROIDEA Novas, 1989
Family: NOASAURIDAE Bonaparte & Powell, 1980
Subfamily: VELOCISAURIINAE Bonaparte, 1991 (emend AGNOLIN, NOVAS & APESTEGUIA, 2003)
Masiakasaurus knopfleri
_____________________________________________________________________________________
MADAGASCAR
MAJUNGA PROVINCE
MAHAJANGA BASIN
MAD 05-36
MAEVARANO FORMATION
ANEMBALEMBA MEMBER
MAASTRICHTIAN STAGE
SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: CRUROTARSI SERENO & ARCUCCI, 1990
COHORT: CROCODYLOTARSI BENTON & CLARK, 1988
MAGNORDER: SUCHIA KREBS, 1974
SUPERORDER: CROCODYLOMORPHA WALKER, 1968
GRANDORDER: CROCODILIFORMES HAY, 1930
MICRORDER: MESOEUROCROCODYLIA WHETSTONE & WHYBROW, 1983
ORDER: MESOSUCHIA EMEND HUXLEY, 1875
PARVORDER: METASUCHIA BENTON & CLARK, 1988
GONDWANASUCHIA DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004
NOTOSUCHIMORPHA DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004
NOTOSUCHIFORMES DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004
TERRIASUCHIA DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004
INFRAORDER: NOTOSUCHIA GASPARINI, 1971
Family: NOTOSUCHIDAE Dollo, 1924
Simosuchus clarki
_____________________________________________________________________________________
MADAGASCAR
MAJUNGA PROVINCE
MAHAJANGA BASIN
MAD 05-37
MAEVARANO FORMATION
ANEMBALEMBA MEMBER
MAASTRICHTIAN STAGE
SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: CRUROTARSI SERENO & ARCUCCI, 1990
COHORT: CROCODYLOTARSI BENTON & CLARK, 1988
MAGNORDER: SUCHIA KREBS, 1974
SUPERORDER: CROCODYLOMORPHA WALKER, 1968
GRANDORDER: CROCODILIFORMES HAY, 1930
MICRORDER: MESOEUROCROCODYLIA WHETSTONE & WHYBROW, 1983
ORDER: MESOSUCHIA EMEND HUXLEY, 1875
PARVORDER: METASUCHIA BENTON & CLARK, 1988
GONDWANASUCHIA DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004
NOTOSUCHIMORPHA DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004
NOTOSUCHIFORMES DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004
TERRIASUCHIA DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004
INFRAORDER: NOTOSUCHIA GASPARINI, 1971
Family: NOTOSUCHIDAE Dollo, 1924
Simosuchus clarki
_____________________________________________________________________________________
MADAGASCAR
MAJUNGA PROVINCE
MAHAJANGA BASIN
BOENY REGION
MAD 05-38
MAEVARANO FORMATION
ANEMBALEMBA MEMBER
MAASTRICHTIAN STAGE
SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: CHELONOMORPHA KUHN, 1960
SUPERCOHORT: TESTUDINES LINNAEUS, 1758
COHORT: CASICHELYDIA emend GAFFNEY & MEYLAN, 1988
MAGNORDER: PLEURODIRA GAFFNEY & MEYLAN, 1988
ORDER: PLEURODIRIDA COPE, 1864
SUBORDER: EUPLEURODIRA GAFFNEY & MEYLAN, 1988
Superfamily: PELOMEDUSOIDES Cope, 1868 Cope, 1868
Hyperfamily: PODOCNEMIDERA Cope, 1868
CLADE: PAN-PODOCNEMIDIDAE
Family: SAHONACHELYIDAE Joyce, Rollot, Evers, Lyson, Rahantariosa & Krause, 2021
*Sahonachelys mailakavava
_____________________________________________________________________________________
MADAGASCAR
MAJUNGA PROVINCE
MAHAJANGA BASIN
MAD 05-42
MAEVARANO FORMATION
ANEMBALEMBA MEMBER
MAASTRICHTIAN STAGE
SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: CRUROTARSI SERENO & ARCUCCI, 1990
COHORT: CROCODYLOTARSI BENTON & CLARK, 1988
MAGNORDER: SUCHIA KREBS, 1974
SUPERORDER: CROCODYLOMORPHA WALKER, 1968
GRANDORDER: CROCODILIFORMES HAY, 1930
MICRORDER: MESOEUROCROCODYLIA WHETSTONE & WHYBROW, 1983
ORDER: MESOSUCHIA EMEND HUXLEY, 1875
PARVORDER: METASUCHIA BENTON & CLARK, 1988
GONDWANASUCHIA DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004
NOTOSUCHIMORPHA DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004
NOTOSUCHIFORMES DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004
TERRIASUCHIA DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004
INFRAORDER: NOTOSUCHIA GASPARINI, 1971
Family: NOTOSUCHIDAE Dollo, 1924
Simosuchus clarki
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
AVEPODA PAUL, 2002
SUPERORDER: NEOTHEROPODA BAKKER, 1986
GRANDORDER: CERATOSAURIA NOVAS, 1992
AVEROSTRA PAUL, 2001
ORDER: CERATOSAURIA MARSH, 1884
Superfamily: ABELISAUROIDEA Novas, 1989
Family: ABELISAURIDAE Bonaparte & Novas, 1985
Subfamily: CARNOTAURINAE Sereno, 1998
Majungatholus atopus
Family: NOASAURIDAE Bonaparte & Powell, 1980
Subfamily: VELOCISAURIINAE Bonaparte, 1991 (emend AGNOLIN, NOVAS & APESTEGUIA, 2003)
Masiakasaurus knopfleri
CLASS: AVES LINNAEUS, 1758
Incertae sedis
Gen.sp indet.
SUBCLASS: SAURORNITHES NICHOLSON, 1879
AVEBREVICAUDA PAUL, 2001
INFRACLASS: ORNITHOTHORACES NOVA
Incertae sedis
Gen.sp indet.
LEGION: ORNITHURAE NOVA
COHORT: ENANTIORNITHES NOVA
Incertae sedis
*Falcatakely forsterae
Gen.sp indet.
SUPERORDER: EUENANTIORNITHES CHIAPPE, 2002
ORDER: EUORNITHIFORMES KUROCHKIN, 1996
Family: CONCORNITHIDAE Kurochkin, 1996
Vorona berivotrensis
_____________________________________________________________________________________
MADAGASCAR
MAJUNGA PROVINCE
MAHAJANGA BASIN
MASIAKAKOHO STUDY AREA
MAD 05-59
MAEVARANO FORMATION
ANEMBALEMBA MEMBER
MAASTRICHTIAN STAGE
SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: CRUROTARSI SERENO & ARCUCCI, 1990
COHORT: CROCODYLOTARSI BENTON & CLARK, 1988
MAGNORDER: SUCHIA KREBS, 1974
SUPERORDER: CROCODYLOMORPHA WALKER, 1968
GRANDORDER: CROCODILIFORMES HAY, 1930
MICRORDER: MESOEUROCROCODYLIA WHETSTONE & WHYBROW, 1983
ORDER: MESOSUCHIA EMEND HUXLEY, 1875
PARVORDER: METASUCHIA BENTON & CLARK, 1988
GONDWANASUCHIA DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004
NOTOSUCHIMORPHA DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004
NOTOSUCHIFORMES DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004
TERRIASUCHIA DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004
INFRAORDER: NOTOSUCHIA GASPARINI, 1971
Family: NOTOSUCHIDAE Dollo, 1924
Simosuchus clarki
_____________________________________________________________________________________
MADAGASCAR
MAJUNGA PROVINCE
MAHAJANGA BASIN
MASIAKAKOHO STUDY AREA
MAD 05-61
MAEVARANO FORMATION
MAASTRICHTIAN STAGE
SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: CHELONOMORPHA KUHN, 1960
SUPERCOHORT: TESTUDINES LINNAEUS, 1758
COHORT: CASICHELYDIA emend GAFFNEY & MEYLAN, 1988
MAGNORDER: PLEURODIRA GAFFNEY & MEYLAN, 1988
ORDER: PLEURODIRIDA COPE, 1864
SUBORDER: EUPLEURODIRA GAFFNEY & MEYLAN, 1988
Superfamily: PELOMEDUSOIDES Cope, 1868 Cope, 1868
Hyperfamily: PODOCNEMIDERA Cope, 1868
CLADE: PAN-PODOCNEMIDIDAE
Family: SAHONACHELYIDAE Joyce, Rollot, Evers, Lyson, Rahantariosa & Krause, 2021
Sokatra antitra
_____________________________________________________________________________________
MADAGASCAR
MAJUNGA PROVINCE
MAHAJANGA BASIN
MAD 05-64
MAEVARANO FORMATION
ANEMBALEMBA MEMBER
MAASTRICHTIAN STAGE
SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: AMPHIBIA LINNAEUS, 1758
SUBCLASS: LISSAMPHIBIA, HAECKEL, 1866?
SUPERORDER: SALIENTIA LAURENTI, 1768
ORDER: ANURA RAFINESQUE, 1815
SUBORDER: NEOBATRACHIA REIG, 1958
Superfamily: HYLOIDEA Rafinesque, 1815 (sensu Wied, 1856)
Family: LEPTODACTYLIDAE Werner, 1896
Subfamily: CERATOPHYRINAE Tschudi, 1838
Beelzebufo ampinga
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
INFRACLASS: NEODIAPSIDA BENTON, 1985
LEGION: LEPIDOSAUROMORPHA BENTON, 1983
SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)
MAGNORDER: SQUAMATA, OPPEL, 1811
SUPERORDER: SERPENTES LINNAEUS, 1759
ORDER: OPHIDIA BRONGNIART, 1800
SUBORDER: ALETHINOPHIDIA NOPCSCA, 1923
Superfamily: BOOIDEA Gray, 1825
Subfamily: BOOIDINAE Gray, 1825
Infrafamily: BOOINIEA Gray, 1925
Tribe: MATSOINI Hoffstetter, 1961
Madtsoia madagascariensis
_____________________________________________________________________________________
MADAGASCAR
MAJUNGA PROVINCE
MAHAJANGA BASIN
MAD 07-15
MAEVARANO FORMATION
ANEMBALEMBA MEMBER
MAASTRICHTIAN STAGE
SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: AMPHIBIA LINNAEUS, 1758
SUBCLASS: LISSAMPHIBIA, HAECKEL, 1866?
SUPERORDER: SALIENTIA LAURENTI, 1768
ORDER: ANURA RAFINESQUE, 1815
SUBORDER: NEOBATRACHIA REIG, 1958
Superfamily: HYLOIDEA Rafinesque, 1815 (sensu Wied, 1856)
Family: LEPTODACTYLIDAE Werner, 1896
Subfamily: CERATOPHYRINAE Tschudi, 1838
Beelzebufo ampinga
_____________________________________________________________________________________
MADAGASCAR
MAJUNGA PROVINCE
MAHAJANGA BASIN
MAD 07-16
MAEVARANO FORMATION
ANEMBALEMBA MEMBER
MAASTRICHTIAN STAGE
SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
AVEPODA PAUL, 2002
SUPERORDER: NEOTHEROPODA BAKKER, 1986
GRANDORDER: CERATOSAURIA NOVAS, 1992
AVEROSTRA PAUL, 2001
ORDER: CERATOSAURIA MARSH, 1884
Superfamily: ABELISAUROIDEA Novas, 1989
Family: NOASAURIDAE Bonaparte & Powell, 1980
Subfamily: VELOCISAURIINAE Bonaparte, 1991 (emend AGNOLIN, NOVAS & APESTEGUIA, 2003)
Masiakasaurus knopfleri
_____________________________________________________________________________________
MADAGASCAR
MAJUNGA PROVINCE
MAHAJANGA BASIN
MAD 07-17
MAEVARANO FORMATION
ANEMBALEMBA MEMBER
MAASTRICHTIAN STAGE
SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
AVEPODA PAUL, 2002
SUPERORDER: NEOTHEROPODA BAKKER, 1986
GRANDORDER: CERATOSAURIA NOVAS, 1992
AVEROSTRA PAUL, 2001
ORDER: CERATOSAURIA MARSH, 1884
Superfamily: ABELISAUROIDEA Novas, 1989
Family: NOASAURIDAE Bonaparte & Powell, 1980
Subfamily: VELOCISAURIINAE Bonaparte, 1991 (emend AGNOLIN, NOVAS & APESTEGUIA, 2003)
Masiakasaurus knopfleri
_____________________________________________________________________________________
MADAGASCAR
MAJUNGA PROVINCE
MAHAJANGA BASIN
MAD 07-20
MAEVARANO FORMATION
ANEMBALEMBA MEMBER
MAASTRICHTIAN STAGE
SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: AMPHIBIA LINNAEUS, 1758
SUBCLASS: LISSAMPHIBIA, HAECKEL, 1866?
SUPERORDER: SALIENTIA LAURENTI, 1768
ORDER: ANURA RAFINESQUE, 1815
SUBORDER: NEOBATRACHIA REIG, 1958
Superfamily: HYLOIDEA Rafinesque, 1815 (sensu Wied, 1856)
Family: LEPTODACTYLIDAE Werner, 1896
Subfamily: CERATOPHYRINAE Tschudi, 1838
Beelzebufo ampinga
_____________________________________________________________________________________
NORTHEASTERN MADAGASCAR
MAHAJANA PROVINCE
MAHAJANGA BASIN
APPROXIMATELY 2 KM NORTH OF THE WESTERN END OF LAC KINKONY AND 4.9 KM NORTHEAST OF THE VILLAGE OF ANTONGOMENA
LAC KINKONY STUDY AREA
MAD 07-25
UNNAMED MEMBER
MAEVARANO FORMATION
MAASTRICHTIAN STAGE
SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: CHELONOMORPHA KUHN, 1960
SUPERCOHORT: TESTUDINES LINNAEUS, 1758
COHORT: CASICHELYDIA emend GAFFNEY & MEYLAN, 1988
MAGNORDER: PLEURODIRA GAFFNEY & MEYLAN, 1988
ORDER: PLEURODIRIDA COPE, 1864
SUBORDER: EUPLEURODIRA GAFFNEY & MEYLAN, 1988
Superfamily: PELOMEDUSOIDES Cope, 1868 Cope, 1868
Superfamily: BOTHREMYDINURA Baur, 1891
Epifamily: BOTHERMYDINURA Baur, 1891
Family: BOTHREMYDIDAE Baur, 1891
Subfamily: BOTHREMYDINAE Baur, 1891
Infrafamily: BOTHREMYDODDA Baur, 1891
Tribe: KURMADEMYDINI Gaffney, Tong & Meylan, 2006
*Kinkonychelys rogersi
_____________________________________________________________________________________
MADAGASCAR
MAJUNGA PROVINCE
MAHAJANGA BASIN
BERIVOTRA STUDY AREA
MAD 07-66
MAEVARANO FORMATION
MAASTRICHTIAN STAGE
SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: CHELONOMORPHA KUHN, 1960
SUPERCOHORT: TESTUDINES LINNAEUS, 1758
COHORT: CASICHELYDIA emend GAFFNEY & MEYLAN, 1988
MAGNORDER: PLEURODIRA GAFFNEY & MEYLAN, 1988
ORDER: PLEURODIRIDA COPE, 1864
SUBORDER: EUPLEURODIRA GAFFNEY & MEYLAN, 1988
Superfamily: PELOMEDUSOIDES Cope, 1868 Cope, 1868
Hyperfamily: PODOCNEMIDERA Cope, 1868
CLADE: PAN-PODOCNEMIDIDAE
Family: SAHONACHELYIDAE Joyce, Rollot, Evers, Lyson, Rahantariosa & Krause, 2021
Sokatra antitra
_____________________________________________________________________________________
MADAGASCAR
MAJUNGA PROVINCE
MAHAJANGA BASIN
MAD 10-13
MAEVARANO FORMATION
ANEMBALEMBA MEMBER
MAASTRICHTIAN STAGE
SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: AMPHIBIA LINNAEUS, 1758
SUBCLASS: LISSAMPHIBIA, HAECKEL, 1866?
SUPERORDER: SALIENTIA LAURENTI, 1768
ORDER: ANURA RAFINESQUE, 1815
SUBORDER: NEOBATRACHIA REIG, 1958
Superfamily: HYLOIDEA Rafinesque, 1815 (sensu Wied, 1856)
Family: LEPTODACTYLIDAE Werner, 1896
Subfamily: CERATOPHYRINAE Tschudi, 1838
Beelzebufo ampinga
_____________________________________________________________________________________
MADAGASCAR
MAJUNGA PROVINCE
MAHAJANGA BASIN
MAD 10-24
MAEVARANO FORMATION
LAC KINKOY MEMBER
MAASTRICHTIAN STAGE
SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: AMPHIBIA LINNAEUS, 1758
SUBCLASS: LISSAMPHIBIA, HAECKEL, 1866?
SUPERORDER: SALIENTIA LAURENTI, 1768
ORDER: ANURA RAFINESQUE, 1815
SUBORDER: NEOBATRACHIA REIG, 1958
Superfamily: HYLOIDEA Rafinesque, 1815 (sensu Wied, 1856)
Family: LEPTODACTYLIDAE Werner, 1896
Subfamily: CERATOPHYRINAE Tschudi, 1838
Beelzebufo ampinga
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
INFRACLASS: NEODIAPSIDA BENTON, 1985
LEGION: LEPIDOSAUROMORPHA BENTON, 1983
SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)
MAGNORDER: SQUAMATA, OPPEL, 1811
SUPERORDER: SERPENTES LINNAEUS, 1759
Incertae sedis
Gen. sp indet
ORDER: OPHIDIA BRONGNIART, 1800
SUBORDER: ALETHINOPHIDIA NOPCSCA, 1923
Superfamily: BOOIDEA Gray, 1825
Family: ACROCHORDOIDAE Bonaparte, 1838
Subfamily: ?NIGEROPHEINAE Rage, 1975
*Indophis fanambinana
CLASS: MAMMALIA LINNAEUS, 1758
SUBCLASS: THERIIFORMES (ROWE, 1988) McKENNA & BELL, 1997
INFRACLASS: ALLOTHERIA (MARSH, 1880) McKENNA & BELL, 1997
ORDER: MULTITUBERCULATA COPE, 1884
SUBORDER: GONDWANATHERIA Mones, 1987
Family: SUDAMERICICIDAE Scillato-Yane & Prasad, 1984
*Vintana sertichi
_____________________________________________________________________________________
MADAGASCAR
MAJUNGA PROVINCE
MAHAJANGA BASIN
MAD 93-01
MAEVARANO FORMATION
ANEMBALEMBA MEMBER
MAASTRICHTIAN STAGE
SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: AMPHIBIA LINNAEUS, 1758
SUBCLASS: LISSAMPHIBIA, HAECKEL, 1866?
SUPERORDER: SALIENTIA LAURENTI, 1768
ORDER: ANURA RAFINESQUE, 1815
SUBORDER: NEOBATRACHIA REIG, 1958
Superfamily: HYLOIDEA Rafinesque, 1815 (sensu Wied, 1856)
Family: LEPTODACTYLIDAE Werner, 1896
Subfamily: CERATOPHYRINAE Tschudi, 1838
Beelzebufo ampinga
SUBORDER: MESOBATRACHIA LAURENT, 1979
Superfamily: PELOBATOIDEA Stanius, 1856 , nom. corr. Spinar, 1971 (sensu Bolkay, 1919)
Family: PELOBATIDAE Stanius, 1856 , nom. corr. Spinar, 1971
Gen. sp indet.
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
INFRACLASS: NEODIAPSIDA BENTON, 1985
LEGION: LEPIDOSAUROMORPHA BENTON, 1983
SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)
MAGNORDER: SQUAMATA, OPPEL, 1811
SUPERORDER: SERPENTES LINNAEUS, 1759
Incertae sedis
Gen. sp indet.
ORDER: OPHIDIA BRONGNIART, 1800
SUBORDER: ALETHINOPHIDIA NOPCSCA, 1923
Superfamily: BOOIDEA Gray, 1825
Family: BOIDAE Gray, 1825
Subfamily: BOOIDINAE Gray, 1825
Infrafamily: BOOINIEA Gray, 1925
Tribe: MATSOINI Hoffstetter, 1961
Madtsoia madagascariensis
Family; ACROCHORDOIDAE Bonaparte, 1838
Subfamily: ?NIGEROPHEINAE Rage, 1975
Kelyophis hechti
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
AVEPODA PAUL, 2002
SUPERORDER: NEOTHEROPODA BAKKER, 1986
GRANDORDER: CERATOSAURIA NOVAS, 1992
AVEROSTRA PAUL, 2001
ORDER: CERATOSAURIA MARSH, 1884
Superfamily: ABELISAUROIDEA Novas, 1989
Family: NOASAURIDAE Bonaparte & Powell, 1980
Subfamily: VELOCISAURIINAE Bonaparte, 1991 (emend AGNOLIN, NOVAS & APESTEGUIA, 2003)
Masiakasaurus knopfleri
Family: ABELISAURIDAE Bonaparte & Novas, 1985
Subfamily: CARNOTAURINAE Sereno, 1998
Majungatholus atopus
_____________________________________________________________________________________
MADAGASCAR
MAJUNGA PROVINCE
MAHAJANGA BASIN
MAD 93-06
MAEVARANO FORMATION
ANEMBALEMBA MEMBER
MAASTRICHTIAN STAGE
SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: AMPHIBIA LINNAEUS, 1758
SUBCLASS: LISSAMPHIBIA, HAECKEL, 1866?
SUPERORDER: SALIENTIA LAURENTI, 1768
ORDER: ANURA RAFINESQUE, 1815
SUBORDER: NEOBATRACHIA REIG, 1958
Superfamily: HYLOIDEA Rafinesque, 1815 (sensu Wied, 1856)
Family: LEPTODACTYLIDAE Werner, 1896
Subfamily: CERATOPHYRINAE Tschudi, 1838
Beelzebufo ampinga
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: CRUROTARSI SERENO & ARCUCCI, 1990
COHORT: CROCODYLOTARSI BENTON & CLARK, 1988
MAGNORDER: SUCHIA KREBS, 1974
SUPERORDER: CROCODYLOMORPHA WALKER, 1968
GRANDORDER: CROCODILIFORMES HAY, 1930
MICRORDER: MESOEUROCROCODYLIA WHETSTONE & WHYBROW, 1983
ORDER: MESOSUCHIA EMEND HUXLEY, 1875
PARVORDER: METASUCHIA BENTON & CLARK, 1988
GONDWANASUCHIA DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004
NOTOSUCHIMORPHA DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004
NOTOSUCHIFORMES DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004
TERRIASUCHIA DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004
INFRAORDER: NOTOSUCHIA GASPARINI, 1971
Family: NOTOSUCHIDAE Dollo, 1924
Simosuchus clarki
_____________________________________________________________________________________
MADAGASCAR
MAJUNGA PROVINCE
MAHAJANGA BASIN
MAD 93-09
MAEVARANO FORMATION
ANEMBALEMBA MEMBER
MAASTRICHTIAN STAGE
SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
INFRACLASS: NEODIAPSIDA BENTON, 1985
LEGION: LEPIDOSAUROMORPHA BENTON, 1983
SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)
MAGNORDER: SQUAMATA, OPPEL, 1811
SUPERORDER: SERPENTES LINNAEUS, 1759
Incertae sedis
Gen. sp indet.
ORDER: OPHIDIA BRONGNIART, 1800
SUBORDER: ALETHINOPHIDIA NOPCSCA, 1923
Superfamily: BOOIDEA Gray, 1825
Family: BOIDAE Gray, 1825
Subfamily: BOOIDINAE Gray, 1825
Infrafamily: BOOINIEA Gray, 1925
Tribe: MATSOINI Hoffstetter, 1961
Madtsoia madagascariensis
Family; ACROCHORDOIDAE Bonaparte, 1838
Subfamily: ?NIGEROPHEINAE Rage, 1975
Kelyophis hechti
LEGION: ARCHOSAUROMORPHA HUENE, 1946SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
AVEPODA PAUL, 2002
SUPERORDER: NEOTHEROPODA BAKKER, 1986
GRANDORDER: CERATOSAURIA NOVAS, 1992
AVEROSTRA PAUL, 2001
ORDER: CERATOSAURIA MARSH, 1884
Superfamily: ABELISAUROIDEA Novas, 1989
Family: NOASAURIDAE Bonaparte & Powell, 1980
Subfamily: VELOCISAURIINAE Bonaparte, 1991 (emend AGNOLIN, NOVAS & APESTEGUIA, 2003)
Masiakasaurus knopfleri
_____________________________________________________________________________________
MADAGASCAR
MAJUNGA PROVINCE
MAHAJANGA BASIN
MAD 93-09
MAEVARANO FORMATION
ANEMBALEMBA MEMBER
MAASTRICHTIAN STAGE
SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
INFRACLASS: NEODIAPSIDA BENTON, 1985
LEGION: LEPIDOSAUROMORPHA BENTON, 1983
SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)
MAGNORDER: SQUAMATA, OPPEL, 1811
SUPERORDER: SERPENTES LINNAEUS, 1759
ORDER: OPHIDIA BRONGNIART, 1800
SUBORDER: ALETHINOPHIDIA NOPCSCA, 1923
Superfamily: BOOIDEA Gray, 1825
Family: BOIDAE Gray, 1825
Subfamily: BOOIDINAE Gray, 1825
Infrafamily: BOOINIEA Gray, 1925
Tribe: MATSOINI Hoffstetter, 1961
Madtsoia madagascariensis
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
AVEPODA PAUL, 2002
SUPERORDER: NEOTHEROPODA BAKKER, 1986
GRANDORDER: CERATOSAURIA NOVAS, 1992
AVEROSTRA PAUL, 2001
ORDER: CERATOSAURIA MARSH, 1884
Superfamily: ABELISAUROIDEA Novas, 1989
Family: NOASAURIDAE Bonaparte & Powell, 1980
Subfamily: VELOCISAURIINAE Bonaparte, 1991 (emend AGNOLIN, NOVAS & APESTEGUIA, 2003)
Masiakasaurus knopfleri
_____________________________________________________________________________________
MADAGASCAR
MAJUNGA PROVINCE
MAHAJANGA BASIN
MAD 93-14
MAEVARANO FORMATION
ANEMBALEMBA MEMBER
MAASTRICHTIAN STAGE
SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: AMPHIBIA LINNAEUS, 1758
SUBCLASS: LISSAMPHIBIA, HAECKEL, 1866?
SUPERORDER: SALIENTIA LAURENTI, 1768
ORDER: ANURA RAFINESQUE, 1815
SUBORDER: NEOBATRACHIA REIG, 1958
Superfamily: HYLOIDEA Rafinesque, 1815 (sensu Wied, 1856)
Family: LEPTODACTYLIDAE Werner, 1896
Subfamily: CERATOPHYRINAE Tschudi, 1838
Beelzebufo ampinga
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
INFRACLASS: NEODIAPSIDA BENTON, 1985
LEGION: LEPIDOSAUROMORPHA BENTON, 1983
SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)
MAGNORDER: SQUAMATA, OPPEL, 1811
SUPERORDER: SERPENTES LINNAEUS, 1759
ORDER: OPHIDIA BRONGNIART, 1800
SUBORDER: ALETHINOPHIDIA NOPCSCA, 1923
Superfamily: BOOIDEA Gray, 1825
Family: BOIDAE Gray, 1825
Subfamily: BOOIDINAE Gray, 1825
Infrafamily: BOOINIEA Gray, 1925
Tribe: MATSOINI Hoffstetter, 1961
Madtsoia madagascariensis
*Menarana nosymena
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: CRUROTARSI SERENO & ARCUCCI, 1990
COHORT: CROCODYLOTARSI BENTON & CLARK, 1988
MAGNORDER: SUCHIA KREBS, 1974
SUPERORDER: CROCODYLOMORPHA WALKER, 1968
GRANDORDER: CROCODILIFORMES HAY, 1930
MICRORDER: MESOEUROCROCODYLIA WHETSTONE & WHYBROW, 1983
ORDER: MESOSUCHIA EMEND HUXLEY, 1875
PARVORDER: METASUCHIA BENTON & CLARK, 1988
GONDWANASUCHIA DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004
NOTOSUCHIMORPHA DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004
NOTOSUCHIFORMES DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004
TERRIASUCHIA DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004
INFRAORDER: NOTOSUCHIA GASPARINI, 1971
Family: NOTOSUCHIDAE Dollo, 1924
Simosuchus clarki
_____________________________________________________________________________________
MADAGASCARMAHAJANGA BASIN
SOUTH OF BERIVORTA
MAD 93-16
MAEVARANO FORMATION
MAASTRICHTIAN STAGE
UPPER SENONIAN SUBEPOCH
UPPER UPPER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
INFRACLASS: NEODIAPSIDA BENTON, 1985
LEGION: LEPIDOSAUROMORPHA BENTON, 1983
SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)
MAGNORDER: SQUAMATA, OPPEL, 1811
SUPERORDER: SERPENTES LINNAEUS, 1759
ORDER: OPHIDIA BRONGNIART, 1800
SUBORDER: ALETHINOPHIDIA NOPCSCA, 1923
Superfamily: BOOIDEA Gray, 1825
Family: BOIDAE Gray, 1825
Subfamily: BOOIDINAE Gray, 1825
Infrafamily: BOOINIEA Gray, 1925
Tribe: MATSOINI Hoffstetter, 1961
Menarana nosymena
_____________________________________________________________________________________
MADAGASCAR
MAHAJANGA BASIN
SOUTH OF BERIVORTA
MAD 93-17MAEVARANO FORMATION
MAASTRICHTIAN STAGE
UPPER SENONIAN SUBEPOCH
UPPER UPPER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: AMPHIBIA LINNAEUS, 1758
SUBCLASS: LISSAMPHIBIA, HAECKEL, 1866?
SUPERORDER: SALIENTIA LAURENTI, 1768
ORDER: ANURA RAFINESQUE, 1815
SUBORDER: NEOBATRACHIA REIG, 1958
Superfamily: HYLOIDEA Rafinesque, 1815 (sensu Wied, 1856)
Family: LEPTODACTYLIDAE Werner, 1896
Subfamily: CERATOPHYRINAE Tschudi, 1838
Beelzebufo ampinga
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
INFRACLASS: NEODIAPSIDA BENTON, 1985
LEGION: LEPIDOSAUROMORPHA BENTON, 1983
SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)
MAGNORDER: SQUAMATA, OPPEL, 1811
SUPERORDER: SERPENTES LINNAEUS, 1759
ORDER: OPHIDIA BRONGNIART, 1800
SUBORDER: ALETHINOPHIDIA NOPCSCA, 1923
Superfamily: BOOIDEA Gray, 1825
Family: BOIDAE Gray, 1825
Subfamily: BOOIDINAE Gray, 1825
Infrafamily: BOOINIEA Gray, 1925
Tribe: MATSOINI Hoffstetter, 1961
Madtsoia madagascariensis
_____________________________________________________________________________________
MADAGASCAR
MAJUNGA PROVINCE
MAHAJANGA BASIN
MAD 93-17
MAEVARANO FORMATION
ANEMBALEMBA MEMBER
MAASTRICHTIAN STAGE
SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
INFRACLASS: NEODIAPSIDA BENTON, 1985
LEGION: LEPIDOSAUROMORPHA BENTON, 1983
SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)
MAGNORDER: SQUAMATA, OPPEL, 1811
SUPERORDER: SERPENTES LINNAEUS, 1759
ORDER: OPHIDIA BRONGNIART, 1800
SUBORDER: ALETHINOPHIDIA NOPCSCA, 1923
Superfamily: BOOIDEA Gray, 1825
Family: BOIDAE Gray, 1825
Subfamily: BOOIDINAE Gray, 1825
Infrafamily: BOOINIEA Gray, 1925
Tribe: MATSOINI Hoffstetter, 1961
Madtsoia madagascariensis
_____________________________________________________________________________________
MADAGASCAR
MAJUNGA PROVINCE
MAHAJANGA BASIN
MAD 93-18
MAEVARANO FORMATION
ANEMBALEMBA MEMBER
MAASTRICHTIAN STAGE
SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: CHELONOMORPHA KUHN, 1960
SUPERCOHORT: TESTUDINES LINNAEUS, 1758
COHORT: CASICHELYDIA emend GAFFNEY & MEYLAN, 1988
MAGNORDER: PLEURODIRA GAFFNEY & MEYLAN, 1988
ORDER: PLEURODIRIDA COPE, 1864
SUBORDER: EUPLEURODIRA GAFFNEY & MEYLAN, 1988
Superfamily: PELOMEDUSOIDES Cope, 1868 Cope, 1868
Superfamily: BOTHREMYDINURA Baur, 1891
Epifamily: BOTHERMYDINURA Baur, 1891
Family: BOTHREMYDIDAE Baur, 1891
Subfamily: BOTHREMYDINAE Baur, 1891
Infrafamily: BOTHREMYDODDA Baur, 1891
Tribe: KURMADEMYDINI Gaffney, Tong & Meylan, 2006
Kinkonychelys rogersi
SUBCLASS: DIAPSIDA OSBORN, 1903
INFRACLASS: NEODIAPSIDA BENTON, 1985
LEGION: LEPIDOSAUROMORPHA BENTON, 1983
SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)
MAGNORDER: SQUAMATA, OPPEL, 1811
SUPERORDER: SERPENTES LINNAEUS, 1759
ORDER: OPHIDIA BRONGNIART, 1800
SUBORDER: ALETHINOPHIDIA NOPCSCA, 1923
Superfamily: BOOIDEA Gray, 1825
Family: BOIDAE Gray, 1825
Subfamily: BOOIDINAE Gray, 1825
Infrafamily: BOOINIEA Gray, 1925
Tribe: MATSOINI Hoffstetter, 1961
Madtsoia madagascariensis
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: CRUROTARSI SERENO & ARCUCCI, 1990
COHORT: CROCODYLOTARSI BENTON & CLARK, 1988
MAGNORDER: SUCHIA KREBS, 1974
SUPERORDER: CROCODYLOMORPHA WALKER, 1968
GRANDORDER: CROCODILIFORMES HAY, 1930
MICRORDER: MESOEUROCROCODYLIA WHETSTONE & WHYBROW, 1983
ORDER: MESOSUCHIA EMEND HUXLEY, 1875
PARVORDER: METASUCHIA BENTON & CLARK, 1988
GONDWANASUCHIA DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004
NOTOSUCHIMORPHA DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004
NOTOSUCHIFORMES DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004
TERRIASUCHIA DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004
INFRAORDER: NOTOSUCHIA GASPARINI, 1971
Family: NOTOSUCHIDAE Dollo, 1924
Simosuchus clarki
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
AVEPODA PAUL, 2002
SUPERORDER: NEOTHEROPODA BAKKER, 1986
GRANDORDER: CERATOSAURIA NOVAS, 1992
AVEROSTRA PAUL, 2001
ORDER: CERATOSAURIA MARSH, 1884
Superfamily: ABELISAUROIDEA Novas, 1989
Family: ABELISAURIDAE Bonaparte & Novas, 1985
Subfamily: CARNOTAURINAE Sereno, 1998
Majungatholus atopus
GRANDORDER: TETANURAE GAUTHIER, 1986
MICRORDER: COELUROSAURIA HUENE, 1914
ORDER: MANIRAPTORFORMES HOLTZ, 1996
AVETHEROPOA PAUL, 2002
AVEPECTORA PAUL, 2002
SUBORDER: MANIRAPTORA GAUTHIER, 1986
INFRAORDER: PARVES SERENO, 1997
PARVORDER: DEINONYCHOSAURIA COLBERT & D. A. RUSSELL, 1969
Family: DROMAEOSAURIDAE Russell, 1969
Subfamily: UNENLAGIINAE Bonaparte, 1999
*Rahonavis ostromi
= Rahona ostromiCOHORT: ORNITHISCHIFORMES COOPER, 1985SUPERORDER: SAUROPODOMORPHA HUENE, 1932GRANDORDER: BRONTOSAURIA OLSHEVSKY, 1991 (emend)MICRORDER: SAUROPODA MARSH, 1878ORDER: EUSAUROPODASUBORDER: NEOSAUROPODA BONAPARTE, 1986INFRAORDER: MACRONARIA WILSON & SERENO, 1998PARVORDER: TITANOSAURIFORMES SALGADO ET AL, 1997Superfamily: TITANOSAUROIDEA Lydekker, 1885 (sensu Upchurch, 1999)Family: TITANOSAURIDAE Lydekker, 1885Incertae sedisVahiny depereti
CLASS: AVES LINNAEUS, 1758
Incertae sedis
Gen.sp indet.
SUBCLASS: SAURORNITHES NICHOLSON, 1879
AVEBREVICAUDA PAUL, 2001
INFRACLASS: ORNITHOTHORACES NOVA
Incertae sedis
Gen.sp indet.
LEGION: ORNITHURAE NOVA
Incertae sedis
Gen.sp indet.
COHORT: ENANTIORNITHES NOVA
Incertae sedis
Gen.sp indet.
SUPERORDER: EUENANTIORNITHES CHIAPPE, 2002
ORDER: EUORNITHIFORMES KUROCHKIN, 1996
Family: CONCORNITHIDAE Kurochkin, 1996
*Vorona berivotrensis
_____________________________________________________________________________________
MADAGASCAR
MAJUNGA PROVINCE
MAHAJANGA BASIN
MAD 93-19
MAEVARANO FORMATION
ANEMBALEMBA MEMBER
MAASTRICHTIAN STAGE
SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
INFRACLASS: NEODIAPSIDA BENTON, 1985
LEGION: LEPIDOSAUROMORPHA BENTON, 1983
SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)
MAGNORDER: SQUAMATA, OPPEL, 1811
SUPERORDER: SERPENTES LINNAEUS, 1759
Incertae sedis
Gen. sp indet.
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
AVEPODA PAUL, 2002
SUPERORDER: NEOTHEROPODA BAKKER, 1986
GRANDORDER: CERATOSAURIA NOVAS, 1992
AVEROSTRA PAUL, 2001
ORDER: CERATOSAURIA MARSH, 1884
Superfamily: ABELISAUROIDEA Novas, 1989
Family: ABELISAURIDAE Bonaparte & Novas, 1985
Subfamily: CARNOTAURINAE Sereno, 1998
Majungatholus atopus
_____________________________________________________________________________________
MADAGASCAR
MAJUNGA PROVINCE
MAHAJANGA BASIN
MAD 93-25
MAEVARANO FORMATION
ANEMBALEMBA MEMBER
MAASTRICHTIAN STAGE
SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: AMPHIBIA LINNAEUS, 1758
SUBCLASS: LISSAMPHIBIA, HAECKEL, 1866?
SUPERORDER: SALIENTIA LAURENTI, 1768
ORDER: ANURA RAFINESQUE, 1815
SUBORDER: NEOBATRACHIA REIG, 1958
Superfamily: HYLOIDEA Rafinesque, 1815 (sensu Wied, 1856)
Family: LEPTODACTYLIDAE Werner, 1896
Subfamily: CERATOPHYRINAE Tschudi, 1838
*Beelzebufo ampinga
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
INFRACLASS: NEODIAPSIDA BENTON, 1985
LEGION: LEPIDOSAUROMORPHA BENTON, 1983
SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)
MAGNORDER: SQUAMATA, OPPEL, 1811
SUPERORDER: SERPENTES LINNAEUS, 1759
ORDER: OPHIDIA BRONGNIART, 1800
SUBORDER: ALETHINOPHIDIA NOPCSCA, 1923
Superfamily: BOOIDEA Gray, 1825
Family: BOIDAE Gray, 1825
Subfamily: BOOIDINAE Gray, 1825
Infrafamily: BOOINIEA Gray, 1925
Tribe: MATSOINI Hoffstetter, 1961
Madtsoia madagascariensis
_____________________________________________________________________________________
MADAGASCAR
MAJUNGA PROVINCE
MAHAJANGA BASIN
MAD 93-25
MAEVARANO FORMATION
ANEMBALEMBA MEMBER
MAASTRICHTIAN STAGE
SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
INFRACLASS: NEODIAPSIDA BENTON, 1985
LEGION: LEPIDOSAUROMORPHA BENTON, 1983
SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)
MAGNORDER: SQUAMATA, OPPEL, 1811
SUPERORDER: SERPENTES LINNAEUS, 1759
ORDER: OPHIDIA BRONGNIART, 1800
SUBORDER: ALETHINOPHIDIA NOPCSCA, 1923
Superfamily: BOOIDEA Gray, 1825
Family: BOIDAE Gray, 1825
Subfamily: BOOIDINAE Gray, 1825
Infrafamily: BOOINIEA Gray, 1925
Tribe: MATSOINI Hoffstetter, 1961
Madtsoia madagascariensis
_____________________________________________________________________________________
MADAGASCAR
MAJUNGA PROVINCE
MAHAJANGA BASIN
MAD 93-26
MAEVARANO FORMATION
ANEMBALEMBA MEMBER
MAASTRICHTIAN STAGE
SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
AVEPODA PAUL, 2002
SUPERORDER: NEOTHEROPODA BAKKER, 1986
GRANDORDER: CERATOSAURIA NOVAS, 1992
AVEROSTRA PAUL, 2001
ORDER: CERATOSAURIA MARSH, 1884
Superfamily: ABELISAUROIDEA Novas, 1989
Family: NOASAURIDAE Bonaparte & Powell, 1980
Subfamily: VELOCISAURIINAE Bonaparte, 1991 (emend AGNOLIN, NOVAS & APESTEGUIA, 2003)
Masiakasaurus knopfleri
_____________________________________________________________________________________
MADAGASCAR
MAJUNGA PROVINCE
MAHAJANGA BASIN
MAD 93-27
MAEVARANO FORMATION
ANEMBALEMBA MEMBER
MAASTRICHTIAN STAGE
SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
INFRACLASS: NEODIAPSIDA BENTON, 1985
LEGION: LEPIDOSAUROMORPHA BENTON, 1983
SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)
MAGNORDER: SQUAMATA, OPPEL, 1811
SUPERORDER: SERPENTES LINNAEUS, 1759
Incertae sedis
Gen. sp indet.
_____________________________________________________________________________________
MADAGASCARMAHAJANGA BASIN
SOUTH OF BERIVORTA
MAD 93-28
MAEVARANO FORMATION
MAASTRICHTIAN STAGE
UPPER SENONIAN SUBEPOCH
UPPER UPPER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
INFRACLASS: NEODIAPSIDA BENTON, 1985
LEGION: LEPIDOSAUROMORPHA BENTON, 1983
SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)
MAGNORDER: SQUAMATA, OPPEL, 1811
SUPERORDER: SERPENTES LINNAEUS, 1759
ORDER: OPHIDIA BRONGNIART, 1800
SUBORDER: ALETHINOPHIDIA NOPCSCA, 1923
Superfamily: BOOIDEA Gray, 1825
Family: BOIDAE Gray, 1825
Subfamily: BOOIDINAE Gray, 1825
Infrafamily: BOOINIEA Gray, 1925
Tribe: MATSOINI Hoffstetter, 1961
Madtsoia madagascariensis
_____________________________________________________________________________________
MADAGASCAR
MAJUNGA PROVINCE
MAHAJANGA BASIN
MAD 93-30
MAEVARANO FORMATION
ANEMBALEMBA MEMBER
MAASTRICHTIAN STAGE
SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
INFRACLASS: NEODIAPSIDA BENTON, 1985
LEGION: LEPIDOSAUROMORPHA BENTON, 1983
SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)
MAGNORDER: SQUAMATA, OPPEL, 1811
SUPERORDER: SERPENTES LINNAEUS, 1759
ORDER: OPHIDIA BRONGNIART, 1800
SUBORDER: ALETHINOPHIDIA NOPCSCA, 1923
Superfamily: BOOIDEA Gray, 1825
Family: BOIDAE Gray, 1825
Subfamily: BOOIDINAE Gray, 1825
Infrafamily: BOOINIEA Gray, 1925
Tribe: MATSOINI Hoffstetter, 1961
Madtsoia madagascariensis
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
AVEPODA PAUL, 2002
SUPERORDER: NEOTHEROPODA BAKKER, 1986
GRANDORDER: CERATOSAURIA NOVAS, 1992
AVEROSTRA PAUL, 2001
ORDER: CERATOSAURIA MARSH, 1884
Superfamily: ABELISAUROIDEA Novas, 1989
Family: NOASAURIDAE Bonaparte & Powell, 1980
Subfamily: VELOCISAURIINAE Bonaparte, 1991 (emend AGNOLIN, NOVAS & APESTEGUIA, 2003)
Masiakasaurus knopfleri
_____________________________________________________________________________________
MADAGASCAR
MAJUNGA PROVINCE
MAHAJANGA BASIN
MAD 93-33
MAEVARANO FORMATION
ANEMBALEMBA MEMBER
MAASTRICHTIAN STAGE
SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: AMPHIBIA LINNAEUS, 1758
SUBCLASS: LISSAMPHIBIA, HAECKEL, 1866?
SUPERORDER: SALIENTIA LAURENTI, 1768
ORDER: ANURA RAFINESQUE, 1815
SUBORDER: MESOBATRACHIA LAURENT, 1979
Superfamily: PELOBATOIDEA Stanius, 1856 , nom. corr. Spinar, 1971 (sensu Bolkay, 1919)
Family: PELOBATIDAE Stanius, 1856 , nom. corr. Spinar, 1971
Gen. sp indet.
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903 OSBORN, 1903
INFRACLASS: NEODIAPSIDA BENTON, 1985
LEGION: LEPIDOSAUROMORPHA BENTON, 1983
SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)
MAGNORDER: SQUAMATA, OPPEL, 1811
SUPERORDER: LACERTILIA OWEN, 1842
GRANDORDER: SCINCOGEKKONOMORPHA SUKHANOV, 1961
MICRORDER: SCINCOMORPHA CAMP, 1923
ORDER: LEPTOGLOSSA COPE, 1900
SUBORDER: ZONUROMORPHA ALIFANOV, 2000
INFRAORDER: ZONURIA FURBRINGER, 1900
Superfamily: CORDYLOIDEA Fritzinger, 1826
Family: CORDYLIDAE? Gray, 1837
*Konkasaurus mahalana
SUPERORDER: SERPENTES LINNAEUS, 1759
ORDER: OPHIDIA BRONGNIART, 1800
SUBORDER: ALETHINOPHIDIA NOPCSCA, 1923
Superfamily: BOOIDEA Gray, 1825
Family: BOIDAE Gray, 1825
Subfamily: BOOIDINAE Gray, 1825
Infrafamily: BOOINIEA Gray, 1925
Tribe: MATSOINI Hoffstetter, 1961
Madtsoia madagascariensis
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: CRUROTARSI SERENO & ARCUCCI, 1990
COHORT: CROCODYLOTARSI BENTON & CLARK, 1988
MAGNORDER: SUCHIA KREBS, 1974
SUPERORDER: CROCODYLOMORPHA WALKER, 1968
GRANDORDER: CROCODILIFORMES HAY, 1930
MICRORDER: MESOEUROCROCODYLIA WHETSTONE & WHYBROW, 1983
ORDER: MESOSUCHIA EMEND HUXLEY, 1875
PARVORDER: METASUCHIA BENTON & CLARK, 1988
GONDWANASUCHIA DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004
NOTOSUCHIMORPHA DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004
NOTOSUCHIFORMES DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004
Family: URUGUAYSUCHIDAE Gasparini, 1971
*Araripesuchus tsangatsangana
TERRIASUCHIA DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004
INFRAORDER: NOTOSUCHIA GASPARINI, 1971
Family: NOTOSUCHIDAE Dollo, 1924
Simosuchus clarki
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
AVEPODA PAUL, 2002
SUPERORDER: NEOTHEROPODA BAKKER, 1986
GRANDORDER: CERATOSAURIA NOVAS, 1992
AVEROSTRA PAUL, 2001
ORDER: CERATOSAURIA MARSH, 1884
Superfamily: ABELISAUROIDEA Novas, 1989
Family: ABELISAURIDAE Bonaparte & Novas, 1985
Subfamily: CARNOTAURINAE Sereno, 1998
Majungatholus atopus
_____________________________________________________________________________________
MADAGASCAR
MAJUNGA PROVINCE
MAHAJANGA BASIN
MAD 93-33
MAEVARANO FORMATION
ANEMBALEMBA MEMBER
MAASTRICHTIAN STAGE
SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: AMPHIBIA LINNAEUS, 1758
SUBCLASS: LISSAMPHIBIA, HAECKEL, 1866?
SUPERORDER: SALIENTIA LAURENTI, 1768
ORDER: ANURA RAFINESQUE, 1815
SUBORDER: NEOBATRACHIA REIG, 1958
Superfamily: HYLOIDEA Rafinesque, 1815 (sensu Wied, 1856)
Family: LEPTODACTYLIDAE Werner, 1896
Subfamily: CERATOPHYRINAE Tschudi, 1838
Beelzebufo ampinga
_____________________________________________________________________________________
MADAGASCAR
MAJUNGA PROVINCE
MAHAJANGA BASIN
MAD 93-34
MAEVARANO FORMATION
ANEMBALEMBA MEMBER
MAASTRICHTIAN STAGE
SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: AMPHIBIA LINNAEUS, 1758
SUBCLASS: LISSAMPHIBIA, HAECKEL, 1866?
SUPERORDER: SALIENTIA LAURENTI, 1768
ORDER: ANURA RAFINESQUE, 1815
SUBORDER: NEOBATRACHIA REIG, 1958
Superfamily: HYLOIDEA Rafinesque, 1815 (sensu Wied, 1856)
Family: LEPTODACTYLIDAE Werner, 1896
Subfamily: CERATOPHYRINAE Tschudi, 1838
Beelzebufo ampinga
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: CHELONOMORPHA KUHN, 1960
SUPERCOHORT: TESTUDINES LINNAEUS, 1758
COHORT: CASICHELYDIA emend GAFFNEY & MEYLAN, 1988
MAGNORDER: PLEURODIRA GAFFNEY & MEYLAN, 1988
ORDER: PLEURODIRIDA COPE, 1864
SUBORDER: EUPLEURODIRA GAFFNEY & MEYLAN, 1988
Superfamily: PELOMEDUSOIDES Cope, 1868 Cope, 1868
Hyperfamily: PODOCNEMIDERA Cope, 1868
CLADE: PAN-PODOCNEMIDIDAE
Family: SAHONACHELYIDAE Joyce, Rollot, Evers, Lyson, Rahantariosa & Krause, 2021
Sokatra antitra
SUBCLASS: DIAPSIDA OSBORN, 1903
INFRACLASS: NEODIAPSIDA BENTON, 1985
LEGION: LEPIDOSAUROMORPHA BENTON, 1983
SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)
MAGNORDER: SQUAMATA, OPPEL, 1811
SUPERORDER: SERPENTES LINNAEUS, 1759
ORDER: OPHIDIA BRONGNIART, 1800
SUBORDER: ALETHINOPHIDIA NOPCSCA, 1923
Superfamily: BOOIDEA Gray, 1825
Family: BOIDAE Gray, 1825
Subfamily: BOOIDINAE Gray, 1825
Infrafamily: BOOINIEA Gray, 1925
Tribe: MATSOINI Hoffstetter, 1961
Madtsoia madagascariensis
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: CRUROTARSI SERENO & ARCUCCI, 1990
COHORT: CROCODYLOTARSI BENTON & CLARK, 1988
MAGNORDER: SUCHIA KREBS, 1974
SUPERORDER: CROCODYLOMORPHA WALKER, 1968
GRANDORDER: CROCODILIFORMES HAY, 1930
MICRORDER: MESOEUROCROCODYLIA WHETSTONE & WHYBROW, 1983
SUBORDER: MESOEUROCROCODYLIA WHETSTONE & WHYBROW, 1983
MESOEUROCRODYLIA: NEW FAMILES?
GONDWANASUCHIA DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004
NOTOSUCHIMORPHA DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004
NOTOSUCHIFORMES DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004
TERRIASUCHIA DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004
INFRAORDER: NOTOSUCHIA GASPARINI, 1971
Family: NOTOSUCHIDAE Dollo, 1924
Simosuchus clarki
PEIROSAURIMORPHA DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004
PEIROSAURIFORMES DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004
Superfamily: PEIROSAURIDAE Gasparini, 1982 emend de Souza Carvalho, Ribeiro & dos Santos Avilla, 2004
Family: PEIROSAURIDAE Gasparini, 1982
Subfamily: LOMASUCHINAE de Souza Carvalho, Ribeiro & dos Santos Avilla, 2004
Tribe MAHAJANGASUCHINI de Souza Carvalho, Ribeiro & dos Santos Avilla, 2004
Mahajangasuchus insignis
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
AVEPODA PAUL, 2002
SUPERORDER: NEOTHEROPODA BAKKER, 1986
GRANDORDER: CERATOSAURIA NOVAS, 1992
AVEROSTRA PAUL, 2001
ORDER: CERATOSAURIA MARSH, 1884
Superfamily: ABELISAUROIDEA Novas, 1989
Family: NOASAURIDAE Bonaparte & Powell, 1980
Subfamily: VELOCISAURIINAE Bonaparte, 1991 (emend AGNOLIN, NOVAS & APESTEGUIA, 2003)
Masiakasaurus knopfleri
_____________________________________________________________________________________
MADAGASCAR
MAHAJANGA PROVINCE
MAHAJANGA BASIN
NEAR BERIVOTRA
MAD 93-35
MAEVARANO FORMATION
CAMPANIAN STAGE?
SENONIAN SUBEPOCH
UPPER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: AMPHIBIA LINNAEUS, 1758
SUBCLASS: LISSAMPHIBIA, HAECKEL, 1866?
SUPERORDER: SALIENTIA LAURENTI, 1768
ORDER: ANURA RAFINESQUE, 1815
SUBORDER: NEOBATRACHIA REIG, 1958
Superfamily: HYLOIDEA Rafinesque, 1815 (sensu Wied, 1856)
Family: LEPTODACTYLIDAE Werner, 1896
Subfamily: CERATOPHYRINAE Tschudi, 1838
Beelzebufo ampinga
SUBORDER: MESOBATRACHIA LAURENT, 1979
Superfamily: PELOBATOIDEA Stanius, 1856 , nom. corr. Spinar, 1971 (sensu Bolkay, 1919)
Family: PELOBATIDAE Stanius, 1856 , nom. corr. Spinar, 1971
Gen. sp indet.
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: CHELONOMORPHA KUHN, 1960
SUPERCOHORT: TESTUDINES LINNAEUS, 1758
COHORT: CASICHELYDIA emend GAFFNEY & MEYLAN, 1988
MAGNORDER: PLEURODIRA GAFFNEY & MEYLAN, 1988
ORDER: PLEURODIRIDA COPE, 1864
SUBORDER: EUPLEURODIRA GAFFNEY & MEYLAN, 1988
Superfamily: PELOMEDUSOIDES Cope, 1868 Cope, 1868
Hyperfamily: PELOMEDUSERA Cope, 1868
Family: PELOMEDUSIDAE Cope, 1868
Subfamily: PELOMEDUSINAE Cope, 1868
Subfamily: Erymnochelus group
Cf. Erymnochelys
Hyperfamily: PODOCNEMIDERA Cope, 1868
CLADE: PAN-PODOCNEMIDIDAE
Family: SAHONACHELYIDAE Joyce, Rollot, Evers, Lyson, Rahantariosa & Krause, 2021
*Sokatra antitra
SUBCLASS: DIAPSIDA OSBORN, 1903
INFRACLASS: NEODIAPSIDA BENTON, 1985
LEGION: LEPIDOSAUROMORPHA BENTON, 1983
SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)
MAGNORDER: SQUAMATA, OPPEL, 1811
SUPERORDER: SERPENTES LINNAEUS, 1759
Incertae sedis
Gen. sp indet.
ORDER: OPHIDIA BRONGNIART, 1800
SUBORDER: ALETHINOPHIDIA NOPCSCA, 1923
Superfamily: BOOIDEA Gray, 1825
Family: BOIDAE Gray, 1825
Subfamily: BOOIDINAE Gray, 1825
Infrafamily: BOOINIEA Gray, 1925
Tribe: MATSOINI Hoffstetter, 1961
Madtsoia madagascariensis
Menarana nosymena
Family; ACROCHORDOIDAE Bonaparte, 1838
Subfamily: ?NIGEROPHEINAE Rage, 1975
Kelyophis hechti
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: CRUROTARSI SERENO & ARCUCCI, 1990
COHORT: CROCODYLOTARSI BENTON & CLARK, 1988
MAGNORDER: SUCHIA KREBS, 1974
SUPERORDER: CROCODYLOMORPHA WALKER, 1968
GRANDORDER: CROCODILIFORMES HAY, 1930
MICRORDER: MESOEUROCROCODYLIA WHETSTONE & WHYBROW, 1983
SUBORDER: MESOEUROCROCODYLIA WHETSTONE & WHYBROW, 1983
MESOEUROCRODYLIA: NEW FAMILES?
GONDWANASUCHIA DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004
NOTOSUCHIMORPHA DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004
NOTOSUCHIFORMES DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004
TERRIASUCHIA DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004
INFRAORDER: NOTOSUCHIA GASPARINI, 1971
Family: NOTOSUCHIDAE Dollo, 1924
Simosuchus clarki
PEIROSAURIMORPHA DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004
PEIROSAURIFORMES DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004
Superfamily: PEIROSAURIDAE Gasparini, 1982 emend de Souza Carvalho, Ribeiro & dos Santos Avilla, 2004
Family: PEIROSAURIDAE Gasparini, 1982
Subfamily: LOMASUCHINAE de Souza Carvalho, Ribeiro & dos Santos Avilla, 2004
Tribe MAHAJANGASUCHINI de Souza Carvalho, Ribeiro & dos Santos Avilla, 2004
*Mahajangasuchus insignis
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
AVEPODA PAUL, 2002
SUPERORDER: NEOTHEROPODA BAKKER, 1986
GRANDORDER: CERATOSAURIA NOVAS, 1992
AVEROSTRA PAUL, 2001
ORDER: CERATOSAURIA MARSH, 1884
Superfamily: ABELISAUROIDEA Novas, 1989
Family: NOASAURIDAE Bonaparte & Powell, 1980
Subfamily: VELOCISAURIINAE Bonaparte, 1991 (emend AGNOLIN, NOVAS & APESTEGUIA, 2003)
Masiakasaurus knopfleri
CLASS: AVES LINNAEUS, 1758
Incertae sedis
Gen.sp indet.
_____________________________________________________________________________________
MADAGASCAR
MAHAJANGA BASIN
SOUTH OF BERIVORTA
MAD 93-36MAEVARANO FORMATION
MAASTRICHTIAN STAGE
UPPER SENONIAN SUBEPOCH
UPPER UPPER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: AMPHIBIA LINNAEUS, 1758
SUBCLASS: LISSAMPHIBIA, HAECKEL, 1866?
SUPERORDER: SALIENTIA LAURENTI, 1768
ORDER: ANURA RAFINESQUE, 1815
SUBORDER: NEOBATRACHIA REIG, 1958
Superfamily: HYLOIDEA Rafinesque, 1815 (sensu Wied, 1856)
Family: LEPTODACTYLIDAE Werner, 1896
Subfamily: CERATOPHYRINAE Tschudi, 1838
Beelzebufo ampinga
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
INFRACLASS: NEODIAPSIDA BENTON, 1985
LEGION: LEPIDOSAUROMORPHA BENTON, 1983
SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)
MAGNORDER: SQUAMATA, OPPEL, 1811
SUPERORDER: SERPENTES LINNAEUS, 1759
ORDER: OPHIDIA BRONGNIART, 1800
SUBORDER: ALETHINOPHIDIA NOPCSCA, 1923
Superfamily: BOOIDEA Gray, 1825
Family: BOIDAE Gray, 1825
Subfamily: BOOIDINAE Gray, 1825
Infrafamily: BOOINIEA Gray, 1925
Tribe: MATSOINI Hoffstetter, 1961
Madtsoia madagascariensis
CLASS: AVES LINNAEUS, 1758
SUBCLASS: SAURORNITHES NICHOLSON, 1879
AVEBREVICAUDA PAUL, 2001
INFRACLASS: ORNITHOTHORACES NOVA
LEGION: ORNITHURAE NOVA
COHORT: ENANTIORNITHES
SUPERORDER: EUENANTIORNITHES CHIAPPE, 2002
ORDER: EUORNITHIFORMES KUROCHKIN, 1996
Family: CONCORNITHIDAE Kurochkin, 1996
Vorona berivotrensis
_____________________________________________________________________________________
MADAGASCAR
MAHAJANGA PROVINCE
MAHAJANGA BASIN
NEAR BERIVOTRA
MAD 93-37
MAEVARANO FORMATION
CAMPANIAN STAGE?
SENONIAN SUBEPOCH
UPPER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: AMPHIBIA LINNAEUS, 1758
SUBCLASS: LISSAMPHIBIA, HAECKEL, 1866?
SUPERORDER: SALIENTIA LAURENTI, 1768
ORDER: ANURA RAFINESQUE, 1815
SUBORDER: MESOBATRACHIA LAURENT, 1979
Superfamily: PELOBATOIDEA Stanius, 1856 , nom. corr. Spinar, 1971 (sensu Bolkay, 1919)
Family: PELOBATIDAE Stanius, 1856 , nom. corr. Spinar, 1971
Gen. sp indet.
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
INFRACLASS: NEODIAPSIDA BENTON, 1985
LEGION: LEPIDOSAUROMORPHA BENTON, 1983
SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)
MAGNORDER: SQUAMATA, OPPEL, 1811
SUPERORDER: SERPENTES LINNAEUS, 1759
ORDER: OPHIDIA BRONGNIART, 1800
SUBORDER: ALETHINOPHIDIA NOPCSCA, 1923
Superfamily: BOOIDEA Gray, 1825
Family; ACROCHORDOIDAE Bonaparte, 1838
Subfamily: ?NIGEROPHEINAE Rage, 1975
Kelyophis hechti
_____________________________________________________________________________________
MADAGASCAR
MAJUNGA PROVINCE
MAHAJANGA BASIN
MAD 93-38
MAEVARANO FORMATION
ANEMBALEMBA MEMBER
MAASTRICHTIAN STAGE
SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
INFRACLASS: NEODIAPSIDA BENTON, 1985
LEGION: LEPIDOSAUROMORPHA BENTON, 1983
SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)
MAGNORDER: SQUAMATA, OPPEL, 1811
SUPERORDER: SERPENTES LINNAEUS, 1759
ORDER: OPHIDIA BRONGNIART, 1800
SUBORDER: ALETHINOPHIDIA NOPCSCA, 1923
Superfamily: BOOIDEA Gray, 1825
Family: BOIDAE Gray, 1825
Subfamily: BOOIDINAE Gray, 1825
Infrafamily: BOOINIEA Gray, 1925
Tribe: MATSOINI Hoffstetter, 1961
Madtsoia madagascariensis
Gen. sp indet.
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
AVEPODA PAUL, 2002
SUPERORDER: NEOTHEROPODA BAKKER, 1986
GRANDORDER: CERATOSAURIA NOVAS, 1992
AVEROSTRA PAUL, 2001
ORDER: CERATOSAURIA MARSH, 1884
Superfamily: ABELISAUROIDEA Novas, 1989
Family: NOASAURIDAE Bonaparte & Powell, 1980
Subfamily: VELOCISAURIINAE Bonaparte, 1991 (emend AGNOLIN, NOVAS & APESTEGUIA, 2003)
Masiakasaurus knopfleri
_____________________________________________________________________________________
MADAGASCAR
MAJUNGA PROVINCE
MAHAJANGA BASIN
MAD 93-40
MAEVARANO FORMATION
ANEMBALEMBA MEMBER
MAASTRICHTIAN STAGE
SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
INFRACLASS: NEODIAPSIDA BENTON, 1985
LEGION: LEPIDOSAUROMORPHA BENTON, 1983
SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)
MAGNORDER: SQUAMATA, OPPEL, 1811
SUPERORDER: SERPENTES LINNAEUS, 1759
Incertae sedis
Gen. sp indet.
_____________________________________________________________________________________
MADAGASCAR
MAJUNGA PROVINCE
MAHAJANGA BASIN
MAD 93-44
MAEVARANO FORMATION
ANEMBALEMBA MEMBER
MAASTRICHTIAN STAGE
SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
INFRACLASS: NEODIAPSIDA BENTON, 1985
LEGION: LEPIDOSAUROMORPHA BENTON, 1983
SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)
MAGNORDER: SQUAMATA, OPPEL, 1811
SUPERORDER: SERPENTES LINNAEUS, 1759
Incertae sedis
Gen. sp indet.
_____________________________________________________________________________________
MADAGASCAR
MAJUNGA PROVINCE
MAHAJANGA BASIN
MAD 93-73
MAEVARANO FORMATION
ANEMBALEMBA MEMBER
MAASTRICHTIAN STAGE
SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: AMPHIBIA LINNAEUS, 1758
SUBCLASS: LISSAMPHIBIA, HAECKEL, 1866?
SUPERORDER: SALIENTIA LAURENTI, 1768
ORDER: ANURA RAFINESQUE, 1815
SUBORDER: NEOBATRACHIA REIG, 1958
Superfamily: HYLOIDEA Rafinesque, 1815 (sensu Wied, 1856)
Family: LEPTODACTYLIDAE Werner, 1896
Subfamily: CERATOPHYRINAE Tschudi, 1838
Beelzebufo ampinga
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
INFRACLASS: NEODIAPSIDA BENTON, 1985
LEGION: LEPIDOSAUROMORPHA BENTON, 1983
SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)
MAGNORDER: SQUAMATA, OPPEL, 1811
SUPERORDER: SERPENTES LINNAEUS, 1759
ORDER: OPHIDIA BRONGNIART, 1800
SUBORDER: ALETHINOPHIDIA NOPCSCA, 1923
Superfamily: BOOIDEA Gray, 1825
Family: BOIDAE Gray, 1825
Subfamily: BOOIDINAE Gray, 1825
Infrafamily: BOOINIEA Gray, 1925
Tribe: MATSOINI Hoffstetter, 1961
*Adinophis fisaka
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: CRUROTARSI SERENO & ARCUCCI, 1990
COHORT: CROCODYLOTARSI BENTON & CLARK, 1988
MAGNORDER: SUCHIA KREBS, 1974
SUPERORDER: CROCODYLOMORPHA WALKER, 1968
GRANDORDER: CROCODILIFORMES HAY, 1930
MICRORDER: MESOEUROCROCODYLIA WHETSTONE & WHYBROW, 1983
ORDER: MESOSUCHIA EMEND HUXLEY, 1875
PARVORDER: METASUCHIA BENTON & CLARK, 1988
GONDWANASUCHIA DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004
NOTOSUCHIMORPHA DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004
NOTOSUCHIFORMES DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004
TERRIASUCHIA DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004
INFRAORDER: NOTOSUCHIA GASPARINI, 1971
Family: NOTOSUCHIDAE Dollo, 1924
Simosuchus clarki
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
AVEPODA PAUL, 2002
SUPERORDER: NEOTHEROPODA BAKKER, 1986
GRANDORDER: CERATOSAURIA NOVAS, 1992
AVEROSTRA PAUL, 2001
ORDER: CERATOSAURIA MARSH, 1884
Superfamily: ABELISAUROIDEA Novas, 1989
Family: ABELISAURIDAE Bonaparte & Novas, 1985
Subfamily: CARNOTAURINAE Sereno, 1998
Majungatholus atopus
_____________________________________________________________________________________
MADAGASCAR
MAJUNGA PROVINCE
MAHAJANGA BASIN
MAD 93-74
MAEVARANO FORMATION
ANEMBALEMBA MEMBER
MAASTRICHTIAN STAGE
SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
AVEPODA PAUL, 2002
SUPERORDER: NEOTHEROPODA BAKKER, 1986
GRANDORDER: CERATOSAURIA NOVAS, 1992
AVEROSTRA PAUL, 2001
ORDER: CERATOSAURIA MARSH, 1884
Superfamily: ABELISAUROIDEA Novas, 1989
Family: NOASAURIDAE Bonaparte & Powell, 1980
Subfamily: VELOCISAURIINAE Bonaparte, 1991 (emend AGNOLIN, NOVAS & APESTEGUIA, 2003)
Masiakasaurus knopfleri
_____________________________________________________________________________________
MADAGASCAR
MAHAJANGA PROVINCE
MAHAJANGA BASIN
NEAR BERIVOTRA
MAD 93-79
MAEVARANO FORMATION
CAMPANIAN STAGE?
SENONIAN SUBEPOCH
UPPER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: CHELONOMORPHA KUHN, 1960
SUPERCOHORT: TESTUDINES LINNAEUS, 1758
COHORT: CASICHELYDIA emend GAFFNEY & MEYLAN, 1988
MAGNORDER: PLEURODIRA GAFFNEY & MEYLAN, 1988
ORDER: PLEURODIRIDA COPE, 1864
SUBORDER: EUPLEURODIRA GAFFNEY & MEYLAN, 1988
Superfamily: PELOMEDUSOIDES Cope, 1868 Cope, 1868
Family: PELOMEDUSIDAE Cope, 1868
Subfamily: PELOMEDUSINAE Cope, 1868
Bothremyididae gen sp indet
_____________________________________________________________________________________
MADAGASCAR
MAJUNGA PROVINCE
MAHAJANGA BASIN
MAD 93-81
MAEVARANO FORMATION
ANEMBALEMBA MEMBER
MAASTRICHTIAN STAGE
SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
INFRACLASS: NEODIAPSIDA BENTON, 1985
LEGION: LEPIDOSAUROMORPHA BENTON, 1983
SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)
MAGNORDER: SQUAMATA, OPPEL, 1811
SUPERORDER: SERPENTES LINNAEUS, 1759
Incertae sedis
Gen. sp indet.
ORDER: OPHIDIA BRONGNIART, 1800
SUBORDER: ALETHINOPHIDIA NOPCSCA, 1923
Superfamily: BOOIDEA Gray, 1825
Family: BOIDAE Gray, 1825
Subfamily: BOOIDINAE Gray, 1825
Infrafamily: BOOINIEA Gray, 1925
Tribe: MATSOINI Hoffstetter, 1961
Madtsoia madagascariensis
_____________________________________________________________________________________
MADAGASCAR
MAJUNGA PROVINCE
MAHAJANGA BASIN
MAD 93-86
MAEVARANO FORMATION
ANEMBALEMBA MEMBER
MAASTRICHTIAN STAGE
SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
INFRACLASS: NEODIAPSIDA BENTON, 1985
LEGION: LEPIDOSAUROMORPHA BENTON, 1983
SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)
MAGNORDER: SQUAMATA, OPPEL, 1811
SUPERORDER: SERPENTES LINNAEUS, 1759
Incertae sedis
Gen. sp indet.
_____________________________________________________________________________________
MADAGASCAR
MAJUNGA PROVINCE
MAHAJANGA BASIN
MAD 93-101
MAEVARANO FORMATION
ANEMBALEMBA MEMBER
MAASTRICHTIAN STAGE
SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
AVEPODA PAUL, 2002
SUPERORDER: NEOTHEROPODA BAKKER, 1986
GRANDORDER: CERATOSAURIA NOVAS, 1992
AVEROSTRA PAUL, 2001
ORDER: CERATOSAURIA MARSH, 1884
Superfamily: ABELISAUROIDEA Novas, 1989
Family: NOASAURIDAE Bonaparte & Powell, 1980
Subfamily: VELOCISAURIINAE Bonaparte, 1991 (emend AGNOLIN, NOVAS & APESTEGUIA, 2003)
Masiakasaurus knopfleri
_____________________________________________________________________________________
MADAGASCAR
MAJUNGA PROVINCE
MAHAJANGA BASIN
MAD 95-07
MAEVARANO FORMATION
ANEMBALEMBA MEMBER
MAASTRICHTIAN STAGE
SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: CRUROTARSI SERENO & ARCUCCI, 1990
COHORT: CROCODYLOTARSI BENTON & CLARK, 1988
MAGNORDER: SUCHIA KREBS, 1974
SUPERORDER: CROCODYLOMORPHA WALKER, 1968
GRANDORDER: CROCODILIFORMES HAY, 1930
MICRORDER: MESOEUROCROCODYLIA WHETSTONE & WHYBROW, 1983
SUBORDER: MESOEUROCROCODYLIA WHETSTONE & WHYBROW, 1983
MESOEUROCRODYLIA: NEW FAMILES?
GONDWANASUCHIA DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004
PEIROSAURIMORPHA DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004
PEIROSAURIFORMES DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004
Superfamily: PEIROSAURIDAE Gasparini, 1982 emend de Souza Carvalho, Ribeiro & dos Santos Avilla, 2004
Family: PEIROSAURIDAE Gasparini, 1982
Subfamily: LOMASUCHINAE de Souza Carvalho, Ribeiro & dos Santos Avilla, 2004
Tribe MAHAJANGASUCHINI de Souza Carvalho, Ribeiro & dos Santos Avilla, 2004
Mahajangasuchus insignis
____________________________________________________________________________________
MADAGASCAR
MAJUNGA PROVINCE
MAHAJANGA BASIN
MAD 95-11
MAEVARANO FORMATION
ANEMBALEMBA MEMBER
MAASTRICHTIAN STAGE
SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: CRUROTARSI SERENO & ARCUCCI, 1990
COHORT: CROCODYLOTARSI BENTON & CLARK, 1988
MAGNORDER: SUCHIA KREBS, 1974
SUPERORDER: CROCODYLOMORPHA WALKER, 1968
GRANDORDER: CROCODILIFORMES HAY, 1930
MICRORDER: MESOEUROCROCODYLIA WHETSTONE & WHYBROW, 1983
ORDER: MESOSUCHIA EMEND HUXLEY, 1875
PARVORDER: METASUCHIA BENTON & CLARK, 1988
GONDWANASUCHIA DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004
NOTOSUCHIMORPHA DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004
NOTOSUCHIFORMES DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004
TERRIASUCHIA DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004
INFRAORDER: NOTOSUCHIA GASPARINI, 1971
Family: NOTOSUCHIDAE Dollo, 1924
Simosuchus clarki
_____________________________________________________________________________________
MADAGASCAR
MAJUNGA PROVINCE
MAHAJANGA BASIN
MAD 95-14
MAEVARANO FORMATION
ANEMBALEMBA MEMBER
MAASTRICHTIAN STAGE
SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
INFRACLASS: NEODIAPSIDA BENTON, 1985
LEGION: LEPIDOSAUROMORPHA BENTON, 1983
SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)
MAGNORDER: SQUAMATA, OPPEL, 1811
SUPERORDER: SERPENTES LINNAEUS, 1759
ORDER: OPHIDIA BRONGNIART, 1800
SUBORDER: ALETHINOPHIDIA NOPCSCA, 1923
Superfamily: BOOIDEA Gray, 1825
Family: BOIDAE Gray, 1825
Subfamily: BOOIDINAE Gray, 1825
Infrafamily: BOOINIEA Gray, 1925
Tribe: MATSOINI Hoffstetter, 1961
Madtsoia madagascariensis
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
AVEPODA PAUL, 2002
SUPERORDER: NEOTHEROPODA BAKKER, 1986
GRANDORDER: CERATOSAURIA NOVAS, 1992
AVEROSTRA PAUL, 2001
ORDER: CERATOSAURIA MARSH, 1884
Superfamily: ABELISAUROIDEA Novas, 1989
Family: NOASAURIDAE Bonaparte & Powell, 1980
Subfamily: VELOCISAURIINAE Bonaparte, 1991 (emend AGNOLIN, NOVAS & APESTEGUIA, 2003)
Masiakasaurus knopfleri
Family: ABELISAURIDAE Bonaparte & Novas, 1985
Subfamily: CARNOTAURINAE Sereno, 1998
Majungatholus atopus
_____________________________________________________________________________________
MADAGASCAR
MAJUNGA PROVINCE
MAHAJANGA BASIN
MAD 95-16
MAEVARANO FORMATION
ANEMBALEMBA MEMBER
MAASTRICHTIAN STAGE
SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
AVEPODA PAUL, 2002
SUPERORDER: NEOTHEROPODA BAKKER, 1986
GRANDORDER: CERATOSAURIA NOVAS, 1992
AVEROSTRA PAUL, 2001
ORDER: CERATOSAURIA MARSH, 1884
Superfamily: ABELISAUROIDEA Novas, 1989
Family: ABELISAURIDAE Bonaparte & Novas, 1985
Subfamily: CARNOTAURINAE Sereno, 1998
Majungatholus atopus
_____________________________________________________________________________________
MADAGASCAR
MAJUNGA PROVINCE
MAHAJANGA BASIN
MAD 96-01
MAEVARANO FORMATION
ANEMBALEMBA MEMBER
MAASTRICHTIAN STAGE
SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
INFRACLASS: NEODIAPSIDA BENTON, 1985
LEGION: LEPIDOSAUROMORPHA BENTON, 1983
SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)
MAGNORDER: SQUAMATA, OPPEL, 1811
SUPERORDER: SERPENTES LINNAEUS, 1759
ORDER: OPHIDIA BRONGNIART, 1800
SUBORDER: ALETHINOPHIDIA NOPCSCA, 1923
Superfamily: BOOIDEA Gray, 1825
Family: BOIDAE Gray, 1825
Subfamily: BOOIDINAE Gray, 1825
Infrafamily: BOOINIEA Gray, 1925
Tribe: MATSOINI Hoffstetter, 1961
Madtsoia madagascariensis
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: CRUROTARSI SERENO & ARCUCCI, 1990
COHORT: CROCODYLOTARSI BENTON & CLARK, 1988
MAGNORDER: SUCHIA KREBS, 1974
SUPERORDER: CROCODYLOMORPHA WALKER, 1968
GRANDORDER: CROCODILIFORMES HAY, 1930
MICRORDER: MESOEUROCROCODYLIA WHETSTONE & WHYBROW, 1983
ORDER: MESOSUCHIA EMEND HUXLEY, 1875
PARVORDER: METASUCHIA BENTON & CLARK, 1988
GONDWANASUCHIA DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004
NOTOSUCHIMORPHA DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004
NOTOSUCHIFORMES DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004
TERRIASUCHIA DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004
INFRAORDER: NOTOSUCHIA GASPARINI, 1971
Family: NOTOSUCHIDAE Dollo, 1924
Simosuchus clarki
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
AVEPODA PAUL, 2002
SUPERORDER: NEOTHEROPODA BAKKER, 1986
GRANDORDER: CERATOSAURIA NOVAS, 1992
AVEROSTRA PAUL, 2001
ORDER: CERATOSAURIA MARSH, 1884
Superfamily: ABELISAUROIDEA Novas, 1989
Family: NOASAURIDAE Bonaparte & Powell, 1980
Subfamily: VELOCISAURIINAE Bonaparte, 1991 (emend AGNOLIN, NOVAS & APESTEGUIA, 2003)
Masiakasaurus knopfleri
Family: ABELISAURIDAE Bonaparte & Novas, 1985
Subfamily: CARNOTAURINAE Sereno, 1998
Majungatholus atopus
CLASS: MAMMALIA LINNAEUS, 1758
SUBCLASS: THERIIFORMES (ROWE, 1988) McKENNA & BELL, 1997
INFRACLASS: ALLOTHERIA (MARSH, 1880) McKENNA & BELL, 1997
ORDER: MULTITUBERCULATA COPE, 1884
SUBORDER: GONDWANATHERIA Mones, 1987
Family: SUDAMERICICIDAE Scillato-Yane & Prasad, 1984
*Lavanify miolaka
_____________________________________________________________________________________
MADAGASCAR
MAJUNGA PROVINCE
MAHAJANGA BASIN
MAD 96-02
MAEVARANO FORMATION
ANEMBALEMBA MEMBER
MAASTRICHTIAN STAGE
SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: ORNITHISCHIFORMES COOPER, 1985
SUPERORDER: SAUROPODOMORPHA HUENE, 1932
GRANDORDER: BRONTOSAURIA OLSHEVSKY, 1991 (emend)
MICRORDER: SAUROPODA MARSH, 1878
ORDER: EUSAUROPODA
SUBORDER: NEOSAUROPODA BONAPARTE, 1986
INFRAORDER: MACRONARIA WILSON & SERENO, 1998
PARVORDER: TITANOSAURIFORMES SALGADO ET AL, 1997
Superfamily: TITANOSAUROIDEA Lydekker, 1885 (sensu Upchurch, 1999)
Family: NEMEGTOSAURIDAE Upchurch, 1995
Subfamily: NEMEGTOSAURINAE Upchurch, 1995
*Rapetosaurus krausei
_____________________________________________________________________________________
MADAGASCAR
MAJUNGA PROVINCE
MAHAJANGA BASIN
MAD 96-03
MAEVARANO FORMATION
ANEMBALEMBA MEMBER
MAASTRICHTIAN STAGE
SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: CHELONOMORPHA KUHN, 1960
SUPERCOHORT: TESTUDINES LINNAEUS, 1758
COHORT: CASICHELYDIA emend GAFFNEY & MEYLAN, 1988
MAGNORDER: PLEURODIRA GAFFNEY & MEYLAN, 1988
ORDER: PLEURODIRIDA COPE, 1864
SUBORDER: EUPLEURODIRA GAFFNEY & MEYLAN, 1988
Superfamily: PELOMEDUSOIDES Cope, 1868 Cope, 1868
Superfamily: BOTHREMYDINURA Baur, 1891
Epifamily: BOTHERMYDINURA Baur, 1891
Family: BOTHREMYDIDAE Baur, 1891
Subfamily: BOTHREMYDINAE Baur, 1891
Infrafamily: BOTHREMYDODDA Baur, 1891
Tribe: KURMADEMYDINI Gaffney, Tong & Meylan, 2006
Kinkonychelys rogersi
_____________________________________________________________________________________
MADAGASCAR
MAHAJANGA BASIN
SOUTH OF BERIVORTA
MAD 96-04MAEVARANO FORMATION
MAASTRICHTIAN STAGE
UPPER SENONIAN SUBEPOCH
UPPER UPPER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
INFRACLASS: NEODIAPSIDA BENTON, 1985
LEGION: LEPIDOSAUROMORPHA BENTON, 1983
SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)
MAGNORDER: SQUAMATA, OPPEL, 1811
SUPERORDER: SERPENTES LINNAEUS, 1759
ORDER: OPHIDIA BRONGNIART, 1800
SUBORDER: ALETHINOPHIDIA NOPCSCA, 1923
Superfamily: BOOIDEA Gray, 1825
Family: BOIDAE Gray, 1825
Subfamily: BOOIDINAE Gray, 1825
Infrafamily: BOOINIEA Gray, 1925
Tribe: MATSOINI Hoffstetter, 1961
Madtsoia madagascariensis
_____________________________________________________________________________________
MADAGASCAR
MAJUNGA PROVINCE
MAHAJANGA BASIN
MAD 96-07
MAEVARANO FORMATION
ANEMBALEMBA MEMBER
MAASTRICHTIAN STAGE
SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
AVEPODA PAUL, 2002
SUPERORDER: NEOTHEROPODA BAKKER, 1986
GRANDORDER: CERATOSAURIA NOVAS, 1992
AVEROSTRA PAUL, 2001
ORDER: CERATOSAURIA MARSH, 1884
Superfamily: ABELISAUROIDEA Novas, 1989
Family: ABELISAURIDAE Bonaparte & Novas, 1985
Subfamily: CARNOTAURINAE Sereno, 1998
Majungatholus atopus
COHORT: ORNITHISCHIFORMES COOPER, 1985
SUPERORDER: SAUROPODOMORPHA HUENE, 1932
GRANDORDER: BRONTOSAURIA OLSHEVSKY, 1991 (emend)
MICRORDER: SAUROPODA MARSH, 1878
ORDER: EUSAUROPODA
SUBORDER: NEOSAUROPODA BONAPARTE, 1986
INFRAORDER: MACRONARIA WILSON & SERENO, 1998
PARVORDER: TITANOSAURIFORMES SALGADO ET AL, 1997
Superfamily: TITANOSAUROIDEA Lydekker, 1885 (sensu Upchurch, 1999)
Family: TITANOSAURIDAE Lydekker, 1885
Incertae sedis
*Vahiny depereti
_____________________________________________________________________________________
MADAGASCAR
MAJUNGA PROVINCE
MAHAJANGA BASIN
MAD 96-18
MAEVARANO FORMATION
ANEMBALEMBA MEMBER
MAASTRICHTIAN STAGE
SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
AVEPODA PAUL, 2002
SUPERORDER: NEOTHEROPODA BAKKER, 1986
GRANDORDER: CERATOSAURIA NOVAS, 1992
AVEROSTRA PAUL, 2001
ORDER: CERATOSAURIA MARSH, 1884
Superfamily: ABELISAUROIDEA Novas, 1989
Family: ABELISAURIDAE Bonaparte & Novas, 1985
Subfamily: CARNOTAURINAE Sereno, 1998
Majungatholus atopus
_____________________________________________________________________________________
MADAGASCAR
MAJUNGA PROVINCE
MAHAJANGA BASIN
MAD 96-21
MAEVARANO FORMATION
ANEMBALEMBA MEMBER
MAASTRICHTIAN STAGE
SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: AMPHIBIA LINNAEUS, 1758
SUBCLASS: LISSAMPHIBIA, HAECKEL, 1866?
SUPERORDER: SALIENTIA LAURENTI, 1768
ORDER: ANURA RAFINESQUE, 1815
SUBORDER: NEOBATRACHIA REIG, 1958
Superfamily: HYLOIDEA Rafinesque, 1815 (sensu Wied, 1856)
Family: LEPTODACTYLIDAE Werner, 1896
Subfamily: CERATOPHYRINAE Tschudi, 1838
Beelzebufo ampinga
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
AVEPODA PAUL, 2002
SUPERORDER: NEOTHEROPODA BAKKER, 1986
GRANDORDER: CERATOSAURIA NOVAS, 1992
AVEROSTRA PAUL, 2001
ORDER: CERATOSAURIA MARSH, 1884
Superfamily: ABELISAUROIDEA Novas, 1989
Family: ABELISAURIDAE Bonaparte & Novas, 1985
Subfamily: CARNOTAURINAE Sereno, 1998
Majungatholus atopus
_____________________________________________________________________________________
MADAGASCAR
MAJUNGA PROVINCE
MAHAJANGA BASIN
MAD 96-24
MAEVARANO FORMATION
ANEMBALEMBA MEMBER
MAASTRICHTIAN STAGE
SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: AMPHIBIA LINNAEUS, 1758
SUBCLASS: LISSAMPHIBIA, HAECKEL, 1866?
SUPERORDER: SALIENTIA LAURENTI, 1768
ORDER: ANURA RAFINESQUE, 1815
SUBORDER: NEOBATRACHIA REIG, 1958
Superfamily: HYLOIDEA Rafinesque, 1815 (sensu Wied, 1856)
Family: LEPTODACTYLIDAE Werner, 1896
Subfamily: CERATOPHYRINAE Tschudi, 1838
Beelzebufo ampinga
_____________________________________________________________________________________
MADAGASCAR
MAJUNGA PROVINCE
MAHAJANGA BASIN
MAD 96-41
MAEVARANO FORMATION
ANEMBALEMBA MEMBER
MAASTRICHTIAN STAGE
SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
AVEPODA PAUL, 2002
SUPERORDER: NEOTHEROPODA BAKKER, 1986
GRANDORDER: CERATOSAURIA NOVAS, 1992
AVEROSTRA PAUL, 2001
ORDER: CERATOSAURIA MARSH, 1884
Superfamily: ABELISAUROIDEA Novas, 1989
Family: NOASAURIDAE Bonaparte & Powell, 1980
Subfamily: VELOCISAURIINAE Bonaparte, 1991 (emend AGNOLIN, NOVAS & APESTEGUIA, 2003)
Masiakasaurus knopfleri
_____________________________________________________________________________________
MADAGASCAR
MAHAJANGA BASIN
SOUTH OF BERIVORTA
MAD 98-08MAEVARANO FORMATION
MAASTRICHTIAN STAGE
UPPER SENONIAN SUBEPOCH
UPPER UPPER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
INFRACLASS: NEODIAPSIDA BENTON, 1985
LEGION: LEPIDOSAUROMORPHA BENTON, 1983
SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)
MAGNORDER: SQUAMATA, OPPEL, 1811
SUPERORDER: SERPENTES LINNAEUS, 1759
ORDER: OPHIDIA BRONGNIART, 1800
SUBORDER: ALETHINOPHIDIA NOPCSCA, 1923
Superfamily: BOOIDEA Gray, 1825
Family: BOIDAE Gray, 1825
Subfamily: BOOIDINAE Gray, 1825
Infrafamily: BOOINIEA Gray, 1925
Tribe: MATSOINI Hoffstetter, 1961
Madtsoia madagascariensis
_____________________________________________________________________________________
MADAGASCAR
MAHAJANGA PROVINCE
MAJUNGA REGION
SE OF BERIVOTRA
MAD 98-17
MAEVERANO FORMATION
MAASTRICHTIAN (PERHAPS LATE MAASTRICHTIAN?) STAGE
UPPERMOST SENONIAN SUBEPOCH
UPPERMOST GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: CRUROTARSI SERENO & ARCUCCI, 1990
COHORT: CROCODYLOTARSI BENTON & CLARK, 1988
MAGNORDER: SUCHIA KREBS, 1974
SUPERORDER: CROCODYLOMORPHA WALKER, 1968
GRANDORDER: CROCODILIFORMES HAY, 1930
MICRORDER: MESOEUROCROCODYLIA WHETSTONE & WHYBROW, 1983
ORDER: MESOSUCHIA EMEND HUXLEY, 1875
PARVORDER: METASUCHIA BENTON & CLARK, 1988
GONDWANASUCHIA DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004
NOTOSUCHIMORPHA DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004
NOTOSUCHIFORMES DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004
TERRIASUCHIA DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004
INFRAORDER: NOTOSUCHIA GASPARINI, 1971
Family: NOTOSUCHIDAE Dollo, 1924
*Simosuchus clarki
_____________________________________________________________________________________
MADAGASCAR
MAHAJANGA PROVINCE
WHICH LIE APPROXIMATELY 4-5 KM SOUTHEAST OF THE MIAN FIELD STUDY AREA SURROUNDING THE VILLAGE OF BERIVOTRA
BETWEEN THE TWO MIADANA HILLS
MAD98-19
MAEVARANO FORMATION
CAMPANIAN STAGE
SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: CRUROTARSI SERENO & ARCUCCI, 1990
COHORT: CROCODYLOTARSI BENTON & CLARK, 1988
MAGNORDER: SUCHIA KREBS, 1974
SUPERORDER: CROCODYLOMORPHA WALKER, 1968
GRANDORDER: CROCODILIFORMES HAY, 1930
MICRORDER: MESOEUROCROCODYLIA WHETSTONE & WHYBROW, 1983
ORDER: MESOSUCHIA EMEND HUXLEY, 1875
PARVORDER: METASUCHIA BENTON & CLARK, 1988
INFRAORDER: METAMESOSUCHIA HULKE, 1878
NEOSUCHIA CLARK, IN BENTON & CLARK, 1988
Family: TREMATOCHAMPSIDAE Buffetaut, 1974
Miadanasuchus oblita
= Trematochampsa oblita
_____________________________________________________________________________________
MADAGASCAR
MAJUNGA PROVINCE
MAHAJANGA BASIN
MAD 98-25
MAEVARANO FORMATION
ANEMBALEMBA MEMBER
MAASTRICHTIAN STAGE
SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: AMPHIBIA LINNAEUS, 1758
SUBCLASS: LISSAMPHIBIA, HAECKEL, 1866?
SUPERORDER: SALIENTIA LAURENTI, 1768
ORDER: ANURA RAFINESQUE, 1815
SUBORDER: NEOBATRACHIA REIG, 1958
Superfamily: HYLOIDEA Rafinesque, 1815 (sensu Wied, 1856)
Family: LEPTODACTYLIDAE Werner, 1896
Subfamily: CERATOPHYRINAE Tschudi, 1838
Beelzebufo ampinga
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: CRUROTARSI SERENO & ARCUCCI, 1990
COHORT: CROCODYLOTARSI BENTON & CLARK, 1988
MAGNORDER: SUCHIA KREBS, 1974
SUPERORDER: CROCODYLOMORPHA WALKER, 1968
GRANDORDER: CROCODILIFORMES HAY, 1930
MICRORDER: MESOEUROCROCODYLIA WHETSTONE & WHYBROW, 1983
ORDER: MESOSUCHIA EMEND HUXLEY, 1875
PARVORDER: METASUCHIA BENTON & CLARK, 1988
GONDWANASUCHIA DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004
NOTOSUCHIMORPHA DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004
NOTOSUCHIFORMES DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004
TERRIASUCHIA DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004
INFRAORDER: NOTOSUCHIA GASPARINI, 1971
Family: NOTOSUCHIDAE Dollo, 1924
Simosuchus clarki
_____________________________________________________________________________________
MADAGASCAR
MAJUNGA PROVINCE
MAHAJANGA BASIN
MAD 98-26
MAEVARANO FORMATION
ANEMBALEMBA MEMBER
MAASTRICHTIAN STAGE
SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
AVEPODA PAUL, 2002
SUPERORDER: NEOTHEROPODA BAKKER, 1986
GRANDORDER: CERATOSAURIA NOVAS, 1992
AVEROSTRA PAUL, 2001
ORDER: CERATOSAURIA MARSH, 1884
Superfamily: ABELISAUROIDEA Novas, 1989
Family: NOASAURIDAE Bonaparte & Powell, 1980
Subfamily: VELOCISAURIINAE Bonaparte, 1991 (emend AGNOLIN, NOVAS & APESTEGUIA, 2003)
Masiakasaurus knopfleri
_____________________________________________________________________________________
MADAGASCAR
MAHAJANGA BASIN
SOUTH OF BERIVORTA
MAD 98-31MAEVARANO FORMATION
MAASTRICHTIAN STAGE
UPPER SENONIAN SUBEPOCH
UPPER UPPER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
INFRACLASS: NEODIAPSIDA BENTON, 1985
LEGION: LEPIDOSAUROMORPHA BENTON, 1983
SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)
MAGNORDER: SQUAMATA, OPPEL, 1811
SUPERORDER: SERPENTES LINNAEUS, 1759
ORDER: OPHIDIA BRONGNIART, 1800
SUBORDER: ALETHINOPHIDIA NOPCSCA, 1923
Superfamily: BOOIDEA Gray, 1825
Family: BOIDAE Gray, 1825
Subfamily: BOOIDINAE Gray, 1825
Infrafamily: BOOINIEA Gray, 1925
Tribe: MATSOINI Hoffstetter, 1961
Madtsoia madagascariensis
_____________________________________________________________________________________
MADAGASCAR
MAJUNGA PROVINCE
MAHAJANGA BASIN
MAD 99-05
MAEVARANO FORMATION
ANEMBALEMBA MEMBER
MAASTRICHTIAN STAGE
SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
AVEPODA PAUL, 2002
SUPERORDER: NEOTHEROPODA BAKKER, 1986
GRANDORDER: CERATOSAURIA NOVAS, 1992
AVEROSTRA PAUL, 2001
ORDER: CERATOSAURIA MARSH, 1884
Superfamily: ABELISAUROIDEA Novas, 1989
Family: NOASAURIDAE Bonaparte & Powell, 1980
Subfamily: VELOCISAURIINAE Bonaparte, 1991 (emend AGNOLIN, NOVAS & APESTEGUIA, 2003)
Masiakasaurus knopfleri
_____________________________________________________________________________________
MADAGASCAR
MAJUNGA PROVINCE
MAHAJANGA BASIN
MAD 99-06
MAEVARANO FORMATION
ANEMBALEMBA MEMBER
MAASTRICHTIAN STAGE
SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
AVEPODA PAUL, 2002
SUPERORDER: NEOTHEROPODA BAKKER, 1986
GRANDORDER: CERATOSAURIA NOVAS, 1992
AVEROSTRA PAUL, 2001
ORDER: CERATOSAURIA MARSH, 1884
Superfamily: ABELISAUROIDEA Novas, 1989
Family: NOASAURIDAE Bonaparte & Powell, 1980
Subfamily: VELOCISAURIINAE Bonaparte, 1991 (emend AGNOLIN, NOVAS & APESTEGUIA, 2003)
Masiakasaurus knopfleri
_____________________________________________________________________________________
MADAGASCAR
MAJUNGA PROVINCE
MAHAJANGA BASIN
MAD 99-15
MAEVARANO FORMATION
ANEMBALEMBA MEMBER
MAASTRICHTIAN STAGE
SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: MAMMALIA LINNAEUS, 1758
SUBCLASS: THERIIFORMES (ROWE, 1988) McKENNA & BELL, 1997
INFRACLASS: ALLOTHERIA (MARSH, 1880) McKENNA & BELL, 1997
ORDER: MULTITUBERCULATA COPE, 1884
SUBORDER: GONDWANATHERIA Mones, 1987
Incertae sedis
*Adalatherium hui
_____________________________________________________________________________________
MADAGASCAR
MAJUNGA PROVINCE
MAHAJANGA BASIN
MAD 99-26
MAEVARANO FORMATION
ANEMBALEMBA MEMBER
MAASTRICHTIAN STAGE
SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
AVEPODA PAUL, 2002
SUPERORDER: NEOTHEROPODA BAKKER, 1986
GRANDORDER: CERATOSAURIA NOVAS, 1992
AVEROSTRA PAUL, 2001
ORDER: CERATOSAURIA MARSH, 1884
Superfamily: ABELISAUROIDEA Novas, 1989
Family: ABELISAURIDAE Bonaparte & Novas, 1985
Subfamily: CARNOTAURINAE Sereno, 1998
Majungatholus atopus
_____________________________________________________________________________________
MADAGASCAR
MAJUNGA PROVINCE
MAHAJANGA BASIN
MAD 99-29
MAEVARANO FORMATION
ANEMBALEMBA MEMBER
MAASTRICHTIAN STAGE
SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: AMPHIBIA LINNAEUS, 1758
SUBCLASS: LISSAMPHIBIA, HAECKEL, 1866?
SUPERORDER: SALIENTIA LAURENTI, 1768
ORDER: ANURA RAFINESQUE, 1815
SUBORDER: NEOBATRACHIA REIG, 1958
Superfamily: HYLOIDEA Rafinesque, 1815 (sensu Wied, 1856)
Family: LEPTODACTYLIDAE Werner, 1896
Subfamily: CERATOPHYRINAE Tschudi, 1838
Beelzebufo ampinga
_____________________________________________________________________________________
MADAGASCAR
MAJUNGA PROVINCE
MAHAJANGA BASIN
MAD 99-30
MAEVARANO FORMATION
ANEMBALEMBA MEMBER
MAASTRICHTIAN STAGE
SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
AVEPODA PAUL, 2002
SUPERORDER: NEOTHEROPODA BAKKER, 1986
GRANDORDER: CERATOSAURIA NOVAS, 1992
AVEROSTRA PAUL, 2001
ORDER: CERATOSAURIA MARSH, 1884
Superfamily: ABELISAUROIDEA Novas, 1989
Family: NOASAURIDAE Bonaparte & Powell, 1980
Subfamily: VELOCISAURIINAE Bonaparte, 1991 (emend AGNOLIN, NOVAS & APESTEGUIA, 2003)
Masiakasaurus knopfleri
_____________________________________________________________________________________
MADAGASCAR
MAHAJANGA BASIN
SOUTH OF BERIVORTA
MAD 99-31MAEVARANO FORMATION
MAASTRICHTIAN STAGE
UPPER SENONIAN SUBEPOCH
UPPER UPPER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
INFRACLASS: NEODIAPSIDA BENTON, 1985
LEGION: LEPIDOSAUROMORPHA BENTON, 1983
SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)
MAGNORDER: SQUAMATA, OPPEL, 1811
SUPERORDER: SERPENTES LINNAEUS, 1759
ORDER: OPHIDIA BRONGNIART, 1800
SUBORDER: ALETHINOPHIDIA NOPCSCA, 1923
Superfamily: BOOIDEA Gray, 1825
Family: BOIDAE Gray, 1825
Subfamily: BOOIDINAE Gray, 1825
Infrafamily: BOOINIEA Gray, 1925
Tribe: MATSOINI Hoffstetter, 1961
Madtsoia madagascariensis
_____________________________________________________________________________________
MADAGASCAR
MAJUNGA PROVINCE
MAHAJANGA BASIN
MAD 99-31
MAEVARANO FORMATION
ANEMBALEMBA MEMBER
MAASTRICHTIAN STAGE
SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: CHELONOMORPHA KUHN, 1960
SUPERCOHORT: TESTUDINES LINNAEUS, 1758
COHORT: CASICHELYDIA emend GAFFNEY & MEYLAN, 1988
MAGNORDER: PLEURODIRA GAFFNEY & MEYLAN, 1988
ORDER: PLEURODIRIDA COPE, 1864
SUBORDER: EUPLEURODIRA GAFFNEY & MEYLAN, 1988
Superfamily: PELOMEDUSOIDES Cope, 1868 Cope, 1868
Superfamily: BOTHREMYDINURA Baur, 1891
Epifamily: BOTHERMYDINURA Baur, 1891
Family: BOTHREMYDIDAE Baur, 1891
Subfamily: BOTHREMYDINAE Baur, 1891
Infrafamily: BOTHREMYDODDA Baur, 1891
Tribe: KURMADEMYDINI Gaffney, Tong & Meylan, 2006
Kinkonychelys rogersi
SUBCLASS: DIAPSIDA OSBORN, 1903
INFRACLASS: NEODIAPSIDA BENTON, 1985
LEGION: LEPIDOSAUROMORPHA BENTON, 1983
SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)
MAGNORDER: SQUAMATA, OPPEL, 1811
SUPERORDER: SERPENTES LINNAEUS, 1759
ORDER: OPHIDIA BRONGNIART, 1800
SUBORDER: ALETHINOPHIDIA NOPCSCA, 1923
Superfamily: BOOIDEA Gray, 1825
Family: BOIDAE Gray, 1825
Subfamily: BOOIDINAE Gray, 1825
Infrafamily: BOOINIEA Gray, 1925
Tribe: MATSOINI Hoffstetter, 1961
Menarana nosymena
_____________________________________________________________________________________
MADAGASCAR
MAJUNGA PROVINCE
MAHAJANGA BASIN
MAD 99-32
MAEVARANO FORMATION
ANEMBALEMBA MEMBER
MAASTRICHTIAN STAGE
SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
AVEPODA PAUL, 2002
SUPERORDER: NEOTHEROPODA BAKKER, 1986
GRANDORDER: CERATOSAURIA NOVAS, 1992
AVEROSTRA PAUL, 2001
ORDER: CERATOSAURIA MARSH, 1884
Superfamily: ABELISAUROIDEA Novas, 1989
Family: ABELISAURIDAE Bonaparte & Novas, 1985
Subfamily: CARNOTAURINAE Sereno, 1998
Majungatholus atopus
_____________________________________________________________________________________
MADAGASCAR
MAJUNGA PROVINCE
MAHAJANGA BASIN
MAD 99-33
MAEVARANO FORMATION
ANEMBALEMBA MEMBER
MAASTRICHTIAN STAGE
SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
AVEPODA PAUL, 2002
SUPERORDER: NEOTHEROPODA BAKKER, 1986
GRANDORDER: CERATOSAURIA NOVAS, 1992
AVEROSTRA PAUL, 2001
ORDER: CERATOSAURIA MARSH, 1884
Superfamily: ABELISAUROIDEA Novas, 1989
Family: NOASAURIDAE Bonaparte & Powell, 1980
Subfamily: VELOCISAURIINAE Bonaparte, 1991 (emend AGNOLIN, NOVAS & APESTEGUIA, 2003)
Masiakasaurus knopfleri
Family: ABELISAURIDAE Bonaparte & Novas, 1985
Subfamily: CARNOTAURINAE Sereno, 1998
Majungatholus atopus
_____________________________________________________________________________________
MADAGASCAR
MAJUNGA PROVINCE
MAHAJANGA BASIN
MAD 99-35
MAEVARANO FORMATION
ANEMBALEMBA MEMBER
MAASTRICHTIAN STAGE
SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: CRUROTARSI SERENO & ARCUCCI, 1990
COHORT: CROCODYLOTARSI BENTON & CLARK, 1988
MAGNORDER: SUCHIA KREBS, 1974
SUPERORDER: CROCODYLOMORPHA WALKER, 1968
GRANDORDER: CROCODILIFORMES HAY, 1930
MICRORDER: MESOEUROCROCODYLIA WHETSTONE & WHYBROW, 1983
SUBORDER: MESOEUROCROCODYLIA WHETSTONE & WHYBROW, 1983
MESOEUROCRODYLIA: NEW FAMILES?
GONDWANASUCHIA DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004
PEIROSAURIMORPHA DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004
PEIROSAURIFORMES DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004
Superfamily: PEIROSAURIDAE Gasparini, 1982 emend de Souza Carvalho, Ribeiro & dos Santos Avilla, 2004
Family: PEIROSAURIDAE Gasparini, 1982
Subfamily: LOMASUCHINAE de Souza Carvalho, Ribeiro & dos Santos Avilla, 2004
Tribe MAHAJANGASUCHINI de Souza Carvalho, Ribeiro & dos Santos Avilla, 2004
Mahajangasuchus insignis
____________________________________________________________________________________
MADAGASCAR
MAHAJANGA BASIN
SOUTH OF BERIVORTA
MAD 99-39MAEVARANO FORMATION
MAASTRICHTIAN STAGE
UPPER SENONIAN SUBEPOCH
UPPER UPPER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
INFRACLASS: NEODIAPSIDA BENTON, 1985
LEGION: LEPIDOSAUROMORPHA BENTON, 1983
SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)
MAGNORDER: SQUAMATA, OPPEL, 1811
SUPERORDER: SERPENTES LINNAEUS, 1759
ORDER: OPHIDIA BRONGNIART, 1800
SUBORDER: ALETHINOPHIDIA NOPCSCA, 1923
Superfamily: BOOIDEA Gray, 1825
Family: BOIDAE Gray, 1825
Subfamily: BOOIDINAE Gray, 1825
Infrafamily: BOOINIEA Gray, 1925
Tribe: MATSOINI Hoffstetter, 1961
Madtsoia madagascariensis
_____________________________________________________________________________________
MADAGASCAR
MAJUNGA PROVINCE
MAHAJANGA BASIN
MAD 01-04
MAEVARANO FORMATION
ANEMBALEMBA MEMBER
MAASTRICHTIAN STAGE
SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: CRUROTARSI SERENO & ARCUCCI, 1990
COHORT: CROCODYLOTARSI BENTON & CLARK, 1988
MAGNORDER: SUCHIA KREBS, 1974
SUPERORDER: CROCODYLOMORPHA WALKER, 1968
GRANDORDER: CROCODILIFORMES HAY, 1930
MICRORDER: MESOEUROCROCODYLIA WHETSTONE & WHYBROW, 1983
SUBORDER: MESOEUROCROCODYLIA WHETSTONE & WHYBROW, 1983
MESOEUROCRODYLIA: NEW FAMILES?
GONDWANASUCHIA DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004
PEIROSAURIMORPHA DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004
PEIROSAURIFORMES DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004
Superfamily: PEIROSAURIDAE Gasparini, 1982 emend de Souza Carvalho, Ribeiro & dos Santos Avilla, 2004
Family: PEIROSAURIDAE Gasparini, 1982
Subfamily: LOMASUCHINAE de Souza Carvalho, Ribeiro & dos Santos Avilla, 2004
Tribe MAHAJANGASUCHINI de Souza Carvalho, Ribeiro & dos Santos Avilla, 2004
Mahajangasuchus insignis
____________________________________________________________________________________
MADAGASCAR
MAHAJANGA BASIN
MASIAKAKOHO STUDY AREA LOCALITY
MAD03-15
MAEVARANO FORMATION
MAASTRICHTIAN STAGE
UPPER SENONIAN SUBEPOCH
UPPER UPPER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
INFRACLASS: NEODIAPSIDA BENTON, 1985
LEGION: LEPIDOSAUROMORPHA BENTON, 1983
SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)
MAGNORDER: SQUAMATA, OPPEL, 1811
SUPERORDER: SERPENTES LINNAEUS, 1759
ORDER: OPHIDIA BRONGNIART, 1800
SUBORDER: ALETHINOPHIDIA NOPCSCA, 1923
Superfamily: BOOIDEA Gray, 1825
Family; ACROCHORDOIDAE Bonaparte, 1838
Subfamily: ?NIGEROPHEINAE Rage, 1975
Kelyophis hechti
_____________________________________________________________________________________
MADAGASCAR
MAHAJANGA BASIN
MASIAKAKOHO STUDY AREA LOCALITY
MAD03-23
MAEVARANO FORMATION
MAASTRICHTIAN STAGE
UPPER SENONIAN SUBEPOCH
UPPER UPPER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
INFRACLASS: NEODIAPSIDA BENTON, 1985
LEGION: LEPIDOSAUROMORPHA BENTON, 1983
SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)
MAGNORDER: SQUAMATA, OPPEL, 1811
SUPERORDER: SERPENTES LINNAEUS, 1759
ORDER: OPHIDIA BRONGNIART, 1800
SUBORDER: ALETHINOPHIDIA NOPCSCA, 1923
Superfamily: BOOIDEA Gray, 1825
Family: BOIDAE Gray, 1825
Subfamily: BOOIDINAE Gray, 1825
Infrafamily: BOOINIEA Gray, 1925
Tribe: MATSOINI Hoffstetter, 1961
Madtsoia madagascariensis
____________________________________________________________________________________
MADAGASCAR
MAJUNGA PROVINCE
MAHAJANGA BASIN
MAD05-59
MAEVARANO FORMATION
ANEMBALEMBA MEMBER
MAASTRICHTIAN STAGE
SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
INFRACLASS: NEODIAPSIDA BENTON, 1985
LEGION: LEPIDOSAUROMORPHA BENTON, 1983
SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)
MAGNORDER: SQUAMATA, OPPEL, 1811
SUPERORDER: SERPENTES LINNAEUS, 1759
Incertae sedis
Gen. sp indet.
ORDER: OPHIDIA BRONGNIART, 1800
SUBORDER: ALETHINOPHIDIA NOPCSCA, 1923
Superfamily: BOOIDEA Gray, 1825
Family: BOIDAE Gray, 1825
Subfamily: BOOIDINAE Gray, 1825
Infrafamily: BOOINIEA Gray, 1925
Tribe: MATSOINI Hoffstetter, 1961
Menarana nosymena
Family; ACROCHORDOIDAE Bonaparte, 1838
Subfamily: ?NIGEROPHEINAE Rage, 1975
Kelyophis hechti
_____________________________________________________________________________________
MADAGASCAR
MAJUNGA PROVINCE
MAHAJANGA BASIN
MAD07-37
MAEVARANO FORMATION
ANEMBALEMBA MEMBER
MAASTRICHTIAN STAGE
SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
INFRACLASS: NEODIAPSIDA BENTON, 1985
LEGION: LEPIDOSAUROMORPHA BENTON, 1983
SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)
MAGNORDER: SQUAMATA, OPPEL, 1811
SUPERORDER: SERPENTES LINNAEUS, 1759
ORDER: OPHIDIA BRONGNIART, 1800
SUBORDER: ALETHINOPHIDIA NOPCSCA, 1923
Superfamily: BOOIDEA Gray, 1825
Family: BOIDAE Gray, 1825
Subfamily: BOOIDINAE Gray, 1825
Infrafamily: BOOINIEA Gray, 1925
Tribe: MATSOINI Hoffstetter, 1961
Menarana nosymena
_____________________________________________________________________________________