SOUTH AFRICA
FREE STATE PROVINCE
HARRISMITH DISTRICT
NORMANDIEN FORMATION
BEAUFORT GROUP
LYSTROSAURUS ASSEMBLAGE ZONE (LYSTROSAURUS ZONE, WATSON, 1914, BROOM, 1906)
EARLY SCYTHIAN EPOCH
EARLY TRIASSIC
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: SYNAPSIDA OSBORN, 1903
SUPERORDER: THERASPIDA BROOM, 1905
GRANDORDER: EUTHERAPSIDA
MICRORDER: NEOTHERAPSIDA
ORDER: THERIODONTIA OWEN, 1881
EUTHERIODONTIA
SUBORDER: SCALOPOSAURIA BRINK, 1963
INFRAORDER: SCALOPOSAURINI BRINK, 1963
Superfamily: SCALOPOSAUROIDEA Broom, 1914 (sensu Brink, 1986)
Family: SCALOPOSAURIDAE Broom, 1914
Scaloposaurus constrictus
_____________________________________________________________________________________
SOUTH AFRICA
FREE STATE PROVINCE
HARRISMITH DISTRICT
GEORGINA 276
NORMANDIEN OR VERKYKERSKOP FORMATION?
BEAUFORT GROUP
LYSTROSAURUS ASSEMBLAGE ZONE (LYSTROSAURUS ZONE, WATSON, 1914, BROOM, 1906)
EARLY SCYTHIAN EPOCH
EARLY TRIASSIC
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: SYNAPSIDA OSBORN, 1903
SUPERORDER: THERASPIDA BROOM, 1905
GRANDORDER: EUTHERAPSIDA
MICRORDER: NEOTHERAPSIDA
ORDER: THERIODONTIA OWEN, 1881
EUTHERIODONTIA
SUBORDER: SCALOPOSAURIA BRINK, 1963
INFRAORDER: SCALOPOSAURINI BRINK, 1963
Superfamily: SCALOPOSAUROIDEA Broom, 1914 (sensu Brink, 1986)
Family: SCALOPOSAURIDAE Broom, 1914
Scaloposaurus constrictus
_____________________________________________________________________________________
SOUTH AFRICA
FREE (ORANGE FREE) STATE
HARRISMITH DISTRICT
JISREEL 419
BURGERSDORP FORMATION
TARKASTAD SUBGROUP
CYNOGNATHUS ZONE
SPATHIAN STAGE
UPPERMOST SCYTHIAN EPOCH TO ANISIAN STAGE
LOWERMOST MID TRIASSIC EPOCH
EARLY TO MIDDLE TRIASSIC
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: SYNAPSIDA OSBORN, 1903
SUPERORDER: THERASPIDA BROOM, 1905
GRANDORDER: EUTHERAPSIDA
MICRORDER: NEOTHERAPSIDA
ORDER: THERIODONTIA OWEN, 1881
EUTHERIODONTIA
SUBORDER: SCALOPOSAURIA BRINK, 1963
INFRAORDER: BAURINI WATSON, 1917
Superfamily: BAUROIDEA Watson, 1911 (sensu Brink, 1986)
Family: BAURIIDAE Watson, 1911
Microgomphodon oligocynus
_____________________________________________________________________________________
SOUTH AFRICA
FREE (ORANGE FREE) STATE PROVINCE
HARRISMITH DISTRICT
GEGUND
DICYNODON ZONE (UPPER CISTECEPHALUS ZONE WATSON, 1914)
TATARIAN AGE
LOPINGIAN SUBEPOCH
ZECHSTEIN EPOCH
LATE PERMIAN
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: AVEMETATARSALIA BENTON, 1999
COHORT: PROLACERTIFORMES CAMP, 1945 NOVA
MAGNORDER: PROTEROSUCHIFORMES nova
ORDER: PROTEROSUCHIA BROOM, 1906
Family: PROTEROSUCHIDAE Huene, 1914
Proterosuchus sp?
_____________________________________________________________________________________
SOUTH AFRICA
FREE (ORANGE FREE) STATE PROVINCE
HARRISMITH DISTRICT
E OF HARRISMITH
IN A DONGA
EARLY TRIASSIC
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: SYNAPSIDA OSBORN, 1903
SUPERORDER: THERASPIDA BROOM, 1905
GRANDORDER: EUTHERAPSIDA
MICRORDER: NEOTHERAPSIDA
ORDER: THERIODONTIA OWEN, 1881
EUTHERIODONTIA
SUBORDER: CYNODONTIA OWEN, 1860
INFRAORDER: PROCYNOSUCHIA
EPICYNODONTIA HOPSON & KITCHING, 2001
Family: THRINAXODONTIDAE Seeley, 1894
Thrinaxodon liorhinus
_____________________________________________________________________________________
SOUTH AFRICA
FREE STATE
HARRISMITH
WONDERKRANTZ
HARRISMITH MEMBER
NORMADIEN FORMATION
LYSTROSAURUS ASSEMBLAGE ZONE
EARLY SCYTHIAN EPOCH
EARLY TRIASSIC
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: SYNAPSIDA OSBORN, 1903
SUPERORDER: THERASPIDA BROOM, 1905
GRANDORDER: EUTHERAPSIDA
MICRORDER: NEOTHERAPSIDA
ORDER: THERIODONTIA OWEN, 1881
EUTHERIODONTIA
SUBORDER: SCALOPOSAURIA BRINK, 1963
INFRAORDER: SCALOPOSAURINI BRINK, 1963
Superfamily: SILPHOLESTIDAE Watson & Romer, 1956 (sensu Brink, 1986)
Family: SILPHOLESTIDAE Watson & Romer, 1956
Tetracynodon darti
_____________________________________________________________________________________
SOUTH AFRICAFREE (ORANGE FREE) STATE
HARRISMITH DISTRICT
WONDERKRANTZ
LYSTROSAURUS ZONE
EARLY SCYTHIAN EPOCH
EARLY TRIASSIC
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: AMPHIBIA LINNAEUS, 1758
SUBCLASS: LABYRINTHODONTIA OWEN, 1860
SUPERORDER: TEMNOSPONDYLIA ZITTEL, 1887
MICROORDER: EUSKELIA YATES & WARREN, 2000
ORDER: EYROPODIA
SUPERFAMILY: DISSOROPHOIDEA BOULENGER, 1902 (SENSU BOLT, 1969)
Family: AMPHIBAMIDAE Moodie, 1910
Micropholis stowii
_____________________________________________________________________________________
SOUTH AFRICA
FREE (ORANGE FREE) STATE PROVINCE
HARRISMITH DISTRICT
QUEEN’S HILL
LOWER LYSTROSAURUS ZONE
EARLY SCYTHIAN EPOCH
EARLY TRIASSIC
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: AMPHIBIA LINNAEUS, 1758
SUBCLASS: LABYRINTHODONTIA OWEN, 1860
SUPERORDER: TEMNOSPONDYLIA ZITTEL, 1887
MICROORDER: EUSKELIA YATES & WARREN, 2000
ORDER: EYROPODIA
SUPERFAMILY: DISSOROPHOIDEA BOULENGER, 1902 (SENSU BOLT, 1969)
Family: AMPHIBAMIDAE Moodie, 1910
Micropholis stowii
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: AVEMETATARSALIA BENTON, 1999
COHORT: PROLACERTIFORMES CAMP, 1945 NOVA
MAGNORDER: PROTOROSAURIA HUXLEY, 1871
Superfamily: PROTOROSAUROIDEA Lydekker, 1888 (sensu Kuhn, 1969)
Family: PROLACERTIDAE Parrington, 1935
Prolacerta broomi
_____________________________________________________________________________________
SOUTH AFRICA
FREE (ORANGE FREE) STATE PROVINCE
HARRISMITH DISTRICT
NORTH OF BRICK-WORKS
KATBERG FORMATION
BEAUFORT GROUP
LYSTROSAURUS ASSEMBLAGE ZONE
EARLY SCYTHIAN EPOCH
EARLY TRIASSIC
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: SYNAPSIDA OSBORN, 1903
SUPERORDER: THERASPIDA BROOM, 1905
GRANDORDER: EUTHERAPSIDA
MICRORDER: NEOTHERAPSIDA
ORDER: THERIODONTIA OWEN, 1881
EUTHERIODONTIA
SUBORDER: CYNODONTIA OWEN, 1860
INFRAORDER: PROCYNOSUCHIA
EPICYNODONTIA HOPSON & KITCHING, 2001
Family: GALESAURIDAE Lydekker, 1890
Galesaurus planiceps
= *Glochinodon detinens
_____________________________________________________________________________________
SOUTH AFRICA
FREE (ORANGE FREE) STATE PROVINCE
HARRISMITH DISTRICT
KATBERG FORMATION
BEAUFORT GROUP
LYSTROSAURUS ASSEMBLAGE ZONE
EARLY SCYTHIAN EPOCH
EARLY TRIASSIC
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: AMPHIBIA LINNAEUS, 1758
SUBCLASS: LABYRINTHODONTIA OWEN, 1860
SUPERORDER: TEMNOSPONDYLIA ZITTEL, 1887
MICROORDER: LIMNARCHIA YATES & WARREN, 2000
ORDER: STEREOSPONDYLOMORPHA YATES & WARREN, 2000
Family: LYDEKKERINIDAE Watson, 1919
*Lydekkerina huxleyi
= Bothriceps huxleyi
= Lydekkerina dutoiti
= *Broomulus dutoiti
= *Limnoiketes paludinatans
Lydekkerina spGen. Sp indet.
SUBORDER: RHINESUCHOIDEA ROMER, 1947
Family: RHINESUCHIDAE Watson, 1919
*Broomistega putterilli
= Lydekkerina putterilliCLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
SUPERLEGION: NEODIAPSIDA BENTON, 1985
Incertae sedis
*Aenigmasaurus grallator
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: AVEMETATARSALIA BENTON, 1999
COHORT: PROLACERTIFORMES CAMP, 1945 NOVA
MAGNORDER: PROTOROSAURIA HUXLEY, 1871
Superfamily: PROTOROSAUROIDEA Lydekker, 1888 (sensu Kuhn, 1969)
Family: PROLACERTIDAE Parrington, 1935
Prolacerta broomi
MAGNORDER: PROTEROSUCHIFORMES nova
ORDER: PROTEROSUCHIA BROOM, 1906
Family: PROTEROSUCHIDAE Huene, 1914
*Proterosuchus fergusi
= Chasmatosaurus vanhoepeniSUBCLASS: SYNAPSIDA OSBORN, 1903
SUPERORDER: THERASPID BROOM, 1905
GRANDORDER: EUTHERAPSIDA
MICRORDER: NEOTHERAPSIDA
ORDER: ANOMODONTIA WATSON & ROMER, 1956
INFRAORDER: DICYNODONTIA OWEN, 1860
Superfamily: PRISTERODONTIA Cluver & King, 1983
Family: KANNEMEYERIDAE Huene, 1948
Subfamily: LYSTROSAURINAE Cope, 1870 (sensu Broom, 1903)
Lystrosaurus declivis
= Dicynodon latirostris
= Ptychognathus latirostris
= Lystrosaurus latirostris
= Ptychosaigum latirostris
= *Lystrosaurus wagneri
Lystrosaurus murrayi
= *Dicynodon strigops
= Prolystrosaurus strigops
= *Lystrosaurus jeppei
Lystrosaurus curvatus
= Dicynodon curvatus
= *Lystrosaurus wageri
= *Lystrosaurus jorisseni
= *Lystrosaurus theileri
Lystrosaurus mccaigi
= Rhabdocephalus mccaigi
= *Lystrosaurus putterilli
Lystrosaurus oviceps
= *Lystrosaurus breyeri
Lystrosarurus spSuperfamily: DIICTODONTIA Cluver & King, 1983
Family: EMYDOPOIDEA Cluver & King, 1983
Subfamily: MYOSAURINAE Cluver & King, 1983
*Myosaurus gracilis
ORDER: THERIODONTIA OWEN, 1881
EUTHERIODONTIA
SUBORDER: SCALOPOSAURIA BRINK, 1963
INFRAORDER: SCALOPOSAURINI BRINK, 1963
Superfamily: SCALOPOSAUROIDEA Broom, 1914 (sensu Brink, 1986)
Family: SCALOPOSAURIDAE Broom, 1914
*Scalopolacerta hoffmani
= Scaloposaurus hoffmaniINFRAORDER: BAURINI WATSON, 1917
Superfamily: BAUROIDEA Watson, 1911 (sensu Brink, 1986)
Famliy: REGISAURIDAE Hopson & Barghusen, 1986
Regisaurus sp
Family: ERICIOLACERTIDAE Watson & Romer, 1956
*Ericiolacerta parva
SUBORDER: CYNODONTIA OWEN, 1860
INFRAORDER: PROCYNOSUCHIA
EPICYNODONTIA HOPSON & KITCHING, 2001
Family: GALESAURIDAE Lydekker, 1890
Galesaurus planiceps
= *Glochinodontoides gracilis
= Galesaurus gracilisFamily: THRINAXODONTIDAE Seeley, 1894
*Platycraniellus elegans
= Platycranion elegans
= Platycranium elegansThrinaxodon liorhinus
= *Ictidopsis elegans
= *Micrictodon marionae
= *Thrinaxodon putterilli
_____________________________________________________________________________________
SOUTH AFRICA
FREE (ORANGE FREE) STATE PROVINCE
HARRISMITH DISTRICT
NEAR HARRISMITH
KATBERG FORMATION
BEAUFORT GROUP
LYSTROSAURUS ASSEMBLAGE ZONE
EARLY SCYTHIAN EPOCH
EARLY TRIASSIC
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: SYNAPSIDA OSBORN, 1903
SUPERORDER: THERASPIDA BROOM, 1905
GRANDORDER: EUTHERAPSIDA
MICRORDER: NEOTHERAPSIDA
ORDER: THERIODONTIA OWEN, 1881
EUTHERIODONTIA
SUBORDER: CYNODONTIA OWEN, 1860
INFRAORDER: PROCYNOSUCHIA
EPICYNODONTIA HOPSON & KITCHING, 2001
Family: THRINAXODONTIDAE Seeley, 1894
Platycraniellus elegans
= Platycranion elegans
= Platycranium elegans
_____________________________________________________________________________________
SOUTH AFRICA
FREE (ORANGE FREE) STATE PROVINCE
HARRISMITH DISTRICT
OLD BRICKFIELDS DONGA
KATBERG FORMATION
BEAUFORT GROUP
LYSTROSAURUS ASSEMBLAGE ZONE
EARLY SCYTHIAN EPOCH
EARLY TRIASSIC
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: AVEMETATARSALIA BENTON, 1999
COHORT: PROLACERTIFORMES CAMP, 1945 NOVA
MAGNORDER: PROTOROSAURIA HUXLEY, 1871
Superfamily: PROTOROSAUROIDEA Lydekker, 1888 (sensu Kuhn, 1969)
Family: PROLACERTIDAE Parrington, 1935
*Prolacerta broomi
SUBCLASS: SYNAPSIDA OSBORN, 1903
SUPERORDER: THERASPID BROOM, 1905
GRANDORDER: EUTHERAPSIDA
MICRORDER: NEOTHERAPSIDA
ORDER: ANOMODONTIA WATSON & ROMER, 1956
INFRAORDER: DICYNODONTIA OWEN, 1860
Superfamily: PRISTERODONTIA Cluver & King, 1983
Family: KANNEMEYERIDAE Huene, 1948
Subfamily: LYSTROSAURINAE Cope, 1870 (sensu Broom, 1903)
Lystrosaurus murrayi
_____________________________________________________________________________________
SOUTH AFRICA
FREE (ORANGE FREE) STATE PROVINCE
HARRISMITH DISTRICT
1 M E OF HARRISMITH
KATBERG FORMATION
BEAUFORT GROUP
LYSTROSAURUS ASSEMBLAGE ZONE
EARLY SCYTHIAN EPOCH
EARLY TRIASSIC
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: AVEMETATARSALIA BENTON, 1999
COHORT: PROLACERTIFORMES CAMP, 1945 NOVA
MAGNORDER: PROTOROSAURIA HUXLEY, 1871
Superfamily: PROTOROSAUROIDEA Lydekker, 1888 (sensu Kuhn, 1969)
Family: PROLACERTIDAE Parrington, 1935
Prolacerta broomi
_____________________________________________________________________________________
SOUTH AFRICA
FREE STATE (ORANGE FREE STATE) PROVINCE
HARRISMITH
LATE TRIASSIC
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
ORDER: Incertae sedis
Gen. sp indet.
____________________________________________________________________________________
SOUTH AFRICA
FREE STATE (ORANGE FREE STATE) PROVINCE
HARRISMITH
RED BEDS
STROMBERG SERIES
LATE TRIASSIC
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: ORNITHISCHIFORMES COOPER, 1985
SUPERORDER: SAUROPODOMORPHA HUENE, 1932
ORDER: PROSAUROPODA HUENE, 1920 (emend)
SUBORDER: PLATEOSAURIA TORNIER, 1913
Family: MASSOSPONDYLIDAE Huene, 1914
Massospondylus carinatus
= *Pachyspondylus orpenii
_____________________________________________________________________________________
SOUTH AFRICA
FREE STATE (ORANGE FREE STATE) PROVINCE
HARRSMITH
BEAUCHERF ABBEY
UPPER ELLIOT FORMATION
HETTANGIAN-LOWER SINEMURIAN STAGE
LOWER LIAS EPOCH
EARLY JURASSIC
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: ORNITHISCHIFORMES COOPER, 1985
SUPERORDER: SAUROPODOMORPHA HUENE, 1932
ORDER: PROSAUROPODA HUENE, 1920 (emend)
SUBORDER: PLATEOSAURIA TORNIER, 1913
Family: MASSOSPONDYLIDAE Huene, 1914
*Massospondylus carinatus
= *Leptospondylus capensis
_____________________________________________________________________________________